A BRYOPHYTE & LICHEN SURVEY

OF TOWNSEND NATURE RESERVE

Bryan Edwards

Dorset Environmental Records Centre

April 2013 For

Dorset Wildlife Trust

DERC Contract 2013_659 Data Log DL 00326

CONTENTS Methods Page 2 Previous records Page 2 Site Description Page 2 esult R s Page 4 Bryophyte and lichen habitats Page 4 Closed limestone grassland Page 4 Open limestone grassland Page 4 Scrub Page 4 Limestone rocks Page 5 Rar e and scarce species Page 5 Red List species Page 5 Nationally Scarce species Page 5 Indicator species Page 5 ssessment Page 9 anagement for bryophytes and lichens Page 11 AM References Page 12 Appendix I: Bryophytes recorded from Townsend NR April 2013 Page 13 ppendix II: Lichens recorded from Townsend NR April 2013 Page 15 A

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METHODS The survey was carried out on the 3rd and 10th April 2013. The area was walked and a species list compiled noting the habitat and substrate they were growing on plus their abundance using the DAFOR scale. Most species were identified in the field using a x10 hand lens, but small pieces of several lichens and liverworts were collected and identified by microscopic examination. or vascular plants nomenclature follows Stace 2010, for bryophytes Hill et al, 2008 and lichens Smith

t al, 2009. Fe PREVIOUS RECORDS Prior to this survey 41 species of lichen and 30 bryophytes had been recorded from the Reserve. Mr G. Barker recorded a short list of lichens in 1977 but there are no further records until August 1994 when I undertook a vegetation survey of the reserve and recorded a number of species in the quadrats. A more thorough survey was carried out in 1995. Bryophytes have received more attention with C. adley and R. Stern both visiting the site in the 1980s. I visited the site in April 1998 and recorded 28 pecies including the local Ditrichum gracile. Hs SITE DESCRIPTION A small site of 12.4 hectares on the Purbeck limestone plateau at Herston on the southern edge of wanage. It comprises limestone grassland with scattered stands of scrub on old quarry workings. The Sgrassland is largely unimproved and managed by pony grazing. Townsend is a Site of Special Scientific Interest and is included in the St Alban’s Heath to Durlston ead Special Area of Conservation (SAC) for its vegetation and populations of Early Gentian

entianella anglica and Early Spider-orchid Ophrys sphegodes. HG

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MAP 1. Survey area with recording compartments

©GetMapping

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RESULTS A total of 4 liverworts, 39 mosses and 80 lichens were recorded from the Reserve. Of these, two ichens are Red Listed and one moss and four lichens are Nationally Scarce being restricted to between 6 and 100 10-km squares in Britain; one lichen is newly recorded for Dorset. l1 BRYOPHYTE AND LICHEN HABITATS

CLOSED LIMESTONE GRASSLAND The typical limestone grassland is short and quite coarse with a few widespread mosses such as Brachythecium rutabulum, Ctenidium molluscum, Homalothecium lutescens, Oxyrrhynchium hians and Pseudoscleropodium purum, plus in more sheltered areas Eurhynchium striatum, Fissidens taxifolius nd Plagiomnium affine. Bare soil on anthills provides a habitat for small acrocarps including Barbula onvoluta, B. unguiculata and Didymodon fallax. a

c OPEN LIMESTONE GRASSLAND A feature of the site is open, summer-parched limestone grassland on the sides and tops of small hummocks and on south-facing banks of the old workings which is referable to the CG7 Festuca ovina-Hieracium pilosella-Thymus spp. grassland. Bryophytes are locally abundant and terricolous lichens are sometimes prominent. The pleurocarpous mosses Hypnum cupressiforme var. lacunosum and Scorpiurium circinatum are most abundant with Aloina aloides, Barbula convoluta, Bryum capillare, Didymodon fallax, Fissidens dubius, Syntrichia ruralis ssp. ruralis, Weissia brachycarpa var. obliqua and W. longifolia var. angustifolia also present. Of particular note are Tortula protobryoides and the Nationally Scarce Didymodon acutus which has only one other recent record in the county. Terricolous lichens are typically present in soil pockets and overgrowing acrocarpous mosses on the top of old spoil heaps. Peltigera rufescens is the most obvious species and is locally frequent to the south of rchid Bastion. Other species include Bilimbia sabuletorum, Cladonia pocillum, C. rangiformis, Collema

