a verification of reports of marine algal species from the
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Philippine Journal of Science142: 5-49, Special IssueISSN 0031 - 7683Date Received: 2 May 2013
Marine Science LaboratorySchool of Life Sciences, Chinese University of Hong Kong
Shatin, N.T., Hong Kong SAR, CHINA
Put O. Ang, Jr., Sin Man Leung, and Mei Mei Choi
A Verification of Reports of Marine Algal Species from the Philippines
Corresponding Author: [email protected]
Records of marine macroalgae reported from the Philippines were checked against AlgaeBase, the international database for algal nomenclatures, and Index Nominum Algarum (INA) Bibliographia Phycologica Universalis of the University of California at Berkeley Silva Center for Phycological Documentation to verify their present nomenclature, status of taxonomy and bibliographic reference. To date, 306 names of taxa (including species, varieties and forms) of greens (Chlorophyta), 234 names of taxa of browns (Ochrophyta, Phaeophyceae) and 751 names of taxa of reds (Rhodophyta), or a total of 1291 published names of taxa have been reported from the Philippines. Of these, 231 taxa representing 197 species in 20 families for green algae, 171 taxa representing 153 species in 10 families for brown algae, and 564 taxa representing 543 species in 52 families for red algae are considered valid records listed with their currently accepted names. All in all, 966 currently accepted taxa, representing 893 species in 82 families of marine macroalgae have been reported from the Philippines. Among the greens, 15 taxa have their type localities in the Philippines. This number is 40 for the browns and 33 for the reds. Proportionally, this is 6.5% of the total for the greens, 23.4% for the browns and 5.9% for the reds.
Key Words: Philippine marine macroalgae, algal taxonomic status, algal type species from the Philippines, AlgaeBase, Index Nominum Algarum (INA)
INTRoDUCTIoN Philippines is known for its rich marine flora. over the years, several efforts have been attempted to compile and annotate the records of marine algae collected from the Philippines, including those by Silva et al. (1987), the two volume works by Trono (1997, 2004), and more recently, the checklist with a cut off date at 1999 by Ganzon-Fortes (2012). The last mentioned literature compiled a list of 949 algal species, including 61 Cyanobacteria, 212 Chlorophytes, 154 ochrophytes (Phaeophyceaens) and 522 Rhodophytes. Several recent works since 1999 have contributed to the long list of Philippine marine algal records. Notable of these included the works of Geraldino
et al. (2005) on Padina, Belleza and Liao (2007) on Caulerpa, Geraldino et al. (2010) on Hypnea and Payo et al. (2013) on Portieria.
With the extensive application of molecular techniques in studies on algal systematics in recent years, major revisions have been made in many groups of tropical marine algae. There is therefore a need to update the nomenclatures of Philippine marine algal records. This nomenclature validation is important to algal biodiversity, conservation and biogeographical studies in order to provide a more accurate picture of floristic connection and the relatedness of Philippine flora with those of other neighbouring regions. The main objective of this current work is therefore to compile all known records of Philippine macroalgae from the literature till April of
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2013 and check them against AlgaeBase (Guiry and Guiry 2013), the international database of algal nomenclatures, and Index Nominum Algarum (INA) Bibliographia Phycologica Universalis of the University of California at Berkeley Silva Center for Phycological Documentation (Silva 1996-to date) to come up with the most updated Philippine records. Furthermore, species with their type locality within the territorial waters of the Philippines are also checked and listed.
MATeRIALS AND MeTHoDSThis compilation work relied initially on the main cata-logue of marine algae reported from the Philippines by Silva et al. (1987). Further updates were based on works by Trono (1997, 2004) and Ganzon-Fortes (2012). In ad-dition, all relevant, more recent (post 1999) literatures were consulted. In case of doubts, original literatures were also consulted for confirmation of the records. These literatures are all included in the Reference list.
The compiled list of algae was then double checked for duplication. The revised list was further checked for synonymy and invalidity of taxa based on AlgaeBase (Guiry and Guiry 2013) and INA Database (Silva 1996-to date). The most recent information on family clas-sification, name of authorities, and type localities were also checked and accepted to come up with the final list. Where type localities of species are reported to be in the Philippines, the protologues were consulted, or reference was made to Silva (1996-to date), for verification.
ReSULTS AND DISCUSSIoN
This compilation includes only taxa of marine macroal-gae belonging to the three major phyla, the Chlorophyta (Green), ochrophyta (Pheophyceae, Brown) and Rho-dophyta (Red) listed in Tables 1 to 3 respectively. For ease of reference, the scientific names of listed taxa are arranged alphabetically first by the family classification, followed by the generic and species epithets. The abbre-viation of the authority names of all binomials follows that used in AlgaeBase. For consideration of nomencla-tural priority, the original year of publication was added at the end of each currently accepted taxon listed.
In total, 306 published names of taxa (including species, varieties and forms) of greens, 234 names of browns and 751 names of reds, or a total of 1291 names of algal taxa have been reported from the Philippines. of these, there are 231 validly published taxa representing 197 species in 20 families for green algae (Table 1), 171 taxa repre-senting 153 species in 10 families for brown algae (Table
2), and 564 taxa representing 543 species in 52 fami-lies for red algae (Table 3). Among the reds, four taxa are generic records only (Euptilota sp., Callithamnion sp., Dudresnaya sp. and Rhabdonia sp.). These genera are unique records, although with no specific names re-ported for the Philippine specimens, and are therefore counted as part of the 543 species recorded. Family Rhabdoniaceae, where Rhabdonia sp. belongs, also rep-resents a unique family with only one known species re-corded from the Philippines. The family of Callophycus serratus cannot be ascertained, hence, is classified un-der “Rhodophyta incertae sedis” by AlgaeBase. All in all, 966 currently accepted valid taxa, representing 893 species in 82 families of marine macroalgae have been reported from the Philippines.
Synonyms or invalid names recorded for different spe-cies under the three major groups of algae are also listed respectively in Tables 1 to 3, across their currently ac-cepted names. Some invalid names are clearly due to spelling error (e.g. Sargassum siliquosum vs. S. siloquo-sum), some are simply due to incorrect gender agreement with the generic name (e.g. Kappaphycus striatus vs. K. striatum). These variations of nomenclature in print are listed here for record purposes. Among the greens, Enteromorpha aragoensis Bliding is considered illegiti-mate due to lack of a type citation (Table 1). Under each phylum, there are published records of taxa which have no matching records in AlgaeBase. These are separately listed. These taxa have also been checked against INA Database. Those which are not also found in INA Da-tabase probably represent invalid names. Furthermore, some taxa are classified under “Preliminary Entry” in Al-gaeBase. These are taxa where detailed studies on their taxonomic status are not available. These are all listed in the respective Tables for record purposes and are not included in the total count of species.
Among the greens, 15 taxa have their type localities in the Philippines. This number is 40 for the browns and 33 for the reds. Proportionally, this is 6.5% of the total for the greens, 23.4% for the browns and 5.9% for the reds. The high proportion of type species (taxa) in browns is mainly due to the relatively high number of Sargassum taxa reported as being new to science by Grunow (1915, 1916) (16 taxa) and Trono (1994) (12 species). Two taxa (Sargassum cinctum var. jagorii Grunow and S. crassi-folium var. subduplicatum (‘subduplicata’) Grunow) de-scribed by Grunow (1915) based on materials from the Philippines are not recorded in AlgaeBase (see Table 2). To include these would increase the proportion of brown algae with type or syntype locality in the Philippines to 24.3%. on the other hand, the green alga Cladophora
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Table 1. Listing of green (Chlorophyta) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed.
List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality
1 Anadyomenaceae Anadyomene eseptata Gilbert 1961Dalupiri Island, Babuyan Islands, Cagayan, Luzon, Philippines (Gilbert 1961: 425).
2 Anadyomene leclancheri Decaisne 1844 Microdictyon clathratum G. Martens
Syntype localities: Zamboanga, Min-danao, Philippines; Seram and Flores, Indonesia (Martens 1866: 66-67).
3 Anadyomene plicata C. Agardh 1823
4 Anadyomene stellata (Wulfen) C. Agardh 1823
Anadyomene brownii Schroder
5 Anadyomene wrightii Harvey ex J. e. Gray 1866
6 Microdictyon boergesenii Setchell 1925
7 Microdictyon japonicum Setchell 1925
8 Microdictyon okamurae Setchell 1925
9 Microdictyon tenuius J. e. Gray 1866 Microdictyon agardhianum Decaisne
10 Boodleaceae Boodlea coacta (Dickie) G. Murray & De Toni 1889
11 Boodlea composita (Harvey) F. Brand 1904
12 Boodlea montagnei (Harvey ex J. e. Gray) egerod 1952
13 Boodlea struveoide M. A. Howe 1918
14 Cladophora herpestica (Montagne) Kützing 1849
Cladophoropsis zollingeri (Kützing) Reinbold
15 Cladophoropsis fasciculata (Kjellman) Wille 1910
Cladophoropsis sundanensis Reinbold
16 Cladophoropsis gracillima e. Y. Dawson 1950
17 Cladophoropsis membranacea (Hofman Bang ex C. Agardh) Børgesen 1905
18 Cladophoropsis philippinensis W. R. Taylor 1961
Little Santa Cruz Island, opposite Zamboanga, The Philippines (Taylor 1961: 58).
19 Cladophoropsis vaucheriiformis (Areschoug) Papenfuss 1958
Cladophoropsis dichotoma (Zanardini) Papenfuss; Cladophoropsis neocaledon-ica (Grunow ex G. Murray & Boodle) Papenfuss
20 Phyllodictyon anastomosans (Harvey) Kraft & M. J. Wynne 1996
Struvea anastomosans (Harvey) Piccone & Grunow ex Piccone
21 Phyllodictyon pulcherrimum J. e. Gray 1866
Microdictyon curtissiae W. R. Taylor; Struvea ramosa Dickie
22 Struvea okamurae Leliaert 2007 Chamaedoris orientalis okamura & Higashi
23 Bryopsidaceae Bryopsis corticulans Setchell 1899
24 Bryopsis indica A. Gepp & e. S. Gepp 1908
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25 Bryopsis pennata J. V. Lamouroux 1809
26 Bryopsis pennata var. secunda (Harvey) Collins & Hervey 1917
27 Bryopsis plumosa (Hudson) C. Agardh 1823
28 Caulerpaceae Caulerpa antoensis Yamada 1940 Caulerpa arenicola W. R. Taylor
29 Caulerpa brachypus Harvey 1860
30 Caulerpa brachypus f. parvifolia (Harvey) A. B. Cribb 1958 Caulerpa parvifolia Harvey
31 Caulerpa cupressoides (M. Vahl) C. Agardh 1817
32 Caulerpa cupressoides f. amicorum (Harvey) Weber-van Bosse 1898
33 Caulerpa cupressoides f. disticha Weber-van Bosse 1898
34Caulerpa cupressoides var. [lycopodium] f. elegans (P. L. Crouan & H. M. Crouan) Weber-van Bosse 1898
Caulerpa cupressoides var. lycopodium f. elegans (P. L. & H. M. Crouan) Weber-van Bosse
35 Caulerpa cupressoides var. ericifolia (Turner) Weber-van Bosse 1898
36 Caulerpa cupressoides var. lycopodium Weber-van Bosse 1898
37Caulerpa cupressoides var. urvilleana (Montagne) L. J. Hodgson, Pham Huu Tri, Lewmanomont & McDermid 2004
Caulerpa urvilleana Mon-tagne
38 Caulerpa elongata Weber-van Bosse 1898
39 Caulerpa fastigiata Montagne 1837
40 Caulerpa fergusonii G. Murray 1891
41 Caulerpa filicoides Yamada 1936
42 Caulerpa lentillifera J. Agardh 1837
43 Caulerpa lentillifera var. compacta G. C. Trono & Ang 1982
Bugsuk, Palawan, Philippines (Trono and Ang 1982: 7)
44 Caulerpa lessonii Bory de Saint-Vincent 1828
45 Caulerpa mexicana Sonder ex Kützing 1849
46 Caulerpa mexicana var. pluriseriata W. R. Taylor 1975
47 Caulerpa microphysa (Weber-van Bosse) Feldmann 1955
48 Caulerpa peltata J. V. Lamouroux 1809
Caulerpa racemosa var. laetevirens (Montagne) Weber-van Bosse; Caulerpa racemosa var. peltata (J. V. Lamouroux) eubank
49 Caulerpa peltata var. macrodisca (Decaisne) Weber-van Bosse 1898
Caulerpa macrodisca Decaisne
50 Caulerpa prolifera (Forsskål) J. V. Lamouroux 1809
51 Caulerpa racemosa (Forsskål) J. Agardh 1873
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52 Caulerpa racemosa var. lamourouxii (Turner) Weber-van Bosse 1898
53 Caulerpa racemosa var. macra Weber-van Bosse 1913
54 Caulerpa racemosa var. macrophysa (Sonder ex Kützing) W. R. Taylor 1928
55 Caulerpa racemosa var. occidentalis (J. Agardh) Børgesen 1907
56 Caulerpa racemosa var. turbinata (J. Agardh) eubank 1946
57 Caulerpa reyesii Menez & H. P. Calumpong 1982
58 Caulerpa selago (Turner) C. Agardh 1817
59 Caulerpa serrulata (Forsskål) J. Agardh 1837
60 Caulerpa serrulata f. lata (Weber-van Bosse) C. K. Tseng 1936
61 Caulerpa serrulata f. spiralis (Weber-van Bosse) Gilbert 1942
62 Caulerpa serrulata var. boryana (J. Agardh) Gilbert 1942
63 Caulerpa serrulata var. boryana f. longifolia Gilbert 1942
64Caulerpa serrulata var. boryana f. oc-cidentalis (Weber-van Bosse) Yamada & Tanaka 1938
65 Caulerpa serrulata var. pectinata (Weber-van Bosse) W. R. Taylor 1960
66 Caulerpa sertularioides (S. G. Gmelin) M. A. Howe 1905
67 Caulerpa sertularioides f. brevipes (J. Agradh) Svedelius 1906
68 Caulerpa sertularioides f. farlowii (Weber-van Bosse) Børgesen 1907
69 Caulerpa sertularioides f. longipes (J. Agardh) Collins 1909
70 Caulerpa sertularioides f. longiseta (Bory de Saint-Vincent) Svedelius 1906
71 Caulerpa subserrata okamura 1897
72 Caulerpa taxifolia (M. Vahl) C. Agardh 1817
73 Caulerpa urvilleana var. vitiensis Weber-van Bosse 1898
74 Caulerpa verticillata J. Agardh 1847
75 Caulerpa vesiculifera (Harvey) Harvey 1863
76 Caulerpa webbiana Montagne 1837
77 Caulerpa webbiana var. pickeringii (Har-vey & Bailey) eubank 1946
78 Caulerpella ambigua (okamura) Prud'homme van Reine & Lokhorst 1992 Caulerpa ambigua okamura
79 Cladophoraceae Chaetomorpha aerea (Dillwyn) Kützing 1849
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80 Chaetomorpha antennina (Bory de Saint-Vincent) Kützing 1847
81 Chaetomorpha brachygona Harvey 1858
82 Chaetomorpha crassa (C. Agardh) Kützing 1845
83 Chaetomorpha gracilis Kützing 1845
84 Chaetomorpha inflata Kützing 1849
85 Chaetomorpha kellersii M. A. Howe 1932 Panay Island, Philippines (Howe 1932: 169)
86 Chaetomorpha ligustica (Kützing) Kützing 1849
87 Chaetomorpha linum (o. F. Müller) Kützing 1845
88 Chaetomorpha philippinensisLeliaert 2011
Dapdap, Siquijor,Philippines (Leliaert etal. 2011: 385)
89 Chaetomorpha spiralis okamura 1903
90 Chaetomorpha vieillardii(Kützing) M. J. Wynne 2011
91 Cladophora albida (Nees) Kützing 1843
92 Cladophora aokii Yamada 1925
93 Cladophora catenata (Linnaeus) Kützing 1843
94 Cladophora conferta P. L. Crouan & H. M. Crouan 1865
95 Cladophora crispula Vickers 1905
96 Cladophora crucigera Grunow 1867
97 Cladophora cymopoliae Børgesen 1925
98 Cladophora dalmatica Kützing 1843
99 Cladophora japonica Yamada 1931
100 Cladophora laetevirens (Dillwyn) Kützing 1843
101 Cladophora liebetruthii Grunow 1884
102 Cladophora pellucida (Hudson) Kützing 1843
103 Cladophora prolifera (Roth) Kützing 1843
104 Cladophora quisumbingii Manza 1939 Batan Island, Batanes, Luzon, Philip-pines (Manza 1939: 109)
105 Cladophora rupestris (Linnaeus) Kützing 1843
106 Cladophora sakaii I. A. Abbott 1972
107 Cladophora sericea (Hudson) Kützing 1843
Cladophora crystallina (Roth) Kützing
108 Cladophora sibogae Reinbold 1905
109 Cladophora vagabunda (Linnaeus) Hoek 1963 Cladophora inserta Dickie
110 Cladophora wrightiana Harvey 1860
111 Rhizoclonium africanum Kützing 1853
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112 Rhizoclonium crassipellitum var. robustum G. S. West 1904
113 Rhizoclonium grande Børgesen 1935
114 Rhizoclonium riparium (Roth) Harvey 1849 Rhizoclonium kerneri Stock-mayer
115 Rhizoclonium riparium var. implexum (Dillwyn) Rosenvinge 1893
Rhizoclonium kochianum Kützing
116 Rhizoclonium setaceum Kützing 1847
117 Codiaceae Codium arabicum Kützing 1856
118 Codium bartlettii C. K. Tseng & W. J. Gilbert 1942
Puerto Galera Bay, Mindoro oriental, Philippines (Tseng and Gilbert 1942: 291-293).
