a verification of reports of marine algal species from the

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5 Philippine Journal of Science 142: 5-49, Special Issue ISSN 0031 - 7683 Date Received: 2 May 2013 Marine Science Laboratory School of Life Sciences, Chinese University of Hong Kong Shatin, N.T., Hong Kong SAR, CHINA Put O. Ang, Jr., Sin Man Leung, and Mei Mei Choi A Verification of Reports of Marine Algal Species from the Philippines Corresponding Author: [email protected] Records of marine macroalgae reported from the Philippines were checked against AlgaeBase, the international database for algal nomenclatures, and Index Nominum Algarum (INA) Bibliographia Phycologica Universalis of the University of California at Berkeley Silva Center for Phycological Documentation to verify their present nomenclature, status of taxonomy and bibliographic reference. To date, 306 names of taxa (including species, varieties and forms) of greens (Chlorophyta), 234 names of taxa of browns (Ochrophyta, Phaeophyceae) and 751 names of taxa of reds (Rhodophyta), or a total of 1291 published names of taxa have been reported from the Philippines. Of these, 231 taxa representing 197 species in 20 families for green algae, 171 taxa representing 153 species in 10 families for brown algae, and 564 taxa representing 543 species in 52 families for red algae are considered valid records listed with their currently accepted names. All in all, 966 currently accepted taxa, representing 893 species in 82 families of marine macroalgae have been reported from the Philippines. Among the greens, 15 taxa have their type localities in the Philippines. This number is 40 for the browns and 33 for the reds. Proportionally, this is 6.5% of the total for the greens, 23.4% for the browns and 5.9% for the reds. Key Words: Philippine marine macroalgae, algal taxonomic status, algal type species from the Philippines, AlgaeBase, Index Nominum Algarum (INA) INTRODUCTION Philippines is known for its rich marine flora. Over the years, several efforts have been attempted to compile and annotate the records of marine algae collected from the Philippines, including those by Silva et al. (1987), the two volume works by Trono (1997, 2004), and more recently, the checklist with a cut off date at 1999 by Ganzon-Fortes (2012). The last mentioned literature compiled a list of 949 algal species, including 61 Cyanobacteria, 212 Chlorophytes, 154 Ochrophytes (Phaeophyceaens) and 522 Rhodophytes. Several recent works since 1999 have contributed to the long list of Philippine marine algal records. Notable of these included the works of Geraldino et al. (2005) on Padina, Belleza and Liao (2007) on Caulerpa, Geraldino et al. (2010) on Hypnea and Payo et al. (2013) on Portieria. With the extensive application of molecular techniques in studies on algal systematics in recent years, major revisions have been made in many groups of tropical marine algae. There is therefore a need to update the nomenclatures of Philippine marine algal records. This nomenclature validation is important to algal biodiversity, conservation and biogeographical studies in order to provide a more accurate picture of floristic connection and the relatedness of Philippine flora with those of other neighbouring regions. The main objective of this current work is therefore to compile all known records of Philippine macroalgae from the literature till April of

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Philippine Journal of Science142: 5-49, Special IssueISSN 0031 - 7683Date Received: 2 May 2013

Marine Science LaboratorySchool of Life Sciences, Chinese University of Hong Kong

Shatin, N.T., Hong Kong SAR, CHINA

Put O. Ang, Jr., Sin Man Leung, and Mei Mei Choi

A Verification of Reports of Marine Algal Species from the Philippines

Corresponding Author: [email protected]

Records of marine macroalgae reported from the Philippines were checked against AlgaeBase, the international database for algal nomenclatures, and Index Nominum Algarum (INA) Bibliographia Phycologica Universalis of the University of California at Berkeley Silva Center for Phycological Documentation to verify their present nomenclature, status of taxonomy and bibliographic reference. To date, 306 names of taxa (including species, varieties and forms) of greens (Chlorophyta), 234 names of taxa of browns (Ochrophyta, Phaeophyceae) and 751 names of taxa of reds (Rhodophyta), or a total of 1291 published names of taxa have been reported from the Philippines. Of these, 231 taxa representing 197 species in 20 families for green algae, 171 taxa representing 153 species in 10 families for brown algae, and 564 taxa representing 543 species in 52 families for red algae are considered valid records listed with their currently accepted names. All in all, 966 currently accepted taxa, representing 893 species in 82 families of marine macroalgae have been reported from the Philippines. Among the greens, 15 taxa have their type localities in the Philippines. This number is 40 for the browns and 33 for the reds. Proportionally, this is 6.5% of the total for the greens, 23.4% for the browns and 5.9% for the reds.