uriforme, Leptogium gelatinosum and L. schraderi. Oa SCRUB Blocks of scrub are scattered throughout the site with the largest blocks in the north. It is the richest habitat for lichens with 51 species and is also important for bryophytes with15 species recorded. On the more exposed twigs of Blackthorn Arthonia radiata, Lecanora confusa, Lecidella elaeochroma, Ramalina farinacea and Xanthoria parietina are the characteristic species with larger leafy species Flavoparmelia caperata, F. soredians, Hypotrachyna afrorevoluta, Melanelixia subaurifera, Parmelia sulcata, Parmotrema perlatum and Punctelia subrudecta on older mature stems in more sheltered areas. The shrubby species Evernia prunastri, Hypogymnia physodes and Ramalina fastigiata. The highly basic bark of old Elder supports abundant Xanthoria parietina, but has a number of interesting species including Bacidia delicata, Caloplaca cerina, C. cerinelloides, C. phlogina, and C. ulcerosa. In places these species have spread onto nearby old Traveller’s Joy stems. In the south of the site in more sheltered stands of scrub the smooth bark of Hawthorn and Sycamore support Arthonia spadicea, Dimerella pineti, Opegrapha ochrocheila, O. vulgata, Porina aenea, P. leptalea and Pyrenula macrospora. Lichens of particular note include Bacidia sulphurella found new to Dorset on a Hawthorn, and a single thallus of the rare and beautiful Teloschistes chrysophthalmus was found on an exposed Blackthorn twig. Bryophytes prefer more sheltered conditions than lichens and are therefore restricted to the larger stands of scrub, particularly in the lower northern half of the site. The lower trunks support pleurocarps including Hypnum cupressiforme var. cupressiforme, H. cupressiforme var. resupinatum and Rhynchostegium confertum, with the liverworts Cololejeunea minutissima, Frullania dilatata, Metzgeria furcata and Radula complanata plus the Zygodon conoideus further up. The better lit upper branches, specially horizontal Sycamore branches, have Cryphaea heteromalla, Orthotrichum affine, O.

iaphanum and O. tenellum. 4

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LIMESTONE ROCKS There are very few exposures of natural limestone rock and most of the limestone walls are too shaded to support many lichens or bryophytes; there are numerous small limestone pebbles in the grassland that support a few species. 16 lichens were recorded from limestone rock and most are idespread calcicoles such as Aspicilia calcarea, A. contorta, Caloplaca flavescens, Lecanora albescens,

errucaria baldensis, V. hochstetteri and V. nigrescens. wV RARE AND SCARCE SPECIES R ed List species

Teloschistes chrysophthalmus This very distinctive and beautiful shrubby lichen was thought until recently to be extinct in the British Isles. In 2008 it was found on Blackthorn twigs in the Corfe River valley and since has been found at a further six sites in Dorset and in scattered sites in along the south coast east to Kent. In most sites only one or two thalli are normally present. At Townsend one small thallus was found on the south side of a mature block of Blackthorn (GR SZ02475 78079) in the eastern part of the southern plateau. It was growing in a typical exposed, slightly enriched, twig community with Physcia aipolia, P. enella, Ramalina farinacea and Xanthoria parietina. Despite a lot of suitable habitat it was not found lsewhere.