119 Codium contractum Kjellman 1857
120 Codium cylindricum Holmes 1896
121 Codium decorticatum (Woodward) M. A. Howe 1911
Codium elongatum Aucto-rum
122 Codium edule P. C. Silva 1952
123 Codium fragile (Suringar) Hariot 1889
124 Codium geppiorum o. C. Schmidt 1923 Codium geppii o. C. Schmidt
125 Codium intricatum okamura 1913
126 Codium muelleri Kützing 1856
127 Codium ovale Zanardini 1878
128 Codium papillatum C. K. Tseng & W. J. Gilbert 1942
Puerto Galera, Mindoro oriental, Philippines (Tseng and Gilbert 1942: 293-295).
129 Codium platyclados R. Jones & Kraft 1984
130 Codium pungniforme okamura 1915
131 Codium tenue (Kützing) Kützing 1856
132 Codium tomentosum Stackhouse 1797
133 Dasycladaceae Bornetella nitida Munier-Chalmas ex Sonder 1880
134 Bornetella oligospora Solms-Laubach 1892
135 Bornetella sphaerica (Zanardini) Solms-Laubach 1892 Bornetella ovalis Yamada
136 Chloroclados australasicus Sonder 1871
Chloroclados philippinensis Gilbert; Chlorocladus australasicus Sonder; Chlorocladus philippinensis Gilbert
137 Cymopolia van-bosseae Solms-Laubach 1892
Cymopolia vanbosseae Solms-Laubach
138 Dasycladus vermicularis (Scopoli) Krasser 1898
139 Neomeris annulata Dickie 1874
140 Neomeris bilimbata Koster 1937
141 Neomeris vanbosseae M. A. Howe 1909
142 Derbesiaceae Derbesia attenuata Funk 1955
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143 Derbesia marina (Lyngbye) Solier 1846
144 Derbesia tenuissima (Moris & De Notaris) P. L. Crouan & H. M. Crouan 1867
145 Pedobesia ryukyuensis (Yamada & T. Tanaka) Kobara & Chihara 1984
146 Dichotomosiphona-ceae
Avrainvillea erecta (Berkeley) A. Gepp & e. S. Gepp 1911
Philippine Islands (Berkeley 1842: 157).
147 Avrainvillea lacerata Harvey ex J. Agardh 1887
148 Avrainvillea longicaulis (Kützing) G. Murray & Boodle 1889
149 Avrainvillea nigricans Decaisne 1842
150 Avrainvillea obscura (C. Agardh) J. Agardh 1887
Avrainvillea capituliformis Tanaka
151 Gayraliaceae Gayralia oxysperma (Kützing) K. L. Vinogradova ex Scagel 1989
Monostroma oxyspermum (Kützing) Doty
152 Gomontiaceae Monostroma latissimum Wittrock 1866
153 Monostroma nitidum Wittrock 1866 Porphyra crispata Kjellman
154 Halimedaceae Halimeda copiosa Goreau & e. A. Graham 1967
155 Halimeda cuneata Hering 1846
156 Halimeda cylindracea Decaisne 1842
157 Halimeda discoidea Decaisne 1842
158 Halimeda fragilis W. R. Taylor 1950
159 Halimeda gigas W. R. Taylor 1950
160 Halimeda goreauii W. R. Taylor 1962
161 Halimeda gracilis Harvey ex J. Agardh 1887
162 Halimeda incrassata (J. ellis) J. V. Lamouroux 1816
163 Halimeda lacunalis W. R. Taylor 1950
164 Halimeda macroloba Decaisne 1841
165 Halimeda macrophysa Askenasy 1888
166 Halimeda micronesica Yamada 1941 Halimeda orientalis Gilbert
167 Halimeda opuntia (Linnaeus) J. V. Lamouroux 1816
168 Halimeda renschii Hauck 1886 Halimeda batanensis W. R. Taylor
169 Halimeda simulans M. A. Howe 1907
170 Halimeda stuposa W. R. Taylor 1950
171 Halimeda taenicola W. R. Taylor 1950
172 Halimeda tuna (J. ellis & Solander) J. V. Lamouroux 1816
173 Halimeda velasquezii W. R. Taylor 1962 Sta. Ana, Cagayan, Luzon Island, Philippines (Taylor 1962: 177).
174 Halimeda velasquezii var. robusta P. A. Cordero 1974
Cadadalman, Camiguin Island, Cagayan, Philippines (Cordero 1974: 115).
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175 Ostreobiaceae Ostreobium quekettii Bornet & Flahault 1889
176 Phaeophilaceae Phaeophila dendroides (P. L. Crouan & H. M. Crouan) Batters 1902
177 Polyphysaceae Acetabularia caliculus J. V. Lamouroux 1824
178 Acetabularia crenulata J. V. Lamouroux 1816
179 Acetabularia dentata Solms-Laubach 1895
180 Acetabularia major G. Martens 1866
181 Acetabularia ryukyuensis okamura & Yamada 1932
Acetabularia philippinensis Gilbert Acetabularia roxasii G. C. Trono, Santiago & Ganzon-Fortes Acetabularia ryukyuensis var. philippinensis (Gilbert) Valet
182 Halicoryne wrightii Harvey 1860
183
Parvocaulis clavatus (Yamada) S. Berger, U. Fettweiss, S.Gleissberg, L. B. Liddle, U.Richter, H. Sawitzky & G. C.Zuccarello 2003
Acetabularia clavata Yamada
184
Parvocaulis parvulus (Solms-Laubach) S. Berger, U. Fettweiss, S. Gleissberg, L. B. Liddle, U. Richter, H. Sawitzky & G. C. Zuccarello 2003
Acetabularia minutissima okamura; Acetabularia moebii Solms-Laubach; Acetabularia parvula Solms-Laubach; Acetabularia velasquezii G. C. Trono, Santiago & Ganzon-Fortes
185
Parvocaulis polyphysoides (P. L. Crouan & H. M. Crouan) S. Berger, U. Fettweiss, S. Gleissberg, L. B. Liddle, U. Richter, H. Sawitzky & G.C. Zuccarello 2003
Acetabularia exigua Solms-Laubach
186 Rhipiliaceae Rhipilia crassa Millar & Kraft 2001
187 Rhipilia orientalis A. Gepp & e. S. Gepp 1911
188 Rhipilia tomentosa Kützing 1858
189 Rhipiliopsis carolyniae Kraft 1986 Bulusan, Sorsogon, Philippines (Kraft 1986: 51)
190 Siphonocladaceae Boergesenia forbesii (Harvey) Feldmann 1938
191 Dictyosphaeria cavernosa (Forsskål) Børgesen 1932
192 Dictyosphaeria cavernosa var. bullata Børgesen 1952
193 Dictyosphaeria intermedia Weber-van Bosse 1905
194 Dictyosphaeria ocellata (M. A. Howe) olsen-Stojkovich 1985
195 Dictyosphaeria versluysii Weber-van Bosse 1905
196 Ernodesmis verticillata (Kützing) Børgesen 1912
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197 Udoteaceae Boodleopsis pusilla (F. S. Collins) W. R. Taylor, A. B. Joly & Bernatowicz 1953
198 Boodleopsis verticillata e. Y. Dawson 1960
199 Chlorodesmis caespitosa J. Agardh 1887
200 Chlorodesmis fastigiata (C. Agardh) S. C. Ducker 1969
Chlorodesmis comosa Har-vey & Bailey
201 Chlorodesmis hildebrandtii A. Gepp & e. S. Gepp 1911
202 Chlorodesmis major Zanardini 1874
203 Rhipidosiphon javensis Montagne 1842 Udotea javensis (Montagne) A. Gepp & e. S. Gepp
204 Tydemania expeditionis Weber-van Bosse 1901
205 Udotea argentea Zanardini 1858
206 Udotea argentea var. spumosa A. Gepp & e. S. Gepp 1911
Syntype localities: Tanah Djampeah and Salayer Islands, Indonesia; Pearl Bank, Tawitawi, Sulu Archipelago, Philippines (Coppejans et al. 2001: 429)
207 Udotea flabellum (J. ellis & Solander) M. A. Howe 1904
208 Udotea geppiorum Yamada 1930 Udotea geppii Yamada
209 Udotea glaucescens Harvey ex J. Agardh 1887
210 Udotea indica A. Gepp & e. S. Gepp 1911
211 Udotea occidentalis A. Gepp & e. S. Gepp 1911
212 Udotea orientalis A. Gepp & e. S. Gepp 1911
Syntype localities: various in Indian and Pacific oceans; Indonesia; Philip-pine Islands (Gepp and Gepp 1911: 119-120).
213 Ulvaceae Ulva australis Areschoug 1854 Ulva pertusa Kjellman
214Ulva chaetomorphoides (Børgesen) Hayden, Blomster, Maggs, P. C. Silva, M. J. Stanhope & J. R. Waaland 2003
Enteromorpha chaetomor-phoides Børgesen
215 Ulva clathrata (Roth) C. Agardh 1811
Enteromorpha clathrata (Roth) Greville; Enteromorpha ramulosa (Smith) Carmichael
216 Ulva compressa Linnaeus 1753 Enteromorpha compressa (Linnaeus) Nees
217 Ulva flexuosa Wulfen 1803
Enteromorpha flexuosa (Wulfen) J. Agardh; Enteromorpha lingulata J. Agardh; Enteromorpha tubulosa (Kützing) Kützing
218 Ulva flexuosa f. submarina (F. S. Collins & Hervey) M. J. Wynne 2005
Enteromorpha flexuosa f. submarina Collins & Hervey
219 Ulva flexuosa subsp. paradoxa (C.Agardh) M. J. Wynne 2005
Enteromorpha flexuosa subsp. paradoxa (C. Agardh) Bliding
220 Ulva intestinalis Linnaeus 1753 Enteromorpha intestinalis (Linnaeus) Nees
221Ulva kylinii (Bliding) Hayden, Blomster, Maggs, P. C. Silva, M. J. Stanhope & J. R.Waaland 2003
Enteromorpha kylinii Bliding
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222 Ulva lactuca Linnaeus 1753 Ulva fasciata Delile
223 Ulva prolifera o. F. Müller 1778 Enteromorpha prolifera (o. F. Müller) J. Agardh
224 Ulva reticulata Forsskål 1775
225 Ulva rigida C. Agardh 1823
226 Valoniaceae Valonia aegagropila C. Agardh 1823
227 Valonia fastigiata Harvey ex J. Agardh 1887
228 Valonia macrophysa Kützing 1843
229 Valonia utricularis (Roth) C. Agardh 1823
230 Valonia ventricosa J. Agardh 1887 Ventricaria ventricosa (J. Agardh) olsen & J. A. West
231 Valoniopsis pachynema (G. Martens) Børgesen 1934
Valonia confervoides Za-nardini
Invalid Species Name
Family Taxon Recorded Remarks Reference
Ulvaceae Enteromorpha aragoensis Bliding 1960, nom. illeg.
*Enteromopha has now been included under Ulva. This species is currently consid-ered illegitimate because no type material was designated.
Silva et al. 1987;Silva 1996-to date
Species with No Matching Record in AlgaeBase
Family Taxon Recorded Remarks Reference
Caulerpaceae Caulerpa cupressoides var. cupressoides Not listed also in INA Database
Belleza and Liao 2007
Caulerpa cupressoides var. lycopodium f. disticha Weber-van Bosse
Same as Caulerpa cupres-soides f. disticha Weber-van Bosse 1898
Belleza and Liao 2007
Caulerpa mexicana var. mexicana Not listed also in INA Database Belleza and Liao 2007
Caulerpa racemosa var. racemosa (Forsskål) J. Agardh
Listed in INA Database as Caulerpa racemosa var. racemosa J. Agardh
Caulerpa racemosa var. serrulata (Fors-skål) J. Agardh
Not listed also in INA Database
Caulerpa serrulata var. serrulata f. lata (Weber-van Bosse) C. K. Tseng
Same as Caulerpa serrulata f. lata (Weber-van Bosse) C. K. Tseng
Belleza and Liao 2007
Caulerpa sertularioides var. sertularioides f. brevipes (J. Agardh) Svedelius
Same as Caulerpa sertulari-oides f. brevipes (J. Agardh) Svedelius
Belleza and Liao 2007
Cladophoraceae Cladophora filipendula No authority of taxon listed;Not listed also in INA Database
Ganzon-Fortes 2012
Species Listed as Preliminary Entry in AlgaeBase
Family Taxon Recorded Remarks Reference
Caulerpaceae Caulerpa sertularioides f. flagellata (Weber-van Bosse) Weber-van Bosse
Listed in INA Database Silva et al. 1987
Cladophoraceae Cladophora luzoniensis V. Martens Listed in INA Database; Type locality: San Mateo River, Luzon, Philippines (Martens 1866: 20)
Guiry and Guiry 2013;Silva 1996-to date
Halimedaceae Halimeda bikinensis W. R. Taylor Listed in INA Database Ganzon-Fortes 2012
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Table 2. Listing of brown (ochrophyta, Phaeophyceae) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed.
List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality
1 Acinetosporaceae Asteronema breviarticulatum (J. Agardh) ouriques & Bouzon 2000
Hincksia breviarticulata (J. Agardh) P. C. Silva
2 Ectocarpus rallsiae Vickers 1905 Hincksia rallsiae (Vickers) P. C. Silva
3 Feldmannia columellaris (Børgesen) Islam 1976
4 Feldmannia formosana (Yamada) Itono 1973
5 Feldmannia indica (Sonder) Womers-ley & A. Bailey 1970
6 Feldmannia irregularis (Kützing) G. Hamel 1939
7 Feldmannia mitchelliae (Harvey) H. S. Kim 2010
Hincksia mitchelliae (Harvey) P. C. Silva
8 Chordariaceae Leathesia marina (Lyngbye) Decaisne 1842 Leathesia difformis Areschoug
9 Cutleriaceae Mutimo cylindricus (okamura) H. Kawai & T. Kitayama 2012 Cutleria cylindrica okamura
10 Dictyotaceae Canistrocarpus cervicornis (Kützing) De Paula & De Clerck 2006
Dictyota cervicornis Kützing; Dictyota indica Sonder ex Kützing
11 Dictyopteris camiguinensis Tanaka 1967
San Pio Quinto, Camiguin Island, Ba-buyan Islands, Cagayan, Luzon, The Philippines (Tanaka 1967: 16)
12 Dictyopteris delicatula J. V. Lamour-oux 1809
13 Dictyopteris divaricata (okamura) okamura 1932
14 Dictyopteris jamaicensis W. R. Taylor 1960
15 Dictyopteris pacifica (Yendo) I. K. Hwang, H. S. Kim & W. J. Lee 2004 Spatoglossum pacificum Yendo
16 Dictyopteris polypodioides (A. P. De Candolle) J. V. Lamouroux 1809
17 Dictyopteris repens (okamura) Børge-sen 1924
18 Dictyopteris undulata Holmes 1896
19 Dictyota bartayresiana J. V. Lamour-oux 1809 Dictyota patens J. Agardh
20 Dictyota ceylanica Kützing 1859
21 Dictyota ciliolata Sonder ex Kützing 1859
22 Dictyota crenulata J. Agardh 1847 Dictyota bartayresiana var. denticulata Kützing
23 Dictyota crispata J. V. Lamouroux 1809 Dictyota bidentata Harvey & Bailey
24 Dictyota dichotoma (Hudson) J. V. Lamouroux 1809
25 Dictyota friabilis Setchell 1926
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26 Dictyota implexa (Desfontaines) J. V. Lamouroux 1809
Dictyota divaricata J. V. Lamouroux Dictyota linearis (C. Agardh) Greville
27 Dictyota lata J. V. Lamouroux 1809
28 Dictyota major W. R. Taylor 1945
29 Dictyota mertensii (Martius) Kützing 1859 Dictyota dentata J. V. Lamouroux
30Lobophora variegata (J. V. Lamou-roux) Womersley ex e. C. oliveira 1977
Pocockiella variegata (J. V. Lamou-roux) Papenfuss
31 Padina antillarum (Kützing) Piccone 1886
32 Padina arborescens Holmes 1896
33 Padina australis Hauck 1887
34 Padina boryana Thivy ex W. R. Taylor 1966
35 Padina distromatica Hauck 1887
36 Padina fernandeziana Skottsberg & Levring 1941
37 Padina fraseri (Greville) Greville 1830
38 Padina gymnospora (Kützing) Sonder 1871 Padina crassa Yamada
39 Padina japonica Yamada 1931
40 Padina jonesii Tsuda 1972
41 Padina minor Yamada 1925
42 Padina moffittiana I. A. Abbott & Huisman 2003
43 Padina pavonica (Linnaeus) Thivy 1960
44 Padina sanctae-crucis Børgesen 1914
45 Padina tetrastromatica Hauck 1887
46Rugulopteryx okamurae (e. Y. Daw-son) I. K. Hwang, W. J. Lee & H. S. Kim 2009
Dilophus okamurae e. Y. Dawson
47 Spatoglossum asperum J. Agardh 1894
48 Spatoglossum variabile Figari & De Notaris 1853
49 Stypopodium flabelliforme Weber-van Bosse 1913
50 Stypopodium zonale (J. V. Lamouroux) Papenfuss 1940
Stypopodium lobatum (C. Agardh) Kützing
51 Zonaria diesingiana J. Agardh 1841
52 Zonaria flabellata (okamura) Papen-fuss 1944
53 LaminariaceaeSaccharina latissima (Linnaeus) C. e. Lane, C. Mayes, Druehl & G. W. Saunders 2006
Ulva latissima Linnaeus
54 Mesosporaceae Mesospora negrosensis J. A. West & H. P. Calumpong 1996
Lalaan, San Jose, Negros oriental, Phil-ippines (West and Calumpong 1996: 5).
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55 Ralfsiaceae Ralfsia fungiformis (Gunnerus) Setch-ell & N. L. Gardner 1924
56 Ralfsia verrucosa (Areschoug) Are-schoug 1845
57 Sargassaceae Hormophysa cuneiformis (J. F. Gme-lin) P. C. Silva 1987
Hormophysa triquetra (C.Agardh) Kützing
58 Myagropsis myagroides (Mertens ex Turner) Fensholt 1955
59 Sargassum abbottiae G. C. Trono 1994 Pulong Bakaw, Calatagan, Batangas, Philippines (Trono 1994: 5).
60 Sargassum agardhianum Farlow 1889
61 Sargassum angii L. M. Liao 1998 Sargassum yamadae G. C. Trono Barrio Pangil, Currimao, Ilocos Norte, Philippines (Trono 1994: 4).
62 Sargassum aquifolium (Turner) C. Agardh 1820
Sargassum binderi Sonder ex J. Agardh; Sargassum biserrula J.Agardh; Sargassum crassifolium J. Agardh; Sargassum heterocystum (Kützing) Montagne; Sargassum oocyste J. Agardh
63 Sargassum baccularia (Mertens) C. Agardh 1824
64 Sargassum balingasayense G. C. Trono 1994
Barrio Balingasay, Bolinao, Pangas-inan, Philippines (Trono 1994: 5).
65 Sargassum bataanense G. C. Trono 1994
Pulong Bato, orion, Bataan, Philippines (Trono 1994: 6).
66 Sargassum belangeri Bory de Saint-Vincent 1834
Syntype localities: Sunda Strait and Java, Indonesia; Philippine Islands (Silva et al. 1996: 659).
67 Sargassum cervicorne Greville 1849 Sargassum binderi var. angustifolium Sonder
68 Sargassum cinctum J. Agardh 1848
69 Sargassum cinctum var. gracilentum Grunow 1915
Sargassum cinctum var. gracilenta Grunow
Syntype localities: Coast of Australia: Goodie Isl., Balina, Rockinham; near Manila, Philippines (Grunow 1915: 419).
70 Sargassum cinctum var. mixtum Grunow 1915 Sargassum cinctum var. mixta Grunow Near Manila, Luzon, Philippines (Grun-
ow 1915: 419).
71 Sargassum confusum C. Agardh 1824
72 Sargassum corderoi R. B. Modelo Jr., I. Umezaki & L. M. Liao 1998
Polo, New Washington, Aklan, Panay Is., Philippines (Modelo et al. 1998: 79).
73 Sargassum crispifolium Yamada 1931
74 Sargassum currimaoense G. C. Trono 1994
Pangil, Currimao, Ilocos Norte, Philip-pines (Trono 1994: 6).
75 Sargassum cystocarpum C. Agardh 1820
76 Sargassum cystophyllum Montagne 1842
Manila, Luzon, Philippines (Montagne 1842: 249).
77 Sargassum distichum Sonder 1845 Sargassum aemulum Sonder
78 Sargassum dotyi G. C. Trono 1994 Honduras, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 3).
79Sargassum droserifolium var. spathu-latum Grunow 1915
Sargassum droserifolium var. spathu-lata Grunow
Strait Bernardino, Philippines (Grunow 1915: 411).
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80 Sargassum duplicatum var. rotunda-tum Grunow 1915
Sargassum duplicatum var. rotundata Grunow
Syntype localities: Philippine Islands; Java, Indonesia; Seychelles (Grunow 1915: 392-393).
81 Sargassum esperi C. Agardh 1820
82 Sargassum feldmannii Pham-Hoang Ho 1967
83 Sargassum filifolium (C. Agardh) C. Agardh 1824
84 Sargassum filiforme Montagne 1844 "in insulis Philippinensibus” (Montagne 1844: 660) (Silva et al. 1996: 672).
85 Sargassum fluitans (Børgesen) Børge-sen 1914
86 Sargassum fulvellum (Turner) C. Agardh 1820
87 Sargassum furcatum Kützing 1843
88 Sargassum gaudichaudii Montagne 1842
Syntype localities: Mauritius; Philip-pine Islands (Montagne 1842: 249) (Silva et al. 1996: 673).
89 Sargassum giganteifolium Yamada 1925
90 Sargassum gracile J. Agardh 1848
91 Sargassum gracillimum Reinbold 1913
92 Sargassum granuliferum C. Agardh 1820
93 Sargassum hemiphyllum (Turner) C. Agardh 1820
94 Sargassum hombronianum var. ma-nilense (Grunow) Grunow 1916
Sargassum polycystum var. manilense Grunow
Cavite, Luzon, Philippines (Grunow 1916: 8)
95 Sargassum horneri (Turner) C. Agardh 1820 Sargassum filicinum Harvey
96 Sargassum ilicifolium (Turner) C. Agardh 1820
Sargassum berberifolium J. Agardh; Sargassum cristaefolium C. Agardh; Sargassum droserifolium Bory de Saint-Vincent; Sargassum duplicatum Bory de Saint-Vincent; Sargassum duplicatum (J. Agardh) J. Agardh; Sargassum sandei Reinbold; Sargassum turbinatifolium C.K.Tseng & Lu
97 Sargassum ilicifolium f. benkulense Grunow 1915
Sargassum ilicifolium f. benkulensis Grunow
Syntype localities: Pulau Tikus, near Bengkulu, Sumatra, Indonesia; Philip-pine Islands (Grunow 1915: 407).
98 Sargassum ilicifolium var. compactum (Bory de Saint-Vincent) Grunow 1915
99 Sargassum ilicifolium var. pseudospi-nulosum Grunow 1915
100 Sargassum kushimotense Yendo 1907
101 Sargassum latifolium (Turner) C. Agardh 1820
102 Sargassum microcystum J. Agardh 1848Syntype localities: "in marfi Indico"; Moluccas, Indonesia; Manila, Philip-pines; Singapore (Silva et al. 1996: 685).
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103 Sargassum microcystum f. luzonense Grunow 1915
Sargassum microcystum f. luzonensis Grunow
Near Manila, Luzon, Philippines (Grun-ow 1915: 413)
104 Sargassum microcystum var. euryphyl-lum (Grunow) Grunow 1915
Sargassum myriocystum var. eury-phylla (Grunow) Grunow; Sargassum myriocystum var. euryphyl-lum (Grunow) Grunow; Sargassum polycystum var. euryphyl-lum Grunow
Ticao Island, Masbate, Philippines (Grunow 1915: 440)
105 Sargassum microcystum var. microtis Grunow 1915
Syntype localities: Java, Indonesia; Basilan Strait, Philippines (Grunow 1915: 414).
106 Sargassum microphyllum C. Agardh 1820
107 Sargassum miyabei Yendo 1907
108 Sargassum natans (Linnaeus) Gaillon 1828
109 Sargassum nigrifolium Yendo 1907
110 Sargassum notarisii Zanardini 1858
111 Sargassum ohnoi G. C. Trono 1994 Big Balatero, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 7).
112 Sargassum oligocystum Montagne 1845
113 Sargassum oligocystum var. bernar-dium Grunow 1915
"in freto Bernardino" (San Bernardino Strait between Sorsogon, Luzon, and Northern Samar, Philippines) (Grunow 1915: 386)
114 Sargassum oligocystum var. subflexuo-sum Grunow 1915
Sargassum oligocystum var. subflex-uosa Grunow
“Fretum Bernardium, Macassar” (San Bernardino Strait, Philippines; Makas-sar, Indonesia) (Grunow 1915: 385)
115 Sargassum oocyste var. bernardinum Grunow 1915
Sargassum oocyste var. bernardina Grunow 1915
"in freto Bernardino" (San Bernardino Strait, Philippines)(Grunow 1915: 437).
116 Sargassum paniculatum J. Agardh 1848
117 Sargassum parvifolium (Turner) C. Agardh 1820
118 Sargassum patens C. Agardh 1820
119 Sargassum patens var. schizophyllum (Kützing) Yendo 1905
120 Sargassum philippinense Grunow 1916
San Bernardino Strait, Philippines (Grunow 1916: 166)
121 Sargassum piluliferum (Turner) C. Agardh 1820
122 Sargassum polyceratium Montagne 1837
123 Sargassum polycystum C. Agardh 1824 Sargassum myriocystum J. Agardh
124 Sargassum pteropleuron Grunow 1868 Sargassum pteropleron Grunow
125 Sargassum pulchellum Grunow 1888
126 Sargassum sagamianum Yendo 1907
127 Sargassum samarense G. C. Trono 1994
Borangan, eastern Samar, Philippines (Trono 1994: 4)
128 Sargassum serratifolium (C. Agardh) C. Agardh 1820
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129 Sargassum siliquosum J. Agardh 1848 Sargassum siloquosum J. Agardh
130 Sargassum siliquosum var. basilani-cum Grunow 1916
Sargassum siliquosum var. basilanica Grunow
Basilan Strait (between Mindanao and Basilan Island) (Grunow 1916: 175)
131 Sargassum siliquosum var. bicornutum Grunow 1916
Sargassum siliquosum var. bicornuta Grunow
China Sea; Ticao Island, Masbate, Phil-ippines (Grunow 1916: 175)
132 Sargassum siliquosum var. manipaense Grunow 1916
Sargassum siliquosum var. mani-paensis Grunow;Spelled in AlgaeBase as Sargassum siliquosum var. manipense Grunow
"ad insulas Philippinas, Manipa" (Grun-ow 1916: 175)
133 Sargassum spinifex C. Agardh 1820
134 Sargassum subspathulatum (Grunow) Grunow 1916
135 Sargassum sullivanii G. C. Trono 1994 Palawig, Zambales, Philippines (Trono 1994: 6)
136 Sargassum tenerrimum J. Agardh 1848
137 Sargassum tristichum Sonder 1845 Sargassum biforme Sonder
138 Sargassum turbinarioides Grunow 1915
139 Sargassum umezakii G. C. Trono 1994 Medio Island, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 7)
140 Sargassum velasquezii G. C. Trono 1994
San Isidro, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 4)
141 Sargassum vulgare C. Agardh 1820
142 Sargassum vulgare var. indicum C. Agardh 1820
143 Sargassum yendoi okamura & Ya-mada 1938
144 Sargassum yoshidae G. C. Trono 1994 Cabugao Bay, Virac, Catanduanes, Philippines (Trono 1994: 5)
145 Sirophysalis trinodis (Forsskål) Kütz-ing 1849
Cystoseira trinodis (Forsskål) C.Agardh
146Stephanocystis hakodatensis (Yendo) Draisma, Ballesteros, F. Rousseau & T. Thibaut 2010
Cystoseira hakodatensis (Yendo) Fensholt
147 Turbinaria condensata Sonder 1860
148 Turbinaria conoides (J. Agardh) Kütz-ing 1860
149 Turbinaria conoides f. laticuspidata W. R. Taylor 1964 Cebu, Philippines (Taylor 1964: 481)
150 Turbinaria conoides f. retroflexa W. R. Taylor 1964
Tandayong Island, Luzon, Philippines (Taylor 1964: 481-482)
151 Turbinaria decurrens Bory de Saint-Vincent 1828
152 Turbinaria denudata Bory de Saint-Vincent 1828
153 Turbinaria filamentosa Yamada 1925
154 Turbinaria luzonensis W. R. Taylor 1964
Mulanay, Luzon, Philippines (Taylor 1964: 482).
155 Turbinaria murrayana e. S. Barton 1891
156 Turbinaria ornata (Turner) J. Agardh 1848
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157 Turbinaria ornata f. evesiculosa (e. S. Barton) W. R. Taylor 1964
158 Turbinaria trialata var. capensis Kütz-ing 1860
159 Turbinaria turbinata (Linnaeus) Kuntze 1898 Turbinaria trialata (J. Agardh) Kützing
160 Turbinaria vulgaris J. Agardh 1848 nom. illeg.
161 Scytosiphonaceae Chnoospora implexa J. Agardh 1848
162 Chnoospora minima (Hering) Papen-fuss 1956
163 Chnoospora pannosa J. Agardh 1848
164 Colpomenia sinuosa (Mertens ex Roth) Derbes & Solier 1851
165 Hydroclathrus clathratus (C. Agardh) M. A. Howe 1920
166 Hydroclathrus tenuis C. K. Tseng & Lu 1983
167 Rosenvingea intricata (J. Agardh) Børgesen 1914
168 Rosenvingea orientalis (J. Agardh) Børgesen 1914
Hab. in mari Indico ad Manillam (Hb. Binder!) [Manila, the Philippines] (Agardh 1848: 78).