Critically Endangered

te Caloplaca flavorubescens Endangered; BAP Priority This crust-forming lichen was formerly quite widespread on the sunny, dusty bases of veteran trees, especially Ash, but due to atmospheric pollution and the loss of the host tree it has declined drastically. It is now confined to a widely scattered sites in southern England and central and western Scotland. A very surprising find during the survey was a large thallus present on the hard wood of an old step on he boundary of Cpts 13 and 14. This is an atypical habitat for the species and it will be difficult to onserve it here in the long term. tc N B

ationally Scarce species

ryophytes Didymodon acutus Pointed Beard-moss A small acrocarpous moss found in open chalk and limestone grassland and old quarries. It is scattered through England and Wales and in suitable habitat but is typically found in small quantity. There are just four other records for Dorset and only one recently. At Townsend it was found as scattered patches in three places; one the Southern Plateau SZ02331 78063, in Cpt 3 SZ02315 78339 and in Cpt 6 SZ02460 78318. It is found on south facing banks and knolls in open CG7-type summer parched imestone grassland with Barbula convoluta, B. unguiculata, Didymodon fallax, Hypnum cupressiforme ar. lacunosum, Scorpiurium circinatum and Tortula protobryoides. lv Lichens Caloplaca cerinelloides The tiny (0.2 - 0.3mm) yellow fruits of this crust-forming lichen are found on the exposed twigs on basic barked trees and shrubs such as Ash, Aspen, Elder, Elm. It has only recently been recognised as occurring in the British Isles due to past confusion with C. cerinella. In Dorset it is very local and ainly found on or near the coast. It was noted in the south-east of the Reserve on Elder twigs with

aloplaca cerina and Lecania cyrtella and had spread on to adjacent Traveller’s Joy stems. 5

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Caloplaca pPart of the Caloplaca citrina complex, this species is found on the basic bark of Ash, Elder, Elm, Maple and Sycamore in parklands, pastures and scrub. Its distribution is not fully known in the British Isles due to past confusion with C. citrina s.l.. In Dorset it is quite local and declined with the loss of old Elms nd may decline further if Ash trees suffer a similar fate. Found with Caloplaca ulcerosa on one mature lder bush in the west of the site. hlogina

aE Strigula taylorii A crust-forming lichen found mainly in southern and western parts of the British Isles on the basic ark of Ash, Maple and Sycamore, mainly sheltered old woodland sites. It is widespread in Dorset. ound at Townsend on Ash trees where it forms quite extensive streaks on the smooth bark. bF INDICATOR SPECIES Certain species, like vascular plants, can be used to assess habitat quality. The following list of 15 osses are good indicators of old chalk and limestone grassland with most preferring short and open ward andms some are ‘winter-annuals’ which grow in soil pockets often on south-facing slopes. TABLE 1. Chalk and limestone grassland bryophyte indicator species recorded from Townsend

Aloina aloides Scorpiurium circinatum Campyliadelphus chry yllus soph Tortula lanceola Ctenidium molluscum var. molluscum Tortula protobryoides Didymodon acutus Trichostomum brachydontium Ditrichum gracile1 Trichostomum crispulum Encalypta streptocarpa Weissia brachycar . obliqua pa varHomalothecium lutescens Weissia longifolia var. angustifolia Hypnum cupressiforme var. lacunosum

TABLE 2. Chalk and limestone grassland lichen indicator species recorded from Townsend Bacidia bagliettoana Leptogium schraderi Bilimbia sabuletorum Peltigera rufescens Leptogium gelatinosum

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PLATE 1. Teloschistes chrysophthalmus RDB Critically Endangered, on Blackthorn twig on the south side of a small block of Blackthorn scrub in the east of Cpt 16. This small shrubby lichen was considered extinct in Britain ntil the 1990s. In the last 5 years there have been a number of records from Dorset east to Kent, most of one or wo thalli only on sunny mature scrub. ut

PLATE 2. Caloplaca flavorubescens, RDB Endangered, BAP Priority. On hard lignum of old wooden steps on the oundary of Cpts 13 & 14. This lichen is now very rare in Great Britain and is mainly found on the well-lit bases f pasture and parkland trees. bo 7