169 Sphacelariaceae Sphacelaria novae-hollandiae G. Sonder 1845
170 Sphacelaria rigidula Kützing 1843
171 Sphacelaria tribuloides Meneghini 1840
Species with No Matching Record in AlgaeBase
Family Taxon Recorded Remarks Reference
Sphacelariaceae Sphacelaria mucifera No authority of taxon listed; Also not listed in INA Database Ganzon-Fortes 2012
Sargassaceae Sargassum cinctum var. jagorii Grunow
Type locality: Luzon, Philippines (Grunow 1915: 421)
Grunow 1915; Silva 1996-to date
Sargassum crassifolium var. subdupli-catum ('subduplicata') Grunow
Syntype localities: Makassar, Indo-nesia; Strait Bernardino, Philippines (Grunow 1915: 421)
Grunow 1915; Silva 1996-to date
Species Listed as Preliminary Entry in AlgaeBase
Family Taxon Recorded Remarks Reference
Sargassaceae Sargassum elongatum No authority of taxon listed; Not listed in INA Database
Phillips 1995; Ganzon-Fortes 2012
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Table 3. Listing of red (Rhodophyta) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed.
List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality
1 Acrochaetiaceae Acrochaetium hancockii (e. Y. Daw-son) Papenfuss 1845
2 Acrochaetium liagorae Børgesen 1915
3 Acrochaetium nitidulum I. A. Abbott 1962
Balabac Island, Gnat Reef, Pala-wan, Philippines (Abbott 1962: 95)
4 Acrochaetium papenfussii I. A. Abbott 1962
Cagayan Sulu Island, Palawan, Philippines (Abbott 1962: 97)
5 Acrochaetium robustum Børgesen 1915
6 Acrochaetium sinicola (e. Y. Dawson) Papenfuss 1945
7 Acrochaetium trichogloeae Børgesen 1952
8 Acrochaetium tuticorinense Børgesen 1937
9Colaconema gracile (Børgesen) Ateweberhan & Prud'homme van Reine 2005
Acrochaetium gracile Børgesen
10Colaconema hallandicum (Kylin) Afonso-Carillo, Sanson, Sangil & Diaz-Villa 2007
Acrochaetium hallandicum (Kylin) Hamel
11 Colaconema hypneae (Børgesen) A. A. Santos & C. W. N. Moura 2010 Acrochaetium seriatum Børgesen
12 Ahnfeltiaceae Ahnfeltiopsis concinna (J.Agardh) P. C. Silva & DeCew 1992 Ahnfeltia concinna J. Agardh
13 Bangiaceae Bangia atropurpurea (Mertens ex Roth) C. Agardh 1824
14 Bangia yamadae Tanaka 1944
15 Porphyra atropurpurea (olivi) De Toni 1897
16 Porphyra marcosii P. A. Cordero 1976
Dirique Bay, Burgos, Ilocos Norte, Philippines (Cordero 1976: 15)
17 Porphyra umbilicalis Kützing 1843 Ulva umbilicalis Linneaus
18 Pyropia denticulata (Levring) J. A. Philips & J. e. Sutherland 2011
Porphyra denticulata Levring
19Pyropia suborbiculata (Kjellman) J.e.Sutherland, H. G. Choi, M. S. Hwang & W. A. Nelson 2011
Porphyra suborbiculata Kjellman
20Pyropia vietnamensis (Tak. Tanaka & P.H. Ho) J. e. Sutherland & Monotilla 2011
Porphyra vietnamensis T. Tanaka & P. H. Ho
21 Wildemania variegata (Kjellman) De Toni 1890
Porphyra variegata (Kjellman) Kjellman
22 Bonnemaisoniaceae Asparagopsis taxiformis (Delile) Trevisan de Saint-Leon 1845
Falkenbergia hillebrandii (Bornet) Falkenberg
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23 Callithamniaceae Crouania attenuata (C. Agardh) J. Agardh 1842 Crouania minutissima Yamada
24 Caulacanthaceae Catenella caespitosa (Withering) L. M. Irvine 1976
25 Catenella impudica (Montagne) J. Agardh 1852
26 Catenella nipae Zanardini 1872
27 Caulacanthus ustulatus (Mertens ex Turner) Kützing 1843
28 Ceramiaceae Acrothamnion preissii (Sonder) e. M. Wollaston 1968
29 Antithamnion antillanum Børgesen 1917
Antithamnion lherminieri (P. L. Crouan & H. M. Crouan) Bornet ex Nasr
30 Balliella subcorticata (Itono) Itono & T. Tanaka 1973
31 Centroceras clavulatum (C. Agardh) Montagne 1846
32 Centroceras minutum Yamada 1944
33 Ceramium affine Setchell & N. L. Gardner 1930
34 Ceramium cimbricum H. e. Petersen 1924 Ceramium fastigiatum Harvey
35 Ceramium cruciatum F. S. Collins & Hervey 1917
36 Ceramium diaphanum (Lightfoot) Roth 1806
Ceramium gracillimum (Kützing) Griffiths & Harvey; Ceramium tenuissimum (Roth) Areschoug
37 Ceramium equisetoides e. Y. Dawson 1944
38 Ceramium loureiri C. Agardh 1824
39 Ceramium luetzelburgii o. C. Schmidt 1924
40 Ceramium marshallense e. Y. Daw-son 1957
41 Ceramium maryae Weber-van Bosse 1923
42 Ceramium mazatlanense e. Y. Daw-son 1950
43 Ceramium multijugum Jaasund 1970
44 Ceramium nitens C. Agardh (J. Agardh) 1851
45 Ceramium personatum Setchell & N. L. Gardner 1930
46 Ceramium procumbens Setchell & N. L. Gardner 1924
Ceramiella procumbens (Setchell & N. L. Gardner) Diaz-Piferrer
47 Ceramium serpens Setchell & N. L. Gardner 1924
48 Ceramium sinicola Setchell & N. L. Gardner 1924
49 Ceramium tenerrimum (G. Martens) okamura 1921
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50 Ceramium vagans P. C. Silva 1987
51 Ceramium zacae Setchell & N. L. Gardner 1937
52 Corallophila kleiwegii Weber-van Bosse 1923 Centroceras apiculatum Yamada
53 Gayliella flaccida (Harvey ex Kütz-ing) T. o. Cho & L. J. McIvor 2008
Ceramium flaccidum (Harvey ex Kützing) Ardissone
54 Gayliella taylorii (e. Y. Dawson) T. o. Cho & S. M. Boo 2008 Ceramium taylorii e. Y. Dawson
55 Herpochondria elegans (okamura) Itono 1977 Microcladia elegans okamura
56 Microcladia glandulosa (Solander ex Turner) Greville 1830
57 Champiaceae Champia bifida okamura 1901
58 Champia caespitosa e. Y. Dawson 1944
59 Champia compressa Harvey 1838
60 Champia disticha e. Y. Dawson 1944
61 Champia japonica okamura 1931
62 Champia parvula (C. Agardh) Harvey 1853
63 Champia salicornoides Harvey 1853
64 Champia spathulata Weber-van Bosse 1928
65 Champia vieillardii Kützing 1866
66 Coelothrix irregularis (Harvey) Børgesen 1920
67 Corallinaceae Amphiroa anastomosans Weber-van Bosse 1904
68 Amphiroa anceps (Lamarck) Decaisne 1842
69 Amphiroa annulata M. Lemoine 1929
70 Amphiroa beauvoisii J. V. Lamouroux 1816 Amphiroa zonata Yendo
71 Amphiroa crassa J. V. Lamouroux 1824
72 Amphiroa cumingii Montagne 1844
73 Amphiroa dimorpha M. Lemoine 1929
74 Amphiroa ephedraea (Lamarck) Decaisne 1842
75 Amphiroa foliacea J. V. Lamouroux 1824
76 Amphiroa foliacea f. erecta Weber- van Bosse 1904
77 Amphiroa fragilissima (Linnaeus) J. V. Lamouroux 1816
78Amphiroa fragilissima f. cyathifera (J. V. Lamouroux) Weber-van Bosse 1904
79 Amphiroa hancockii W. R. Taylor 1942
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80 Amphiroa pacifica Kützing 1858
81 Amphiroa rigida J. V. Lamouroux 1816
82 Amphiroa subcylindrica e. Y. Dawson 1953
83 Amphiroa tribulus (J. ellis & Solan-der) J. V. Lamouroux 1816
84 Amphiroa valonioides Yendo 1902
85 Arthrocardia palmata (J. ellis & Solander) Areschoug 1852
86 Bossiella frondescens (Postels & Ruprecht) e. Y. Dawson 1964
Corallina frondescens Postels & Ruprecht
87Hydrolithon farinosum (J. V. Lamour-oux) D. Penrose & Y. M. Chamberlain 1993
Fosliella farinosa (J. V. Lamouroux) M. A. Howe
88 Hydrolithon reinboldii (Weber-van Bosse & Foslie) Foslie 1909
89 Jania acutiloba (Decaisne) J. H. Kim, Guiry & H. G. Choi 2007
Cheilosporum jungermannioides Ruprecht ex Areschoug
90 Jania adhaerens J. V. Lamouroux 1816
91 Jania capillacea Harvey 1853
92 Jania cubensis Montagne ex Kützing 1849
Haliptilon cubense (Montagne ex Kützing) Garbary & H. W. Johansen
93 Jania cultrata (Harvey) J. H. Kim, Guiry & H. G. Choi 2007
Cheilosporum cultratum (Harvey) Areschoug
94 Jania decussato-dichotoma (Yendo) Yendo 1905
95 Jania longiarthra e. Y. Dawson 1953
96 Jania micrarthrodia J. V. Lamouroux 1816
97 Jania pacifica Areschoug 1852
98 Jania pumila J. V. Lamouroux 1816
99 Jania rubens (Linnaeus) J. V. Lamouroux 1816
100 Jania sagittata (J. V. Lamouroux) Blainville 1834
Cheilosporum sagittatum (J. V. Lamouroux) Areschoug
101 Jania spectabilis (Harvey ex Grunow) J. H. Kim, Guiry & H. G. Choi 2007
Cheilosporum spectabile Harvey ex Grunow
102 Jania tenella (Kützing) Grunow 1874
103 Jania tenella var. zacae e. Y. Dawson 1953
104 Jania ungulata (Yendo) Yendo 1905
105 Jania ungulata f. brevior (Yendo) Yendo 1905
106 Lithophyllum byssoides (Lamarck) Foslie 1900
107 Lithophyllum pallescens (Foslie) Foslie 1900
108 Lithophyllum pygmaeum (Heydrich) Heydrich 1897
Lithophyllum moluccense (Foslie) Foslie
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109 Lithoporella indica (Foslie) W. H. Adey 1970
110 Lithoporella melobesioides (Foslie) Foslie 1909
111 Mastophora rosea (C. Agardh) Setchell 1943
112 Mesophyllum simulans (Foslie) Me. Lemoine 1928
113 Neogoniolithon brassica-florida (Har-vey) Setchell & L. R. Mason 1943
Neogoniolithon frutescens (Foslie) Setchell & L. R. Mason
114Neogoniolithon megalocystum (Weber-van Bosse & Foslie) Setchell & L. R. Mason 1943
115 Porolithon onkodes (Heydrich) Foslie 1909
116 Corynocystaceae Corynocystis prostrata Kraft 1999 Bulusan, Sorsogon, Philippines (Kraft et al. 1999: 28).
117 Cubiculosporaceae Cubiculosporum koronicarpis Kraft 1973
Bulusan, Sorsogon, Philippines. (Kraft 1973: 881).
118 Cystocloniaceae Hypnea boergesenii T. Tanaka 1941
119 Hypnea caespitosa Geraldino & S. M. Boo 2010
Intertidal zone off Panglao Is-land, Bohol, Philippines (Ger-aldino et al. 2010: 338)
120 Hypnea cenomyce J. Agardh 1851
121 Hypnea charoides J. V. Lamouroux 1813
122 Hypnea cornuta (Kützing) J. Agardh 1851
123 Hypnea divaricata (C. Agardh) Greville 1830
124 Hypnea musciformis (Wulfen) J. V. Lamouroux 1813
125 Hypnea musciformis var. esperi J. Agardh 1851
126 Hypnea musciformis var. hippuroides (Kützing) Weber-van Bosse 1928
127 Hypnea nidulans Setchell 1924
128 Hypnea pannosa J. Agardh 1847
129 Hypnea saidana Holmes 1896
130 Hypnea spinella (C. Agardh) Kützing 1847 Hypnea cervicornis J. Agardh
131 Hypnea stellulifera (J. Agardh) Yama-gishi & M. Masuda 2003
Hypnea cornuta var. stellulifera J. Agardh
Lectotype locality: Manila, Phi-lippines (Yamagishi et al. 2003: 535).
132 Hypnea valentiae (Turner) Montagne 1841
133 Rhodophyllis peltata var. lacunosa Grunow 1874
134 Dasyaceae Dasya anastomosans (Weber-van Bosse) M. J. Wynne 2002
Dasya adhaerens Yamada; Eupogodon pilosus (Weber-van Bosse) P. C. Silva
135 Dasya antillarum (M. A. Howe) A.J.K .Millar 1996
Eupogodon antillarum (M. A. Howe) P. C. Silva
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136 Dasya baillouviana (S. G. Gmelin) Montagne 1841
137 Dasya caraibica Børgesen 1919
138 Dasya mollis Harvey 1853
139 Dasya ocellata (Grateloup) Harvey 1833
140 Dasya punicea (Zanardini) Meneghini ex Zanardini 1841
141 Dasya sessilis Yamada 1928
142 Dasya villosa Harvey 1844
143 Eupogodon planus (C. Agardh) Kütz-ing 1845
144 Heterosiphonia crispella (C. Agardh) M. J. Wynne 1985
Heterosiphonia crispella var. laxa (Børgesen) M. J. Wynne
145 Heterosiphonia muelleri (Sonder) De Toni 1903
146 Tapeinodasya bornetii Weber-van Bosse 1904
147 Delesseriaceae Apoglossum spathulatum (Sonder) Womersley & Shepley 1982
Hypoglossum spathulatum (Sonder) Kützing
148 Botryocarpa prolifera Greville 1830
149 Caloglossa bengalensis (G. Martens) R. J. King & Puttock 1994
Caloglossa adnata (Zanardini) De Toni
150 Caloglossa leprieurii (Montagne) G. Martens 1869
151 Caloglossa ogasawaraensis okamura 1897
152 Caloglossa vieillardii (Kützing) Setchell 1924
Caloglossa leprieurii var. hookeri Post
153 Claudea batanensis Tanaka 1967Basco, Batan Island, Batanes, Luzon, Philippines (Tanaka 1967: 18).
154 Claudea multifida Harvey 1854
155 Hypoglossum attenuatum N. L. Gardner 1927
156 Hypoglossum harveyanum (J. Agardh) Womersley & Shepley 1982 Hypoglossum serrulatum J. Agardh
157 Hypoglossum heterocystideum (J. Agardh) J. Agardh 1898
158 Martensia australis Harvey 1855 Martensia speciosa Zanardini
159 Martensia flabelliformis Harvey ex J. Agardh 1863
160 Martensia fragilis Harvey 1854
161 Martensia martensii (F. Schmitz) S. M. Lin, Fredericq & L. M. Liao 2001 Opephyllum martensii F. Schmitz
an der küste der Philippinen’ [Philippines] (Schmitz and Hauptfleisch 1897: 410).
162 Taenioma perpusillum (J. Agardh) J. Agardh 1863
163 Vanvoorstia spectabilis Harvey 1854
164 Zellera tawallina G. Martens 1866
165 Dicranemataceae Dicranema revolutum (C. Agardh) J. Agardh 1852
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166 Dumontiaceae Gibsmithia hawaiiensis Doty 1963
167 Rhodopeltis borealis Yamada 1931
168 Stenopeltis gracilis (Yamada & Tanaka) Itono & Yoshizaki 1992
Rhodopeltis gracilis Yamada & Tanaka
169 Endocladiaceae Gloiopeltis complanata (Harvey) Yamada 1932
170 Gloiopeltis tenax (Turner) Decaisne 1842
171 Erythropeltidaceae Erythrocladia irregularis Rosenvinge 1909
172 Erythrocladia pinnata W. R. Taylor 1942
173 Erythrotrichiaceae Erythrotrichia bangioides Levring 1955
174 Erythrotrichia biseriata Tanaka 1944
175 Erythrotrichia parietalis Tanaka 1952
176 Faucheaceae Gloiocladia ramellifera Hauck 1886
177
Leptofauchea leptophylla (Segawa) Mas. Suzuki, Nozaki, R. Terada, Kita-yama, Tetsu. Hashimoto & Yoshizaki 2012
Fauchea leptophylla Segawa
178 Galaxauraceae Actinotrichia fragilis (Forsskål) Børgesen 1932
179 Dichotomaria apiculata (Kjellman) A. Kurihara & Masuda 2005 Galaxaura apiculata Kjellman
180 Dichotomaria falcata (Kjellman) A. Kurihara & Masuda 2005 Galaxaura falcata Kjellman
181 Dichotomaria marginata (J. ellis & Solander) Lamarck 1816
Galaxaura marginata (J. ellis & Solander) J. V. Lamouroux
182 Dichotomaria obtusata (J. ellis & Solander) Lamarck 1816
Galaxaura obtusata (J. ellis & Solander) J. V. Lamouroux
183 Dichotomaria spathulata (Kjellman) A.Kurihara & Huisman 2006 Galaxaura arborea Kjellman
184 Galaxaura contigua Kjellman 1900
185 Galaxaura divaricata (Linnaeus) Huisman & R.A.Townsend 1993 Galaxaura fasciculata Kjellman
186 Galaxaura filamentosa R. C. Y. Chou 1945
187 Galaxaura kjellmanii Weber-van Bosse 1921
188 Galaxaura rugosa (J. ellis & Solan-der) J. V. Lamouroux 1816
Galaxaura subfruticulosa R. C. Y. Chou; Galaxaura subverticillata Kjellman
189 Galaxaura striata Kjellman 1900
190Tricleocarpa cylindrica (J. ellis & Solander) Huisman & Borowitzka 1990
Galaxaura fastigiata DecaisneLectotype (Galaxaura fastigata Decaisne): Manila, Luzon, Phil-ippines (Huisman 2006: 25).
191 Tricleocarpa fragilis (Linnaeus) Huis-man & R. A. Townsend 1993
Galaxaura oblongata (J. ellis & Solander) J. V. Lamouroux
192 Gelidiaceae Gelidium amansii (J. V. Lamouroux) J. V. Lamouroux 1813
193 Gelidium capense (S. G. Gmelin) P. C. Silva 1987
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194 Gelidium coulteri Harvey 1853
195 Gelidium crinale (Hare ex Turner) Gaillon 1828
196 Gelidium crinale var. perpusillum Piccone & Grunow 1884
197 Gelidium divaricatum G. Martens 1866
198 Gelidium isabelae W. R. Taylor 1945
199 Gelidium kintaroi Yamada 1941
200 Gelidium pulchellum (Turner) Kütz-ing 1868
201 Gelidium pusillum (Stackhouse) Le Jolis 1863
202 Gelidium pusillum var. pacificumW. R. Taylor 1945
203 Gelidium rigens (C. Agardh) Greville ex Kützing 1849
204 Ptilophora scalaramosa (Kraft) R. e. Norris 1987 Beckerella scalaramosa Kraft Bulusan, Sorsogon, Luzon, Phil-
ippines (Kraft 1976: 85).
205 Gelidiellaceae Gelidiella acerosa (Forsskål) Feld-mann & G. Hamel 1934
206 Parviphycus adnatus (e. Y. Dawson) B. Santelices 2004 Gelidiella adnata e. Y. Dawson
207 Gigartinaceae Chondracanthus tenellus (Harvey) Hommersand 1993 Gigartina tenella Harvey
208 Gracilariaceae Gracilaria arcuata Zanardini 1858
209 Gracilaria arcuata var. snackeyi Weber-van Bosse 1928
210 Gracilaria blodgettii Harvey 1853
211 Gracilaria bursa-pastoris (S. G. Gmelin) P. C. Silva 1952
Gracilaria compressa (C.Agardh) Greville
212 Gracilaria canaliculata Sonder 1871 Gracilaria crassa Harvey ex J.Agardh
213Gracilaria changii (B. M. Xia & I. A. Abbott) I. A. Abbott, J. Zhang & B. M. Xia 1991
214 Gracilaria coronopifolia J. Agardh 1852
215 Gracilaria cylindrica Børgesen 1920
216 Gracilaria damaecornis J. Agardh 1852
217 Gracilaria debilis (Forsskål) Børge-sen 1932
Polycavernosa debilis (Forsskål) Fredericq & J. N. Norris
218 Gracilaria disticha (J. Agardh) J. Agardh 1852
219 Gracilaria firma C. F. Chang & B. M. Xia 1976
220 Gracilaria foliifera (Forsskål) Børge-sen 1932
221 Gracilaria foliifera f. aeruginosa Børgesen 1938
222 Gracilaria gigas Harvey 1860
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223 Gracilaria incurvata okamura 1931
224 Gracilaria isabellana Gurgel, Fred-ericq & J. N. Norris 2004
Gracilaria lacinulata (M. Vahl) M. A. Howe
225 Gracilaria manilaensis Yamamoto & G. C. Trono 1994
Hali Beach, Parañaque, Manila Bay, Philippines (Yamamoto and Trono 1994: 95).
226 Gracilaria minor (Sonder) Durairat-nam 1961
227 Gracilaria papenfussii I. A. Abbott 1983
228 Gracilaria punctata (okamura) Yamada 1941
229 Gracilaria salicornia (C. Agardh) e. Y. Dawson 1954
Manila, Luzon, Philippines (Dawson 1954: 4; Millar and Xia 1997; see also Ganzon-Fortes 2012).
230 Gracilaria spinigera e. Y. Dawson 1949
231 Gracilaria spinulosa (okamura) C. F. Chang & B. M. Xia 1976
232Gracilaria srilankia (C. F. Chang & B. M. Xia) Withell, A. J. K. Millar & Kraft 1994
233 Gracilaria sullivanii Yamamoto & G. C. Trono 1994
Matabungkay, Batangas, Luzon Island, Philippines (Yamamoto and Trono 1994: 99).
234 Gracilaria tenuistipitata C. F. Chang & B. M. Xia 1976
235 Gracilaria tenuistipitata var. liui J. Zhang & B. M. Xia 1988
236 Gracilaria textorii (Suringar) De Toni 1895
237 Gracilaria turgida e. Y. Dawson 1949
238 Gracilaria vanbosseae (I. A. Abbott) I. A. Abbott 1991
Polycavernosa vanbosseae I. A. Abbott
239 Gracilaria venezuelensis W. R. Taylor 1942
240 Gracilaria vieillardii P. C. Silva 1987
241 Gracilariopsis bailiniae J. Zhang & B.M.Xia 1991
Gracilaria heteroclada J. Zhang & B. M. Xia 1988 nom. Illeg.
242 Gracilariopsis heteroclada J. Zhang & B. M. Xia 1991
243Gracilariopsis lemaneiformis (Bory de Saint-Vincent) e. Y. Dawson, Acleto & Foldvik 1964
Gracilaria lemaneiformis (Bory de Saint-Vincent) Greville
244Gracilariopsis longissima (S. G. Gmelin) M. Steentoft, L. M. Irvine & W. F. Farnham 1995
Gracilaria verrucosa (Hudson) Papenfuss
245 Hydropuntia edulis (S. G. Gmelin) Gurgel & Fredericq 2004
Gracilaria edulis (S. G. Gmelin) P. C. Silva; Gracilaria lichenoides Greville; Polycavernosa fastigiata C.F.Chang & B.M.Xia
246 Hydropuntia eucheumatoides (Har-vey) Gurgel & Fredericq 2004
Gracilaria eucheumatoides Harvey; Gracilaria eucheumoides Harvey
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247 Hydropuntia urvillei Montagne 1842
248 Halymeniaceae Carpopeltis capitellata (Sonder) F. Schmitz 1897
249 Carpopeltis maillardii (Montagne & Millardet) Chiang 1970
250 Cryptonemia crenulata (J. Agardh) J. Agardh 1851
251 Cryptonemia denticulata J. Agardh 1851
252 Cryptonemia seminervis (C. Agardh) J. Agardh 1846
Cryptonemia luxurians (C. Agardh) J. Agardh
253 Grateloupia acuminata Holmes 1896 Halymenia acuminata (Holmes) okamura
254 Grateloupia angusta (okamura) S. Kawaguchi & H. W. Wang 2001
Carpopeltis angusta (Harvey) okamura;Prionitis angusta (okamura) okamura
255 Grateloupia articulata (okamura) S. Kawaguchi & H. W. Wang 2001
Carpopeltis articulata (okamura) okamura
256 Grateloupia chiangii S. Kawaguchi & H. W. Wang 2001 Carpopeltis divaricata okamura
257 Grateloupia conferta (Kützing) Kütz-ing 1867
258 Grateloupia cornea okamura 1913 Prionitis cornea (okamura) e. Y. Dawson
259 Grateloupia crispata (okamura) Y. P. Lee 2008 Carpopeltis crispata okamura
260 Grateloupia dichotoma J. Agardh 1842
261 Grateloupia divaricata okamura 1895
262 Grateloupia doryphora (Montagne) M. A. Howe 1914
263 Grateloupia filicina (J. V. Lamour-oux) C. Agardh 1822
264 Grateloupia ramosissima okamura 1913
265 Grateloupia schmitziana (okamura) S. Kawaguchi & H. W. Wang 2001
Cryptonemia schmitziana (oka-mura) okamura
266 Halymenia dilatata Zanardini 1851
267 Halymenia durvillei Bory de Saint-Vincent 1828
268 Halymenia elongata C. Agardh 1822
269 Halymenia floresii (Clemente) C. Agardh 1817
270 Halymenia formosa Harvey ex Kütz-ing 1866
271 Halymenia harveyana J. Agardh 1892
272 Halymenia maculata J. Agardh 1885
273 Halymenia microcarpa (Montagne) P. C. Silva 1987
274 Halymenia stipitata I. A. Abbott 1998
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275 Polyopes affinis (Harvey) S. Kawagu-chi & H. W. Wang 2002
Carpopeltis affinis (Harvey) oka-mura
276 Polyopes prolifer (Hariot) S. Kawa-guchi & H. W. Wang 2002
Carpopeltis prolifera (Hariot) S. Kawaguchi & Masuda
277
Spongophloea procumbens (Weber-van Bosse) Huisman, De Clerck, Prud'homme van Reine & Borowitzka 2011
Thamnoclonium procumbens Weber-van Bosse
278
Spongophloea treubii (Weber-van Bosse) Huisman, De Clerck, Prud'homme van Reine & Borowitzka 2011
Thamnoclonium treubii Weber-van Bosse
279 Yonagunia formosana (okamura) S. Kawaguchi & Masuda 2004
Carpopeltis formosana okamura; Prionitis formosana (okamura) S. Kawaguchi & Nguyen
280 Hapalidiaceae Lithothamnion australe f. brachiatum Foslie 1904
281 Lithothamnion australe f. minutulum Foslie 1904
282 Lithothamnion australe f. ubianum Foslie 1904
283 Lithothamnion indicum f. subtile Foslie 1907
284 Mesophyllum erubescens (Foslie) Me. Lemoine 1928
285 Mesophyllum imbricatum (Dickie) W. H. Adey 1970
286 Mesophyllum pulchrum (Weber-van Bosse & Foslie) Lemoine 1928
287 Mesophyllum siamense (Foslie) W. H. Adey 1970
288 Phymatolithon calcareum (Pallas) W. H. Adey & D. L. McKibbin 1970
289Phymatolithon purpureum (P. L. Crouan & H. M. Crouan) Woelkerling & L. M. Irvine 1986
290 HymenocladiaceaeAsteromenia exanimans G. W. Saun-ders, C. e. Lane, C. W. Schneider & Kraft 2006
291 Hymenocladia dactyloides (Sonder) J. Agardh 1870 Gracilaria dactyloides Sonder 1845
292 Kallymeniaceae Callophyllis adhaerens Yamada 1932
293 Callophyllis adnata okamura 1932
294 Callophyllis fastigiata (J. Agardh) J. Agardh 1876 Gracilaria fastigiata J. Agardh
295 Kallymenia callophylloides okamura & Segawa 1935
296 Kallymenia pacifica Kylin 1956
297 Kallymenia perforata J. Agardh 1872
298 Kallymenia rosea Womersley & R. e. Norris 1971
299 Kallymenia rubra Womersley & R. e. Norris 1971
300 Kallymenia sessilis okamura 1934
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301 Kallymenia tasmanica Harvey 1860
302 Pugetia japonica Kylin 1941 Callophyllis okamurae P. C. Silva
303 Liagoraceae Dermonema virens (J. Agardh) Pedro-che & Ávila ortíz 1996
Dermonema frappieri (Montagne & Millardet) Børgesen
304 Ganonema farinosum (J. V. Lamour-oux) K. C. Fan & Yung C. Wang 1974 Liagora farinosa J. V. Lamouroux
305 Helminthocladia australis Harvey 1863
306 Izziella orientalis (J. Agardh) Huis-man & Schils 2002 Liagora orientalis J. Agardh
307 Liagora boergesenii Yamada 1938
308 Liagora canariensis Børgesen 1927
309 Liagora ceranoides J. V. Lamouroux 1816
Liagora ceranoides f. leprosa (J. Agardh) Yamada
310 Liagora fragilis Zanardini 1851
311 Liagora hawaiiana Butters 1911
312 Liagora japonica Yamada 1938
313 Liagora robusta Yamada 1938
314 Liagora segawae Yamada 1938
315Liagoropsis schrammii (P. L. Crouan & H. M. Crouan) Doty & I. A. Abbott 1964
316 Neoizziella divaricata (C. K. Tseng) S. M. Lin, S. Y.Yang & Huisman 2011 Liagora divaricata C. K. Tseng
317 Titanophycus setchellii (Yamada) S. M. Lin, S. Y. Yang & Huisman 2011 Liagora setchellii Yamada
318Titanophycus validus (Harvey) Huisman, G. W. Saunders & A. R. Sherwood 2006
Liagora tenuis J. Agardh; Liagora valida Harvey
319 Trichogloea requienii (Montagne) Kützing 1847
320 Yamadaella caenomyce (Decaisne) I. A. Abbott 1970 Liagora caenomyce Decaisne Manila, Luzon, Philippines (De-
caisne 1842: 119).
321 Lomentariaceae Ceratodictyon intricatum (C. Agardh) R. e. Norris 1987
Gelidiopsis intricata (C. Agardh) Vickers
322 Ceratodictyon repens (Kützing) R. e. Norris 1987
Gelidiopsis repens (Kützing) Weber-van Bosse
323 Ceratodictyon spongiosum Zanardini 1878
324 Ceratodictyon variabile (J.Agardh) R. e. Norris 1987
Gelidiopsis variabilis (Greville ex J. Agardh) F. Schmitz
325 Lomentaria articulata (Hudson) Lyngbye 1819
326 Lomentaria baileyana (Harvey) Farlow 1876
327 Lomentaria hakodatensis Yendo 1920
328 Lomentaria pinnata Segawa 1938
329 Palmariaceae Palmaria palmata (Linnaeus) Weber & Mohr 1805
330 Peyssonneliaceae Peyssonnelia conchicola Piccone & Grunow 1884
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331 Peyssonnelia distenta (Harvey) Yamada 1930
332 Peyssonnelia evae Weber-van Bosse 1921
Pearl Bank, Tawitawi, Sulu Archipelago, Philippines (Weber-van Bosse 1921: 279)
333 Peyssonnelia foveolata (Weber-van Bosse) Denizot 1968
334 Peyssonnelia indica (Weber-van Bosse) Denizot 1968
Syntype localities: Philippine Islands; various in Indonesia, including Semau and Solor (Silva et al. 1996: 212).
335 Peyssonnelia involvens Zanardini 1858
336 Peyssonnelia luzonensis P. A. Cordero 1977
Quezon Is., Hundred Is., Pan-gasinan, Philippines (Cordero 1977: 80).