PLATE 3. Caloplaca phlogina Nationally Scarce crust-forming lichen associated with well-lit basic bark on the bases of trees in parklands and pastures. At ownsend it is found on several isolated old Elder bushes. AT

LATE 4. Didymodon acutus Pointed Beard-moss Nationally Scarce. In open CG7 grassland on south side of mall knoll in Cpt 16. There is only one other recent record of this species from Dorset. Ps

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ASSESSM NT There are two distinct habitats at Townsend that are important for bryophytes and lichens. Scrub supports the largest number of species with 51 lichens and 15 bryophytes. Of most interest are exposed mature Elder bushes, mature Blackthorn stands and the more sheltered Sycamore. One lichen, Teloschistes chrysophthalmus, is Red Listed and a rare species which grows on the sunny twigs of hrubs such as Blackthorn and Hawthorn. A single thallus was found on the south-side of a small stand f Blackth Plateau (Cpt. 16).

E

so orn in the east of the SouthernTABLE 3. Summary of interest features Number Grid Ref Compartment Interest feature 1 SZ02269 78147 15 Mature Elder with Bacidia delicata, Caloplaca ulcerosa,

logina Caloplaca ph and Opegrapha varia. 2 SZ02332 78059 16 Small knoll with open CG7 grassland with the Nationally Scarce moss Didymodon acutus, plus Hypnum cupressiforme ar. lacunosum, Scorpiurium circinatum and Tortula v

protobryoides. 3 SZ02391 78023 16 Elder supporting Caloplaca cerina and Caloplaca cerinelloides twhich are spreading to old Clema is stems. 4 SZ02475 78079 16 Small block of Blackthorn with a single thallus of the Red Listed lichen Teloschistes chrysophthalmus on the south side. 5 SZ02400 78108 16 South of Orchid Bastion. Several small knolls supporting the best examples of open, parched, CG7 grassland. Bryophytes are abundant with Didymodon acutus, Scorpiurium circinatum nd Tortula protobryoides among the species present. The alocal lichen Peltigera rufescens is locally frequent. 6 SZ02332 78242 7 Small spreading Ash tree with the Nationally Scarce lichen

Strigula taylorii. 7 SZ02292 78238 2 Mature Elder with Caloplaca ulcerosa, Caloplaca phlogina and

Opegrapha varia. 8 SZ02285 78292 2 Mature Hawthorns with a diverse range of epiphytes. One has he lichen Bacidia sulphurella which was found here new to tDorset. 9 SZ02315 78339 3 South-facing back of open limestone grassland supporting the Nationally Scarce moss Didymodon acutus, plus Aloina aloides. 10 SZ02460 78318 6 Small knoll of short limestone grassland with the Nationally

tus, plus Scorpiurium circinatum. Scarce moss Didymodon acu 11 SZ02453 78252 12 Isolated mature Elder with Caloplaca cerina. 12 SZ02500 78263 13 Sycamore, within sheltered scrub, with wide spreading lower branches supporting an abundance of bryophytes including

ololejeunea minutissima, Cryphaea heteromalla, COrthotrichum affine, O. tenellum and Ulota phyllantha.

13 SZ02528 78184 13 Ol ld wooden steps with one large thallus of the Red Listed ichen Caloplaca flavorubescens on the top ‘step’. 9

Map 2. Location of interest features (see table 3 for numbers)

The second habitat is open limestone grassland which supports a number of uncommon bryophytes and lichens, most notably the Nationally Scarce Pointed Beard-moss Didymodon acutus which was found in at least four areas on the Reserve and only has one other recent record in the county. The ocal lichen Peltigera rufescens is frequent on top of some of the old spoil heaps in the CG7 grassland, otably south of ‘Orchid Bastion’.