337 Peyssonnelia mariti (Weber-van Bosse) Denizot 1968
338 Peyssonnelia obscura Weber-van Bosse 1921
Syntype localities: Philippine Islands; various in Indonesia, including Semau and Solor (We-ber-van Bosse 1921: 274-275).
339 Peyssonnelia orientalis (Weber-van Bosse) Cormaci & G. Furnari 1987
Peyssonnelia rubra f. orientalis Weber-van Bosse
Syntype localities: Various lo-calities in Indonesia and Philip-pine Islands (Weber-van Bosse 1921: 272).
340 Peyssonnelia rubra (Greville) J. Agardh 1851
341 Peyssonnelia squamaria (S. G. Gme-lin) Decaisne 1842
342 Polystrata dura Heydrich 1905
343 Ramicrusta calcea (Heydrich) K. R. Dixon 2013 Peyssonnelia calcea Heydrich
344 Phyllophoraceae Ahnfeltiopsis divaricata (Holmes) Masuda 1993 Gymnogongrus divaricatus Holmes
345 Ahnfeltiopsis flabelliformis (Harvey) Masuda 1993
Gymnogongrus flabelliformis Harvey
346 Ahnfeltiopsis okamurae P. C. Silva & DeCew 1992 Ahnfeltia furcellata okamura
347 Ahnfeltiopsis pygmaea (J. Agardh) P. C. Silva & DeCew 1992 Gymnogongrus pygmaeus J. Agardh
348 Gymnogongrus dilatatus (Turner) J. Agardh 1851
349 Phyllophora submaritima e. Y. Dawson 1949
350 Plocamiaceae Plocamium cirrhosum (Turner) M. J. Wynne 2002
Plocamium costatum (C. Agardh) J. D. Hooker & Harvey
351 Plocamium patens G. Martens 1868
Syntype localities: Timor, Indo-nesia; Zamboanga, Mindanao, Philippines (Silva et al. 1996: 343).
352 Plocamium telfairiae (W. J. Hooker & Harvey) Harvey ex Kützing 1849
353 Plocamium serrulatum var. pectina-tum P. A. Cordero 1977
Tajojora, Batan Is. Batanes, Philippines (Cordero 1977: 140).
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354 Pterocladiaceae Pterocladiella caloglossoides (M. A. Howe) Santelices 1998
Pterocladia caloglossoides (M. A. Howe) e. Y. Dawson
355 Pterocladiella capillacea (S. G. Gme-lin) Santelices & Hommersand 1997
Pterocladia capillacea (S. G. Gme-lin) Bornet; Pterocladia densa okamura
356 Pterocladiella nana (okamura) Shi-mada, Horiguchi & Masuda 2000 Pterocladia nana okamura
357 Pterocladiella taylorii (Joly) Santeli-ces 2007 Gelidiella taylori Joly
358 Rhizophyllidaceae Portieria hornemannii (Lyngbye) P. C. Silva 1987
Chondrococcus hornemannii (Lyng-bye) F. Schmitz Desmia hornemannii Lyngbye
359 Portieria japonica (Harvey) P. C. Silva 1987
360 Rhodogorgonaceae Renouxia antillana Fredericq & J. N. Norris 1995
361 Rhodomelaceae Acanthophora aokii okamura 1934
362 Acanthophora muscoides (Linnaeus) Bory de Saint-Vincent 1828
363 Acanthophora spicifera (M. Vahl) Børgesen 1910
Acanthophora orientalis J. Agardh; Acanthophora thierryi J. V. Lamour-oux
364 Acrocystis nana Zanardini 1872
365 Alsidium pusillum e. Y. Dawson 1963
366 Amansia pumila (Sonder) J.Agardh 1892
Melanamansia pumila (Sonder) R. Norris
367 Aneurianna lorentzii (Weber-van Bosse) L. e. Phillips 2006
Lenormandiopsis lorentzii (Weber-van Bosse) Papenfuss
368 Bostrychia calliptera (Montagne) Montagne 1842
369 Bostrychia intricata (Bory de Saint-Vincent) Montagne 1852
370 Bostrychia kelanensis Grunow 1936
371 Bostrychia moritziana (Sonder ex Kützing) J. Agardh 1863
372 Bostrychia radicans (Montagne) Montagne 1842
373 Bostrychia simpliciuscula Harvey ex J. Agardh 1863
374 Bostrychia tenella (J. V. Lamouroux) J. Agardh 1863 Bostrychia binderi Harvey
375 Chondria armata (Kützing) okamura 1907
376 Chondria crassicaulis Harvey 1860
377 Chondria curvilineata F. S. Collins & Hervey 1917
378 Chondria dasyphylla (Woodward) C. Agardh 1817
379 Chondria polyrhiza F. S. Collins & Hervey 1917
380 Chondria repens Børgesen 1924
381 Chondria sedifolia Harvey 1853
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382 Chondria seticulosa (Forsskål) C. Agardh 1822
383 Chondria sibogae Weber-van Bosse 1923
Syntype localities: Borneo Bank and Lombok, Indonesia; Sulu Archipelago, Philippines (Silva et al. 1996: 485).
384 Chondrophycus carolinensis (Y. Saito) K. W. Nam 2007 Laurencia carolinensis Y. Saito
385 Chondrophycus cartilagineus (Ya-mada) Garbary & J. T. Harper 1998 Laurencia cartilaginea Yamada
386Chondrophycus ceylanicus (J. Agardh) M. J. Wynne, Serio, Cormaci & G. Furnari 2005
Laurencia ceylanica J. Agardh
387Chondrophycus columellaris (Børge-sen) e. Coppejans & A. J. K. Millar 2000
Laurencia columellaris Børgesen
388 Chondrophycus glandulifer (Kützing) Lipkin & P. C. Silva 2002
Laurencia glandulifera (Kützing) Kützing
389 Chondrophycus intermedius (Ya-mada) Garbary & J. T. Harper 1994 Laurencia intermedia Yamada
390Chondrophycus parvipapillatus (C.K.Tseng) Garbary & J. T. Harper 1994
Laurencia parvipapillata C.K. Tseng
391 Chondrophycus tronoi (Ganzon-Fortes) K. W. Nam 1999 Laurencia tronoi Ganzon-Fortes
North of Pulong Bakaw, Ca-latagan, Batangas, Philippines (Ganzon-Fortes 1982: 404).
392 Chondrophycus undulatus (Yamada) Garbary & J. T. Harper 1998 Laurencia undulata Yamada
393 Digenea simplex (Wulfen) C. Agardh 1822
394 Enantiocladia okamurae Yamada 1930
395 Endosiphonia spinulosa (Harvey) Womersley & Parsons 2003 Endosiphonia spinuligera Zanardini
396 Erythrocolon podagricum J. Agardh 1874
397 Exophyllum wentii Weber-van Bosse 1911
Syntype localities: Stat. 58. Ile de Savu récif; Stat. 80. Banc de Bornéo 40-50 m; Stat. 99. Ile Ubian du nord, Archipel de Sulu 12-33 m [North Ubian Island, Sulu Archipelago, Philippine Islands; Indonesia: Borneo Bank and Sawu] (Weber-van Bosse 1911: 29).
398 Herposiphonia delicatula Hollenberg 1968
399 Herposiphonia dendroidea Hollen-berg 1968
400 Herposiphonia dendroidea var. minor Hollenberg 1968
401 Herposiphonia nuda Hollenberg 1968
402 Herposiphonia obscura Hollenberg 1968
403 Herposiphonia pacifica Hollenberg 1968
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404 Herposiphonia parca Setchell 1926
405 Herposiphonia plumula (J. Agardh) Falkenberg 1901
406 Herposiphonia secunda (C. Agardh) Ambronn 1880
407 Herposiphonia secunda f. tenella (C. Agardh) M. J. Wynne 1985
408 Herposiphonia subdisticha okamura 1899
409 Herposiphonia tenella (C. Agardh) Ambronn 1880
410 Herposiphonia trichia Hollenberg 1968
411 Laurencia brongniartii J. Agardh 1841
412 Laurencia caraibica P. C. Silva 1972
413 Laurencia chondrioides Børgesen 1918
414 Laurencia clavata Sonder 1853
415 Laurencia composita Yamada 1931
416 Laurencia decumbens Kützing 1863
417 Laurencia dendroidea J. Agardh 1852 Laurencia obtusa var. dendroidea (J. Agardh) Yamada
418 Laurencia distichophylla J. Agardh 1852
419 Laurencia flexilis Setchell 1926
420 Laurencia forsteri (Mertens ex Turner) Greville 1830
421 Laurencia intricata J. V. Lamouroux 1813 Laurencia implicata J. Agardh
422 Laurencia majuscula (Harvey) A. H. S. Lucas 1935
423 Laurencia mariannensis Yamada 1931
424 Laurencia nidifica J. Agardh 1852
425 Laurencia nipponica Yamada 1931
426 Laurencia obtusa (Hudson) J. V. Lamouroux 1813
427 Laurencia obtusa var. densa Yamada 1931
428 Laurencia okamurae Yamada 1931 Laurencia japonica Yamada
429 Laurencia pinnata Yamada 1931
430 Laurencia similis K. W. Nam & Saito 1991
431 Laurencia snackeyi (Weber-van Bosse) M. Masuda 1997
Laurencia obtusa var. snackeyi (Weber-van Bosse) Yamada
432 Laurencia subsimplex C. K. Tseng 1943
433 Laurencia tropica Yamada 1931
434 Laurencia venusta Yamada 1931
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435 Leveillea jungermannioides (Hering & G. Martens) Harvey 1855
436 Lophocladia lallemandi (Montagne) F. Schmitz 1893
437 Lophosiphonia cristata Falkenberg 1901
438 Melanamansia glomerata (C. Agardh) R. e. Norris 1995 Amansia glomerata C. Agardh
439 Murrayella periclados (C. Agardh) F. Schmitz 1893
440 Neosiphonia apiculata (Hollenberg) Masuda & Kogame 2003 Polysiphonia apiculata Hollenberg
441 Neosiphonia beaudettei (Hollenberg) M. S. Kim & I. A. Abbott 2006 Polysiphonia beaudettei Hollenberg
442Neosiphonia ferulacea (Suhr ex J. Agardh) S. M. Guimarães & M. T. Fujii 2004
Polysiphonia ferulacea Suhr ex J. Agardh
443 Neosiphonia gorgoniae (Harvey) S. M. Guimarães & M. T. Fujii 2004 Polysiphonia gorgoniae Harvey
444 Neosiphonia hawaiiensis (Hollen-berg) M. S. Kim & I. A. Abbott 2006
Polysiphonia hawaiiensis Hol-lenberg
445 Neosiphonia poko (Hollenberg) I. A. Abbott 2002 Polysiphonia poko Hollenberg
446 Neosiphonia sertularioides (Grate-loup) K. W. Nam & P. J. Kang 2012
Neosiphonia flaccidissima (Hollen-berg) M. S. Kim & I. K. Lee
447 Neosiphonia sphaerocarpa (Børge-sen) M. S. Kim & I. K. Lee 1999
Polysiphonia sphaerocarpa Børge-sen
448 Neosiphonia tepida (Hollenberg) S. M. Guimarães & M. T. Fujii 2004 Polysiphonia tepida Hollenberg
449 Neosiphonia tongatensis (Harvey ex Kützing) M. S. Kim & I. K. Lee 1999
Polysiphonia mollis var. tongatensis (Harvey & Kützing) Hollenberg
450 Neosiphonia upolensis (Grunow) M. S. Kim & Boo 2008 Polysiphonia upolensis Grunow
451 Neurymenia fraxinifolia (Mertens ex Turner) J. Agardh 1863
452 Osmundaria obtusiloba (C. Agardh) R. e. Norris 1991
Vidalia obtusiloba (Mertens ex C. Agardh) J. Agardh
453 Osmundea pinnatifida (Hudson) Stackhouse 1809
Laurencia pinnatifida (Hudson) J. V. Lamouroux
454 Palisada capituliformis (Yamada) K. W. Nam 2007 Laurencia capituliformis Yamada
455 Palisada corallopsis (Montagne) Sentíez, M. T. Fujii & Díaz 2008
Laurencia corallopsis (Montagne) M. A. Howe
456 Palisada flagellifera (J. Agardh) K. W. Nam 2007 Laurencia flagellifera J. Agardh
457 Palisada parvipapillata (C. K. Tseng) K. W. Nam 2007
Chondrophycus parvipapillatus (C. K. Tseng) Garbary & J. T. Harper
458Palisada patentiramea (Montagne) Cassano, Sentíes, Gil-Rodríguez & M. T. Fujii 2009
Chondrophycus patentiramea (Mon-tagne) Nam; Laurencia patentiramea (Montagne) Kützing
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459 Palisada perforata (Bory de Saint-Vincent) K. W. Nam 2007
Chondrophycus papillosus (C. Agardh) D. J. Garbary & J. T. Harper; Laurencia papillosa (C. Agardh) Greville, Setchell & N. L. Gardner
460 Palisada robusta K. W. Nam 2007 Laurencia palisada Yamada
461 Palisada surculigera (C. K. Tseng) K. W. Nam 2007 Laurencia surculigera C. K. Tseng
462Palisada thuyoides (Kützing) Cas-sano, Sentíes, Gil-Rodríguez & M. T. Fujii 2009
Laurencia paniculata (C. Agardh) J. Agardh
463 Palisada yamadana (M. A. Howe) K. W. Nam 2007 Laurencia yamadana M. A. Howe
464 Polysiphonia flabelluta Harvey 1860
465 Polysiphonia forfex Harvey 1859
466 Polysiphonia fragilis Suringar 1867
467 Polysiphonia howei Hollenberg 1945
468 Polysiphonia japonica var. savatieri (Hariot) Yoon 1986 Polysiphonia savatieri Hariot
469 Polysiphonia mollis J. D. Hooker & Harvey 1847
470 Polysiphonia pacifica Hollenberg 1942
471 Polysiphonia pacifica var. delicatula Hollenberg 1942
472 Polysiphonia scopulorum Harvey 1855
473 Polysiphonia scopulorum var. villum (J. Agardh) Hollenberg 1968
474 Polysiphonia sparsa (Setchell) Hol-lenberg 1968
475 Polysiphonia subtillisima Montagne 1840
476 Polysiphonia triton P. C. Silva 1987
477 Polysiphonia tsudana Hollenberg 1968
478 Tolypiocladia calodictyon (Harvey ex Kützing) P. C. Silva 1952
479 Tolypiocladia condensata (Weber-van Bosse) P. C. Silva 1952
480 Tolypiocladia glomerulata (C. Agardh) F. Schmitz 1897
481 Womersleyella setacea (Hollenberg) R. e. Norris 1992 Polysiphonia setacea Hollenberg
482 Yuzurua poiteaui (J. V. Lamouroux) Martin-Lescanne 2010
Laurencia poiteaui (J. V. Lamour-oux) M. A. Howe
483 Rhodymeniaceae Botryocladia botryoides (Wulfen) Feldmann 1941
484 Botryocladia leptopoda (J. Agardh) Kylin 1931
485 Botryocladia pyriformis (Børgesen) Kylin 1931
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486 Botryocladia skottsbergii (Børgesen) Levring 1941
487 Botryocladia uvarioides e. Y. Daw-son 1944
488 Chrysymenia kaernbachii Grunow 1898 Chrysemenia kaernbachii Grunow
489 Rhodymenia californica Kylin 1931
490 Rhodymenia coacta okamura & Segawa 1935
491 Rhodymenia decumbens W. R. Taylor 1945
492 Rhodymenia divaricata e. Y. Dawson 1941
493 Rhodymenia intricata (okamura) okamura 1930
494 Sarcodiaceae Sarcodia montagneana (J. D. Hooker & Harvey) J. Agardh 1852
495 Sarcomeniaceae Cottoniella filamentosa (M. A. Howe) Børgesen 1920
496 Schizymeniaceae Titanophora incrustans (J. Agardh) Børgesen 1949
497 Titanophora mauritiana Børgesen 1949
498 Titanophora weberae Børgesen 1943
499 Scinaiaceae Scinaia aborealis Huisman 1986
500 Scinaia hormoides Setchell 1914
501 Scinaia japonica Setchell 1914
502 Scinaia latifrons M. A. Howe 1911
503 Scinaia moniliformis J. Agardh 1885
504 Scinaia tsinglanensis C. K. Tseng 1941
505 SebdeniaceaeLesleigha yamadae (okamura & Segawa) G. W. Saunders & Kraft 2011
Sebdenia yamadae okamura & Segawa
506 Sebdenia flabellata (J. Agardh) P. G. Parkinson 1980
507 Sebdenia limensis (Kützing) M. A. Howe 1914
508 Solieriaceae Betaphycus gelatinus (esper) Doty ex P. C. Silva 1996
Eucheuma gelatinae (esper) J. Agardh; Eucheuma gelatinum (esper) Doty
509 Betaphycus philippinensis Doty 1995Betaphycus philippinense Doty; Betaphycus philippinensis var. philippinensis Doty
Holotype: eastern Sorsogon, Philippines (Doty 1995: 237).