©GetMapping

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MANAGEMENT FOR BRYOPHYTES AND LICHENS At any grassland sites with stands of scrub there is always a balance of maintaining grazing pressure to control scrub invasion, while retaining some scrub and scrub-grassland interface for invertebrates nd birds. This survey has highlighted two habitats that are important for bryophytes and lichens and he following observations are based on the current management. at Open limestone grassland (CG7)

• The current pony grazing produces a short sward which is ideal for bryophytes and lichens that are poor competitors, and for the rare plants such as Cerastium pumilum, Ophrys sphegodes, Thesium humifusum and Valerianella eriocarpa. The bryophyte and lichen interest is very localised and mainly associated with open summer-parched tops and south-facing sides of old spoil heaps. The main threat appears to be invasion from Burnet Rose Rosa spinosissima.

Scrub

• oWhere possible retain ld isolated Elder bushes which support uncommon lichens. • Sheltered and more mature scrub is most important for both bryophytes and lichens particularly in the lower northern part of the reserve. Therefore it is in these areas that small stands of scrub should be retained. • Sycamore is becoming established in some areas and is potentially a threat to the grassland habitat through shading, a build-up of litter and seedlings becoming established if grazing is reduced. While Sycamore does support a good range of epiphytes most are widespread species and are present on others tree / shrub species. A small number of Sycamore could be retained in scrub areas where grassland habitat is unlikely to be restored, for example in the north-west corner of Cpt 13.

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R EFERENCES

HB ill, MO, Blackstock, TH, Long, DG & Rothero, GP 2008 A Checklist and Census Catalogue of

ritish and Irish Bryophytes. British Bryological Society. SL

mith, CW, Aptroot, A, Coppins, BJ, Fletcher, A, Gilbert, OL, James, PW & Wolseley, PA 2009 The ichens of Great Bri and Ireland. London, British L etain ich n Society. tace, CA 2010 New Flora of the British Isles (3S rd edition). Cambridge. Cambridge University ress. P

APPENDIX I: List of bryophytes recorded from Townsend Nature Reserve, April 2013

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Species English Name Status Habitat Frequency Compartment Liverworts

Cololejeunea minutissima Minute Pouncewort Epiphyte; Ap1, Cm, Cv, Fe, Ps, Sn LF 13 Frullania dilatata Dilated Scalewort Epiphyte; Ap, Cm, Ps O 13 Metzgeria furcata Forked Veilwort Epiphyte; Ap, Cm, Ps, Sn O 13 Radula complanata Even Scalewort Epiphyte; Ap R 11

Mosses Aloina aloides Common Aloe-moss Open limestone grassland R 4 Barbula unguiculata Bird's-claw Beard-moss Ant hills R 16 Brachythecium rutabulum Rough-stalked Feather-moss Grassland, scrub F Calliergonella cuspidata Pointed Spear-moss Grassland R 16 Campyliadelphus chry hyllus sop Golden Feather-moss DN Open limestone grassland R 16 Ctenidium molluscum var. molluscum Chalk Comb-moss Limestone grassland O-LA 16 Cryphaea heteromalla Lateral Cryphaea Epiphyte; Ap, Fe, Ps O 2, 11, 13, 16 Didymodon acutus Pointed Beard-moss NS Open limestone grassland R 4, 6, 16 Didymodon fallax Fallacious Beard-moss Open limestone grassland F 16 Encalypta streptocarpa Spiral Extinguisher-moss DN Limestone grassland R Eurhynchium striat um Common Striated Feather-moss Limestone grassland O-LF Fissidens taxifolius var. taxifolius Common Pocket-moss Limestone grassland O Homalothecium lutescens Yellow Feather-moss Limestone grassland O-LF 16 Hypnum cupressiforme var. cupressiforme Cypress-leaved Plait-moss Scrub O Hypnum cupressiforme var lacunosum . Great Plait-moss DN Open limestone grassland O-LF 16 Hypnum cupressiforme var. resupinatum Supine Plait-moss Epiphyte; Sn O Kindbergia praelonga Common Feather-moss Soil, scrub Orthotrichum affine Wood Bristle-moss Epiphyte; Ap, Ps O 13 Orthotrichum diaphanum White-tipped Bristle-moss Epiphyte; Ap, Cv, Ps R 13 Orthotrichum tenellum Slender Bristle-moss Epiphyte; Ap, Fe R 7 Oxyrrhynchium hians Swartz's Feather-moss Limestone grassland LF 16 Oxyrrhynchium pumilum Dwarf Feather-moss Limestone grassland O Plagiomnium affine Many-fruited Thyme-moss Limestone grassland R 16 Plagiomnium undulatum Hart's-tongue Thyme-moss Scrub O Pseudoscleropodium purum Neat Feather-moss Limestone grassland R Rhynchostegium confertum Clustered Feather-moss Epiphyte; Sn O 13

Species English Name Sta us t Habitat Frequency Compartment Rhynchostegiella tenella Tender Feather-moss Limestone rock R 11, 14 Scorpiurium circinatum Curving Feather-moss DN Open limestone grassland O-LA 16 Syntrichia laevipila Small Hairy Screw-moss Epiphyte; Sn R 7 Syntrichia papillosa Marble Screw-moss Epiphyte; Cm R 12 Syntrichia ruralis var. ruralis Great Hairy Screw-moss Open limestone grassland O 4, 6, 16 Tortula lanceola Lance-leaved Pottia DN Open limestone grassland R Tortula protobryoides Tall Pottia DS Open limestone grassland R 16 Trichostomum brachydontium Variable Crisp-moss Open limestone grassland R Trichostomum crispulum Curly Crisp-moss Open limestone grassland O 16 Ulota phyllantha Frizzled Pincushion Epiphyte; Ap R Weissia brachycar var. obliqua pa Oblique-mouthed Beardless-moss DN Open limestone grassland O 16 Weissia longifolia var angustifolia . Narrow-leaved Beardless-moss DN Open limestone grassland R Zygodon conoideus var. conoideus Green Yoke-moss Epiphyte; Cm, Sn; c.fr R-O 2

1Ap = Acer pseudoplatanus Sycamore Cm = Crataegus mono Hawthorn gynaCv = Clematis vitalba Traveller’s Joy Fe = Fraxinus excels Ash iorPs = Prunus spinosa Blackthorn Sn = Sambucus nigra Elder

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APPENDIX II. List of lichens recorded from Townsend Nature Reserve

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Species Status Dorset status Habitat Frequency Compartment Acrocordia conoidea Limestone rock R 16 Anisomeridium biforme Epiphyte; Ap1 O 14 Anisomeridium polypori Epiphyte; Cv, Fe, Sn O Arthonia radiata Epiphyte; Ps O 16 Arthonia spadicea Epiphyte; Cm R 11 Arthopyrenia punctiformis Epiphyte; O Aspicilia calcarea Limestone rock O 14, 16 Aspicilia contorta Limestone rock R 16 Bacidia bagliettoana DN Limestone grassland R 16 Bacidia delicata Epiphyte; Sn R 15 Bacidia laurocerasi Epiphyte; Ap, Cm, Cv, Sn O 15, 16 Bacidia sulphurella Epiphyte; Cm R 2 Bilimbia sabuletorum Limestone grassland O 16 Caloplaca cerina DN Epiphyte; Cv, Sn O 12, 16 Caloplaca cerinelloides NS Epiphyte; Cv, Sn O 16 Caloplaca flavescens Limestone rock O 4, 16 Caloplaca flavorubescens EN; BAP Lignum R 14 Caloplaca limonia Limestone rock O 4 Caloplaca marmorata DN Limestone rock R 4 Caloplaca obscurella Epiphyte; Cm R 12 Caloplaca phlogina NS Epiphyte; Sn R 2, 15 Caloplaca ulcerosa DN Epiphyte; Sn O 2, 15 Candelariella reflexa Epiphyte; Ap, Ps O 2, 12, 13 Cladonia pocillum Limestone grassland O 16 Cladonia rangiformis Limestone grassland R 2, 16 Cliostomum griffithii Epiphyte; Ap R 14 Collema auriforme Limestone grassland O 16 Dimerella pineti Epiphyte; Cm R 11 Diploicia canescens Limestone rock O 16 Enterographa crassa Epiphyte; Ap, Fe R 7, 11 Evernia prunastri Epiphyte; Ps O 16 Flavoparmelia caperata Epiphyte; Ap, Ps O 13, 14, 16 Flavoparmelia soredians Epiphyte; Ps O 16