510 Eucheuma alvarezii var. ajakii-assii Doty 1985
eucheuma farm, about 1 km southwest of the southern tip of Tumindao Island, Sulu Archipel-ago, Philippines (Doty 1985: 42).
511 Eucheuma amakusaense okamura 1902
512 Eucheuma arnoldii Weber-van Bosse 1928
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513 Eucheuma arnoldii var. alcyonida Kraft 1972
Bulusan, Sorsogon, Philippines (Kraft 1972: 323).
514 Eucheuma crassum Zanardini 1878
515 Eucheuma crustiforme Weber-van Bosse 1928
516 Eucheuma denticulatum (N. L. Bur-man) F. S. Collins & Hervey 1917
Eucheuma muricatum (S.G.Gmelin) Weber-van Bosse
517 Eucheuma denticulatum var. endong Trono & Ganzon-Fortes 2012
Dancalan, Sorsogon, Philippines (Ganzon-Fortes et al. 2012: 1110)
518 Eucheuma edule (Kützing) Weber-van Bosse 1926
519 Eucheuma horridum J. Agardh 1852
520 Eucheuma isiforme (C. Agardh) J. Agardh 1847
521 Eucheuma leeuwenii Weber-van Bosse 1928
522 Eucheuma perplexum Doty 1988
523 Eucheuma serra (J. Agardh) J. Agardh 1847
524 Kappaphycus alvarezii (Doty) Doty ex P. C. Silva 1996
Eucheuma alvarezii Doty; Eucheuma cottonii Weber-van Bosse; Eucheuma cottonii var. erecta ?
525 Kappaphycus alvarezii var. tamba-lang (Doty) nom. inval.
Eucheuma alvarezii var. tambalang Doty
Seaweed farm at Calatagan, Batangas, Philippines (Doty 1985: 41).
526 Kappaphycus cottonii (Weber-van Bosse) Doty ex P. C. Silva 1996 Eucheuma okamurae Yamada
527 Kappaphycus procrusteanus (Kraft) Doty 1988
Eucheuma procrusteanum Kraft; Kappaphycus procrusteanum Kraft
Semirara Islands of the northern Sulu Sea, Philippines (Kraft 1970: 215).
528 Kappaphycus striatus (F. Schmitz) Doty ex P. C. Silva 1996
Eucheuma striatum F. Schmitz; Kappaphycus striatum F. Schmitz
529 Meristotheca coacta okamura 1930
530 Meristotheca papulosa (Montagne) J. Agardh 1872
531 Sarconema filiforme (Sonder) Kylin 1932
Dicranema furcellatum (Montagne) J. Agardh
532 Solieria dura (Zanardini) F. Schmitz 1895
533 Solieria pacifica (Yamada) Yoshida 1989
534 Solieria robusta (Greville) Kylin 1932
535 Wurdemannia miniata (Sprengel) Feldmann & G. Hamel 1934
536 Sporolithaceae Sporolithon erythraeum (Rothpletz) Kylin 1956
537 Sporolithon schmidtii (Foslie) G. D. Gordon, Masaki & Akioka 1976
538 Sporolithon sibogae (Weber-van Bosse & Foslie) P. C. Silva 1987
Archaeolithothamnion sibogae Weber-van Bosse & Foslie
Pearl Bank, Sulu Archipelago, Philippines (Silva et al. 1996: 277)
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539 Sporolithon timorense (Foslie) P. C. Silva 1987
540 Spyridiaceae Spyridia filamentosa (Wulfen) Har-vey 1833
541 Spyridia velasquezii G. C. Trono 1972
542 Stylonemataceae Chroodactylon ornatum (C. Agardh) Basson 1979
543 Rhodosorus marinus Geitler 1930
544 Stylonema alsidii (Zanardini) K. M. Drew 1956
Goniotrichum alsidii (Zanardini) M. A. Howe
545 Wrangeliaceae Anotrichium tenue (C. Agardh) Nageli 1862
546 Dasyphila plumarioides Yendo 1920
547 Gordoniella yonakuniensis (Yamada & T. Tanaka) Itono 1977
548 Griffithsia ovalis Harvey 1855
549 Griffithsia rhizophora Grunow ex Weber-van Bosse 1923
550 Griffithsia subcylindrica okamura 1930
551 Gymnothamnion elegans (Schousboe ex C. Agardh) J. Agardh 1892
552 Haloplegma duperreyi Montagne 1842
553 Pleonosporium globuliferum Levring 1941
554Ptilothamnion cladophorae (Yamada & T. Tanaka) G. Feldmann-Mazoyer 1941
555 Wrangelia argus (Montagne) Mon-tagne 1856
556 Wrangelia bicuspidata Børgesen 1916
557 Wrangelia penicillata (C. Agardh) C. Agardh 1828
558 Wrangelia plumosa Harvey 1844
559 Wrangelia velutina (Sonder) Harvey 1855
560 Rhodophyta incertae sedis
Callophycus serratus (Harvey ex Kützing) P. C. Silva 1957
Taxon with Generic Record Only
List No. Family Taxon Recorded Remarks Reference
561 Callithamniaceae Callithamnion Lyngbye Kraft et al. 1999
562 Euptilota (Kützing) Kützing Kraft et al. 1999
563 Dumontiaceae Dudresnaya P. L. Crouan & H. M. Crouan Kraft et al. 1999
564 Rhabdoniaceae Rhabdonia J. D. Hooker & Harvey Kraft et al. 1999
Species with No Matching Record in AlgaeBase
Family Taxon Recorded Remarks Reference
Corallinaceae Amphiroa fragilissima f. fragilissima (Linaeus) (Lamoroux) Weber van Bosse
Also not listed in INA Database Trono 2004
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Jania tenella var. tenella e. Y. Dawson Also not listed in INA Database Trono 2004
Halymeniaceae Halymenia japonica? Domantay Also not listed in INA Database Silva et al. 1987, Ganzon-Fortes 2012
Kallymeniaceae Kallymenia denticulata No authority of taxon listed;Also not listed in INA Database
Ganzon-Fortes 2012
Species Listed as Preliminary Entry in AlgaeBase
Family Taxon Recorded Remarks Reference
Corallinaceae Jania constricta Kützing, nom. illeg. Also not listed in INA Database Trono 2004
Peyssonneliaceae Peyssonnelia rubra var. orientalis Weber-van Bosse
As Peyssonnelia rubra f. orientalis Weber-van Bosse in INA Database
Silva et al. 1987;Silva 1996-to date
luzoniensis Martens was also described based on Phil-ippine materials (Table 1). But this taxon is listed as a preliminary entry in AlgaeBase so its current taxonomic status remains unchecked.
Although the number of species (893) recorded from the Philippines compiled in the present work remains com-parable with that compiled by Ganzon-Fortez (2012) for records up to 1999 (888 species, excluding Cyanobacte-ria), the details of the species list are quite different. This is so mainly because of the large number of taxa that has been synonymized in recent years. on a purely nu-merical sense, there is a net increase of 21 species among the reds in the new list, compared to a drop of 1 and 15 species respectively among the browns and the greens. Payo et al. (2013) recently suggested naming 21 new cryptic species of the red alga Portieria from the Philip-pines based on three molecular markers they used. These “new” species have not been considered in the present compilation.
As in other attempts to compile a listing of algal records, no actual examination of the algal materials was done. This listing should therefore be viewed simply as a listed record with the validity of the published species names verified. It does not, however, verify the correct identifi-cation of each original record. Readers which find mis-takes in the list are most welcomed to inform the authors so an improved list can be made in the future. Further studies involving actual examination of the original algal materials are expected to verify the species identity.
ACKNoWLeDGeMeNTSThis paper is dedicated to the memory of Leonardo L. Co, a foremost Philippine plant taxonomist, conservationist and botanical activist whose fruitful life was cut short by the wanton gun fire in the forest of Leyte, Philippines on November 15, 2010. We thank Dr. Benito Tan of the
National University of Singapore for his comments and editorial suggestions on the manuscript. Special thanks are also given to Dr. Lawrence Liao of Hiroshima University, Japan, for his very valuable comments, corrections and additions on our earlier list of algal records from the Philippines. This work benefited from funding from the Research Grant Council of Hong Kong GRF Project No. CUHK460012.
ReFeReNCeSABBoTT IA. 1962. Some Liagora-inhabiting species
of Acrochaetium. occasional Papers Bernice Pauahi Bishop Museum 23: 77-120.
ABBoTT IA. 1985. Gracilaria from the Philippines: List and distribution of the species. Taxonomy of economic Seaweeds 1: 89-90.
ABBoTT IA. 1988. Some species of Gracilaria and Polycavernosa from the Siboga expedition. Taxonomy of economic Seaweeds 2: 151-156.
ABBoTT IA. 1994. New records and a reassessment of Gracilaria (Rhodophyta) from the Philippines. Taxonomy of economic Seaweeds 4: 111-118.
ABBoTT IA. 1999. Notes on some species of Halymenia in the southwestern Pacific. Taxonomy of economic Seaweeds 7: 163-172.
AGARDH JG. 1848. Species genera et ordines algarum, seu descriptiones succinctae specierum, generum et ordinum, quibus algarum regnum constituitur. Volumen Primum. Algas fucoideas complectens. Lundae [Lund]: C.W.K. Gleerup. pp. [i-vi], [i]-viii, [1]-363.
AJISAKA T, NoRo T, YoSHIDA T. 1995. Zygocarpic Sargassum species (Subgenus Sargassum) from Japan. Taxonomy of economic Seaweeds 5: 11-44.
ANG Po, TRoNo GC. 1987. The genus Sargassum
Ang et al.: A verification of Philippine marine macroalgaePhilippine Journal of ScienceVol. 142: Special Issue
45
(Phaeophyta, Sargassaceae) from Balibago, Calatagan, Philippines. Botanica Marina 30: 387-397.
AUNGToNYA C, LIAo LM. 2002. Marine flora (algae and seagrasses) in the reference collection of the Phuket Marine Biological Center, Thailand. Phuket Marine Biological Center Research Bulletin 64: 65-80.
BeLLeZA DFC, LIAo LM. 2007. Taxonomic inventory of the marine green algal genus Caulerpa (Chlorophyta, Bryopsidales) at the University of San Carlos (CeBU) Herbarium. Philippine Scientist 44: 71-104.
BeRKeLeY MJ. 1842. enumeration of fungi, collected by H. Cuming, esq. F.L.S. in the Philippine Islands. London Journal of Botany 1: 142-157.
CHIANG YM, YoSHIDA T, AJISAKA T, TRoNo GC, TSeNG CK, LU B. 1992. Distribution and variation in Sargassum polycystum C. A. Agardh (Fucales, Phaeophyta). Taxonomy of economic Seaweeds 3: 35-42.
CHIANG YM. 1985. Gracilaria from Taiwan: Key, list and distribution of the species. Taxonomy of economic Seaweeds 1: 81-83.
CHIRAPART A, oHNo M, JARAYAPUND P. 2003. Marine algae of Ko Si Chang, eastern Thailand. Bulletin of Marine Science and Fisheries, Kochi University No. 22: 107-118.
CoPPeJANS e, LeLIAeRT F, DARGeNT o, De CLeRCK o. 2001. Marine green algae (Chlorophyta) from the north coast of Papua New Guinea. Cryptogamie, Algologie 22(4): 375-443.
CoRDeRo PA JR. 1974. A new variety of Halimeda velasquezii Taylor from Camiguin Island. Kalikasan, Philippine Journal of Biology 2: 114-117.
CoRDeRo PA JR. 1976. Phycological observations, II: Porphyra marcosii, a new species from the Philippines. Acta Manilana, Series A 15(24): 14-24.
CoRDeRo PA JR. 1977. Studies on Philippine marine red algae. Special Publications from the Seto Marine Biological Laboratory 4: 1-258.
DAWSoN eY. 1954. Notes on tropical Pacific marine algae. Bulletin of the Southern California Academy of Sciences 53: 1-7.
DeCAISNe J. 1842. Mémoires sur les corallines ou polypiers calcifères. Annales des Sciences Naturelles, Botanique, Série 2 16: 85-120.
DINH NH, NANG HQ. 1995. Species of Eucheuma and Kappaphycus in Vietnam. Taxonomy of economic
Seaweeds 5: 229-235.
DoTY MS. 1985. Eucheuma alvarezii sp. nov. (Gigartinales, Rhodophyta) from Malaysia. Taxonomy of economic Seaweeds 1: 37-45.
DoTY MS. 1988. Prodromus ad systematica eucheumatoideorum: A tribe of commercial seaweeds related to Eucheuma (Solieriaceae, Gigartinales). Taxonomy of Seaweeds 2: 159-207.
DoTY MS. 1995. Betaphycus philippinensis gen. et sp. nov. and related species (Solieriaceae, Gigartinales). Taxomony of economic Seaweeds 5: 237-245.
FReDeRICQ S, NoRRIS JN. 1992. Studies on cylindrical species of Western Atlantic Gracilaria (Gracilariales, Rhodophyta): G. cylindrica Borgesen and G. blodgettii Harvey. Taxonomy of economic Seaweeds 3: 211-231.
GALLITo PM, LIAo LM. 2006. Asteromenia (Rhodophyta, Rhodymeniales), another new addition to the marine algal flora of the Philippines. Philippine Scientist 43: 77-83.
GANZoN-FoRTeS eT. 1982. Laurencia tronoi (Rhodophyta, Ceramiales), a new species from Calatagan, Batangas, Philippines. Kalikasan, Philippine Journal of Biology 11: 27-38.
GANZoN-FoRTeS eT. 2012. A historical account of biodiversity studies on Philippine seaweeds (1800-1999). Coastal Marine Science 35(1): 182-201.
GANZoN-FoRTeS eT, TRoNo GC JR, VILLANUeVA RD, RoMeRo JB, MoNTAño MNe. 2012. ‘endong’, a rare variety of the farmed carrageenophyte Eucheuma denticulatum (Burman) Collins & Hervey from the Philippines. Journal of Applied Phycology 24: 1107–1111.
GePP A, GePP eS. 1911. The Codiaceae of the Siboga expedition including a monograph of Flabellariaea and Udoteae. Siboga expedite Monographie Vol. 62. e. J. Brill: Leiden. pp. 1-150, 22 plates.
GeRALDINo PJL, LIAo LM, Boo SM. 2005. Morphological study of the marine algal genus Padina (Dictyotales, Phaeophyceae) from Southern Philippines: 3 species new to Philippines. Algae 20(2): 99-112.
GeRALDINo PJL, RIoSMeNA-RoDRIGUeZ R, LIAo LM, Boo SM. 2010. Phylogenetic relationships within the genus Hypnea (Gigartinales, Rhodophyta), with a description of H. caespitosa sp. nov. Journal of Phycology 46: 336-345.
Ang et al.: A verification of Philippine marine macroalgaePhilippine Journal of ScienceVol. 142: Special Issue
46
GeRUNG GS, LoKoLLo FF, KUSeN JD, HARAHAP AP. 2006. Study on the seaweeds of Ambon Island, Indonesia. Coastal Marine Science 30(1): 162-166.
GILBeRT WJ. 1961. An annotated checklist of Philippine marine Chlorophyta. Philippine Journal of Science 88: 413-451.