Species Status Dorset status Habitat Frequency Co

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mpartment Fuscidea lightfootii Epiphyte; Ps R 16 Hyperphyscia adglutinata Epiphyte; Sn R 16 Hypogymnia physodes Epiphyte; Ps R 16 Hypotrachyna afrorevoluta Epiphyte; Ps O 16 Lecania cyrtella Epiphyte; Bd, Cv, Sn O 16 Lecania naegelii Epiphyte; Cv, Sn O Lecanora albescens Limestone rock R Lecanora barkmaniana Epiphyte; Ps R 14 Lecanora chlarotera Epiphyte; Ap, Fe, Ps O Lecanora confusa Epiphyte; Ps O Lecidella elaeochroma Epiphyte; Ap, Cm, Fe, Ps, Sn F Lepraria lobificans Epiphyte; Ps, Sn O 16 Leptogium gelatinosum DN Limestone grassland R 16 Leptogium schraderi Limestone grassland O 16 Melanelixia glabratula Epiphyte; Cm O Melanelixia subaurifera Epiphyte; Ap, Cm, Ps O Opegrapha atra Epiphyte; Ap O 13, 14 Opegrapha calcarea Limestone rock R 16 Opegrapha ochrocheila Epiphyte; Ap R 3 Opegrapha varia Epiphyte; Sn O 2, 15 Opegrapha vulgata Epiphyte; Ap R 11 Parmelia sulcata Epiphyte; Ps O 16 Parmotrema perlatum Epiphyte; Ap, Cm, Fe, Ps O 2, 7, 10, 11, 12, 13 Peltigera rufescens DN Limestone grassland O 2, 16 Phaeophyscia orbicularis Epiphyte; Ap, Cm, Sn O Physcia aipolia Epiphyte; Ps O Physcia tenella Epiphyte; Ap, Cm, Cv, Fe, Ps F Physconia grisea Epiphyte; Sn R Placynthium nigrum Limestone rock R Porina aenea Epiphyte; Ap R 11 Porina leptalea Epiphyte; Cm R 2, 11 Porina linearis Limestone rock R 4 Protoblastenia rupestris Limestone rock R 16 Punctelia borreri Epiphyte; Ps R 16 Punctelia jeckeri Epiphyte; Cm R 16

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Species Status Dorset status Habitat Frequency Compartment Punctelia subrudecta Epiphyte; Ps O 16 Pyrenula macrospora Epiphyte; Ap R 11 Ramalina farinacea Epiphyte; Ap, Cm, Fe, Ps F Ramalina fastigiata Epiphyte; Ap, Cm, Fe, Ps O Sarcogyne regularis Limestone rock R 16 Strigula taylorii NS Epiphyte; Fe R 7 Teloschistes chrysophthalmus CR Epiphyte; Ps R 16 Verrucaria baldensis Limestone rock O Verrucaria hochstetteri Limestone rock R Verrucaria nigrescens Limestone rock O Xanthoria parietina Epiphyte; Ap, Bd, Cv, Cm, Fe, Ps, Sn F-LA Xanthoria polycarpa Epiphyte; Ps O

1A = Acer pseudoplat Sycamore p anusBd = Buddleja davidii Buddleja Cm = Crataegus mono Hawthorn gynaCv = Clematis vitalba Traveller’s Joy Fe = Fraxinus excels Ash iorPs = Prunus spinosa Blackthorn Sn = Sambucus nigra Elder