GRUNoW A. 1915. Additamenta ad cognitionem Sargassorum. Verhandlungen der Kaiserlich-Möniglichen Zoologisch-Botanischen Gesellschaft in Wien 65: 329-448.
GRUNoW A. 1916. Additamenta ad cognitionem Sargassorum. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 66: 1-48,136-185.
GUIRY MD, GUIRY GM. 2013. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. http://www.algaebase.org; last searched on 2 May, 2013.
GURGeL CFD, FReDeRICQ S, NoRRIS JN. 2004. Molecular systematics and taxonomy of flattened species of Gracilaria Greville (Gracilariaceae, Gracilariales, Rhodophyta) from the western Atlantic. Taxonomy of economic Seaweeds 9: 159-199.
GURGeL CFD, LIAo LM, FReDeRICQ S, HoMMeRSAND MH. 2003. Systematics of Gracilariopsis (Gracilariales, Rhodophyta) based on rbcL sequence analyses and morphological evidence. Journal of Phycology 39: 154-171.
HUISMAN JM. 2006. Tricelocarpa. In: Algae of Australia, Nemaliales. (Huisman, J.M. ed.). Canberra: Australian Biological Resources Study, pp. 24-28.
KRAFT GT. 1970. Eucheuma procrusteanum, a new red algal species from the Philippines. Phycologia 8: 215-219.
KRAFT GT. 1972. Preliminary studies of Philippine Eucheuma species (Rhodophyta). Part1. Taxonomy and ecology of Eucheuma arnoldii Weber-van Bosse. Pacific Science 26: 318-334.
KRAFT GT. 1973. The morphology of Cubiculosporum koronicarpis gen. et sp. nov., representing a new family in the Gigartinales (Rhodophyta). American Journal of Botany 60: 872-882.
KRAFT GT. 1976. The morphology of Beckerella scalaramosa, a new species of Gelidiales (Rhodophyta) from the Philippines. Phycologia 15: 85-91.
KRAFT GT, LIAo LM, MILLAR AJK, CoPPeJANS eGG, HoMMeRSAND MH, FReSHWATeR
DW. 1999. Marine benthic red algae (Rhodophyta) from Bulusan, Sorsogon Province, Southern Luzon, Philippines. Philippine Scientist 36: 1-50.
Le NH, NGUYeN HD. 2006 Contribution to the study of Gracilaria and relative genera (Gracilariales, Rhodophyta) from Vietnam. Coastal Marine Science 30(1): 214-221.
Lee AC, LIAo LM, TAN KS. 2009. New records of marine algae on artificial structures and intertidal flats in coastal waters of Singapore. Raffles Bulletin of Zoology Supplement No. 22: 5-40.
LeLIAeRT F, PAYo DA, CALUMPoNG HP, De CLeRCK o. 2011. Chaetomorpha philippinensis (Cladophorales, Chlorophyta), a new marine microfilamentous green alga from trophical waters. Phycologia 50(4): 384-391.
LeWMANoMoNT K. 1998. Species of Hypnea from Thailand. Taxonomy of economic Seaweeds 6: 179-191.
LIAo LM, UY FA, HeYRoSA NA. 2004. Macrobenthic marine algae and seagrasses of the Anambas expedition 2002. Raffles Bulletin of Zoology Supplement No. 11: 19-23.
LIN SM, FReDeRICQ S, LIAo LM. 2001. Neotypification and taxonomic status of Opephyllum martensii Schmitz. in Schmitz et Hauptfleisch (Delesseriaceae, Rhodophyta) from Zamboanga, Southern Philippines. Botanica Marina 44: 589-595.
MANZA AV. 1939. Two new species of Philippine Cladophora. Bulletin of the National Research Council of the Philippines 23: 109.
MARTeNS G. VoN 1866. Die Tange. Die Preussische expedition nach ost-Asien. Nach amtlichen Quellen. Botanischer Theil. Berlin: Verlag de Königlichen Geheimen ober-Hofbuchdruckerei, pp. 1-152, pls I-Viii.
MCDeRMID KJ. 1988. Laurencia from the Hawaiian Islands: Key, annotated list, and distribution of the species. Taxonomy of economic Seaweeds 2: 231-247.
MILLAR AJK. 1997. Some flattened species of Gracilaria from Australia. Taxonomy of economic Seaweeds 6: 111-123.
MILLAR AJK, KRAFT GT. 2001. Monograph of the green macroalgal genus Rhipilia (Udoteaceae, Halimedales), with a description of R. crassa sp. nov. from Australia and the Philippines. Phycologia 40: 21-34.
Ang et al.: A verification of Philippine marine macroalgaePhilippine Journal of ScienceVol. 142: Special Issue
47
MILLAR AJK, XIA BM. 1997. Studies on terete species of Australian Gracilaria. Taxonomy of economic Seaweeds 6: 103-109.
MoDeLo RB, UMeZAKI I, LIAo LM. 1998. Morphology of Sargassum corderoi (Phaeophyta, Fucales), a new seaweed species from central Philippines. Algae 13(1): 79-83.
MoNTAGNe C. 1842. Troisième centurie de plantes cellulaires exotiques nouvelles. Décades V, VI, VII et VIII. Annales des Sciences Naturelles, Botanique, Seconde Série 18:241-282, pl. 7.
MoNTAGNe C. 1844. Plantae cellulares quas in insulis Philippinensibus a cl. Cuming collectae recensuit, observationibus non nullis descriptionibusque illustravit C. Montagne, D.M. London Journal of Botany 3: 658-662.
NoRRIS Re. 1994. Hawaiian Phyllophoraceae. Taxonomy of economic Seaweeds 4: 185-191.
oHNo M, TeRADA R, YAMAMoTo H. 1999. The species of Gracilaria from Vietnam. Taxonomy of economic Seaweeds 7: 99-111.
PAYo DA, LeLIAeRT F, VeRBRUGGeN H, D’HoNDT S, CALUMPoNG HP, De CLeRCK o. 2013. extensive cryptic species diversity and fine-scale endemism in the marine red alga Portieria in the Philippines. Proceedings of the Royal Society of London B Biological Sciences 280: 20122660.
PHAM HT. 2002. Morphological variability of Kappaphycus cottonii. Taxonomy of economic Seaweeds 8: 247-255.
PHAM MN, TAN HTW, MITRoVIC S, Yeo HHT. 2011. A checklist of the algae of Singpapore, 2nd ed. Singapore: Raffles Museum of Biodiversity Research.
PHANG SM. 1994. New records of Malaysian marine algae. Hydrobiologia 285: 123-129.
PHANG SM. 2006. Seaweed resources in Malaysia: Current status and future propects. Aquatic ecosystem Health & Management 9(2): 185-202.
PHANG SM, LIM Pe, ooI JLS, YeoNG HY, NG WS, KUPPeR FC. 2008. Marine algae of Perak Island, Jarak Island and the Sembilan group of Islands in the Straits of Malacca. Malaysian Journal of Science 27(3): 47-60.
PHANG SM, LIM Pe, YeoNG HY, NG WS, SoNG SL. 2010. Marine algae collected during the Scientific expedition to Bachok, Kelantan and the Islands of
Terengganu with one new record, Pterocladiella for Malaysia. Malaysian Journal of Science 29 (Special Issue): 31-45.
PHANG SM, NoRo T, YoSHIDA T. 1995. Variations in Sargassum siliquosum J. Agardh (Fucales, Phaeophyta) found in Malaysia and Singapore. Taxonomy of economic Seaweeds 5: 55-66.
PHILLIPS N. 1995. Biogeography of Sargassum (Phaeophyta) in the Pacific Basin. Taxonomy of economic Seaweeds 5: 107-144.
PRATHeP A, PoNGPARADoN S, DARAKRAI A, WICHACHUCHeRD B, SINUToK S. 2011. Diversity and distribution of seaweed at Khanom-Mu Ko Thale Tai National Park, Nakhon Si Thammarat Province, Thailand. Songklanakarin Journal of Science and Technology 33(6): 633-640.
PRATHeP A, TANTIPRAPAS P. 2006. Preliminary report on the diversity and community structure of macroalgae before and after the 2004 tsunami at Talibong Island, Trang Province, Thailand. Coastal Marine Science 30(1): 189-195.
SANTeLICeS B, STeWART JG. 1985. Pacific species of Gelidium Lamouroux and other Gelidiales (Rhodophyta), with keys and descriptions to the common or economically important species. Taxonomy of economic Seaweeds 1: 17-32.
SIDIK BJ, HARAH ZM, KAWAGUCHI S. 2012. Historical review of seaweeed research in Malaysia before 2001. Coastal Marine Science 35(1): 169-177.
SILVA PC. (1996-to date). Index Nominum Algarum, University Herbarium, University of California, Berkeley http://ucjeps.berkeley.edu/INA.html. Last searched: 2 May, 2013.
SILVA PC, BASSoN PW, Moe RL. 1996. Catalogue of the benthic marine algae of the Indian ocean. University of California Publications in Botany 79:1-1259.
SILVA PC, MeNeZ eG, Moe RL. 1987. Catalog of the benthic marine algae of the Philippines. Smithsonian Contributions to the Marine Sciences 27:1-197.
SCHMITZ F, HAUPTFLeISCH P. 1897. Delesseriaceae. In: Die natürlichen Pflanzenfamilien nebst ihren Gattungen und wichtigeren Arten insbesondere den Nutzpflanzen unter Mitwirkung zahlreicher hervorragender Fachgelehrten, Teil 1, Abteilung 2. Leipzig: (engler, A. & Prantl, K. eds) Wilhelm
Ang et al.: A verification of Philippine marine macroalgaePhilippine Journal of ScienceVol. 142: Special Issue
48
engelmann, pp. 406-417.
TAN J, LIM Pe, PHANG SM. 2013. Phylogenetic relationship of Kappaphycus Doty and Eucheuma J. Agardh (Solieriaceae, Rhodophyta) in Malaysia. Journal of Applied Phycology 25: 13-29.
TANAKA T. 1967. Some marine algae from Batan and Camiguin Islands, Northern Philippines. I. Memoir of Faculty of Fisheries, Kagoshima University 16: 13-27.
TAYLoR WR. 1961. Cladophoropsis philippinensis, a new species from the western Pacific ocean. Botanica Marina 3: 56-59.
TAYLoR WR. 1962. Two undescribed species of Halimeda. Bulletin of the Torrey Botanical Club 89: 172-177.
TAYLoR WR. 1964. The genus Turbinaria in eastern seas. Journal of Linnean Society of London, Botany 58: 475-490.
TeRADA R, YAMAMoTo H. 2002. Additional notes on Gracilaria eucheumatoides Harvey from Vietnam. Taxonomy of economic Seaweeds 8: 225-230.
THoNGRoY P, LIAo L, PRATHeP A. 2007. Diversity, abundance and distribution of macroalgae at Sirinart Marine National Park, Phuket Province, Thailand. Botanica Marina 50(2): 88-96.
TITo oD, LIAo LM. 2000. ethnobotany of Solieria robusta (Gigartinales, Rhodophyta) in Zamboanga, Philippines. Science Diliman 12(2): 75-77.
TRoNo GC. 1992. The genus Sargassum in the Philippines. Taxonomy of economic Seaweeds 3: 43-94.
TRoNo GC. 1994. New species of Sargassum from the Philippines. Taxonomy of economic Seaweeds 4: 3-7.
TRoNo GC. 1997. Field guide & atlas of the seaweed resources of the Philippines. Makati: Bookmark. 306 pp.
TRoNo GC. 2004. Field guide & atlas of the seaweed resources of the Philippines, Volume 2. BAR, Marine environmental Resources Foundation. 261 pp.
TRoNo GC, ANG Po. 1982. Marine benthic algae from Bugsuk Island and vicinity, Palawan, Philippines. Kalikasan, Philippine Journal of Biology 11: 1-26.
TRoNo GC, oHNo M. 1992. Seaweeds collected from Bolinao, Pangasinan, Philippines. Bulletin of Marine Science and Fisheries, Kochi University No. 12: 39-50.
TSeNG CK, GILBeRT WJ. 1942. on new algae of the genus Codium from the South China Sea. Journal of the Washington Academy of Science 32: 291-296.
TSeNG CK, LU B. 1988. Studies on Chinese species of zygocarpic Sargassum. Taxonomy of economic Seaweeds 2: 23-54.
TSeNG CK, LU B. 1992. Studies on the malacocarpic Sargassum of China: II. Racemosae J. Agardh. Taxonomy of economic Seaweeds 3: 11-34.
TSeNG CK, LU B. 1995. Studies on the glomerulate Sargassum of China: III. The species group Binderia. Taxonomy of economic Seaweeds 5: 93-106.
TSeNG CK, LU B. 1997. Studies on the glomerulate Sargassum of China: IV. Series of Platycarpae. Taxonomy of economic Seaweeds 6: 9-25.
TSeNG CK, LU B. 1999. Studies on the biserrulic Sargassum of China: II. The series Coriilfoliae J. G. Agardh. Taxonomy of economic Seaweeds 7: 3-22.
TSeNG CK, LU B. 2002. Studies on the biserrulic Sargassum of China: III. on Sargassum parvifolium (Turner) C. Agardh, with a note on the classification of the Biserrulae. Taxonomy of economic Seaweeds 8: 3-9.
TSeNG CK, LU B. 2002. Studies on the biserrulic Sargassum of China: IV. The series Plagiophyllae Tseng et Lu. Taxonomy of economic Seaweeds 8: 11-33.
TSeNG CK, LU B. 2002. Studies on the biserrulic Sargassum of China: V. The series Ilicifoliae (J. Agardh) Tseng et Lu. Taxonomy of economic Seaweeds 8: 35-76.
VeLASQUeZ GT, LeWMANoMoNT K. 1975. Checklist on the study of the benthic marine algae of Thailand. Kasetsart University Fishery Research Bulletin No. 8.
WeBeR-VAN BoSSe A. 1911. Notice sur quelques genres nouveaux d’algues de l’Archipel Malaisien. Annales du Jardin Botanique de Buitenzorg 24: 25-33.
WeBeR-VAN BoSSe A. 1921. Liste des algues du Siboga. II. Rhodophyceae. Première partie. Protoflorideae, Nemalionales, Cryptonemiales. Siboga expedite Monographie Vol. 59b. e. J. Brill: Leiden. pp. 187-310, Figs. 53-109, Plates VI-VIII.
WeST JA, CALUMPoNG HP. 1996. Mesospora negrosensis sp. nov. (Phaeophyta) from the Philippines. The Philippine Scientist 33: 5-15.
WoNG SC, HARAH ZM, SIDIK BJ, ARSHAD AB. 2012. Comparison of seaweed communities of the two rocky shores in Sarawak, Malaysia. Coastal Marine Science 35(1): 78-84.
Ang et al.: A verification of Philippine marine macroalgaePhilippine Journal of ScienceVol. 142: Special Issue
49
XIA BM, WANG YQ. 1999. Taxonomic studies on Halymenia (Halymeniaceae, Halymeniales, Rhodophyta) from China. Taxonomy of economic Seaweeds 7: 173-176.
XIA BM, WANG YQ. 2004. Presence of a different Gracilaria in China. Taxonomy of economic Seaweeds 9: 249-252.
YAMAGISHI Y, MASUDA M. 1998. Species of Hypnea from Japan. Taxonomy of economic Seaweeds 6: 135-162.
YAMAGISHI Y, MASUDA M, ABe T, UWAI S, KoGAMe K, KAWAGUCHI S, PHANG SM. 2003. Taxonomic notes on marine algae from Malaysia. XI. Four species of Rhodophyceae. Botanica Marina 46: 534-547.
YAMAMoTo H, TeRADA R, MURAoKA D. 1999. on so-called Gracilaria coronopifolia from the Philippines and Japan. Taxonomy of economic Seaweeds 7: 89-97.
YAMAMoTo H, TRoNo GC. 1994. Two new species of Gracilaria from the Philippines. Taxonomy of economic Seaweeds 4: 95-101.
Ang et al.: A verification of Philippine marine macroalgaePhilippine Journal of ScienceVol. 142: Special Issue