Key Words: Philippine marine macroalgae, algal taxonomic status, algal type species from the Philippines, AlgaeBase, Index Nominum Algarum (INA)

INTRoDUCTIoN Philippines is known for its rich marine flora. over the years, several efforts have been attempted to compile and annotate the records of marine algae collected from the Philippines, including those by Silva et al. (1987), the two volume works by Trono (1997, 2004), and more recently, the checklist with a cut off date at 1999 by Ganzon-Fortes (2012). The last mentioned literature compiled a list of 949 algal species, including 61 Cyanobacteria, 212 Chlorophytes, 154 ochrophytes (Phaeophyceaens) and 522 Rhodophytes. Several recent works since 1999 have contributed to the long list of Philippine marine algal records. Notable of these included the works of Geraldino

et al. (2005) on Padina, Belleza and Liao (2007) on Caulerpa, Geraldino et al. (2010) on Hypnea and Payo et al. (2013) on Portieria.

With the extensive application of molecular techniques in studies on algal systematics in recent years, major revisions have been made in many groups of tropical marine algae. There is therefore a need to update the nomenclatures of Philippine marine algal records. This nomenclature validation is important to algal biodiversity, conservation and biogeographical studies in order to provide a more accurate picture of floristic connection and the relatedness of Philippine flora with those of other neighbouring regions. The main objective of this current work is therefore to compile all known records of Philippine macroalgae from the literature till April of

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2013 and check them against AlgaeBase (Guiry and Guiry 2013), the international database of algal nomenclatures, and Index Nominum Algarum (INA) Bibliographia Phycologica Universalis of the University of California at Berkeley Silva Center for Phycological Documentation (Silva 1996-to date) to come up with the most updated Philippine records. Furthermore, species with their type locality within the territorial waters of the Philippines are also checked and listed.

MATeRIALS AND MeTHoDSThis compilation work relied initially on the main cata-logue of marine algae reported from the Philippines by Silva et al. (1987). Further updates were based on works by Trono (1997, 2004) and Ganzon-Fortes (2012). In ad-dition, all relevant, more recent (post 1999) literatures were consulted. In case of doubts, original literatures were also consulted for confirmation of the records. These literatures are all included in the Reference list.

The compiled list of algae was then double checked for duplication. The revised list was further checked for synonymy and invalidity of taxa based on AlgaeBase (Guiry and Guiry 2013) and INA Database (Silva 1996-to date). The most recent information on family clas-sification, name of authorities, and type localities were also checked and accepted to come up with the final list. Where type localities of species are reported to be in the Philippines, the protologues were consulted, or reference was made to Silva (1996-to date), for verification.

ReSULTS AND DISCUSSIoN

This compilation includes only taxa of marine macroal-gae belonging to the three major phyla, the Chlorophyta (Green), ochrophyta (Pheophyceae, Brown) and Rho-dophyta (Red) listed in Tables 1 to 3 respectively. For ease of reference, the scientific names of listed taxa are arranged alphabetically first by the family classification, followed by the generic and species epithets. The abbre-viation of the authority names of all binomials follows that used in AlgaeBase. For consideration of nomencla-tural priority, the original year of publication was added at the end of each currently accepted taxon listed.

In total, 306 published names of taxa (including species, varieties and forms) of greens, 234 names of browns and 751 names of reds, or a total of 1291 names of algal taxa have been reported from the Philippines. of these, there are 231 validly published taxa representing 197 species in 20 families for green algae (Table 1), 171 taxa repre-senting 153 species in 10 families for brown algae (Table

2), and 564 taxa representing 543 species in 52 fami-lies for red algae (Table 3). Among the reds, four taxa are generic records only (Euptilota sp., Callithamnion sp., Dudresnaya sp. and Rhabdonia sp.). These genera are unique records, although with no specific names re-ported for the Philippine specimens, and are therefore counted as part of the 543 species recorded. Family Rhabdoniaceae, where Rhabdonia sp. belongs, also rep-resents a unique family with only one known species re-corded from the Philippines. The family of Callophycus serratus cannot be ascertained, hence, is classified un-der “Rhodophyta incertae sedis” by AlgaeBase. All in all, 966 currently accepted valid taxa, representing 893 species in 82 families of marine macroalgae have been reported from the Philippines.

Synonyms or invalid names recorded for different spe-cies under the three major groups of algae are also listed respectively in Tables 1 to 3, across their currently ac-cepted names. Some invalid names are clearly due to spelling error (e.g. Sargassum siliquosum vs. S. siloquo-sum), some are simply due to incorrect gender agreement with the generic name (e.g. Kappaphycus striatus vs. K. striatum). These variations of nomenclature in print are listed here for record purposes. Among the greens, Enteromorpha aragoensis Bliding is considered illegiti-mate due to lack of a type citation (Table 1). Under each phylum, there are published records of taxa which have no matching records in AlgaeBase. These are separately listed. These taxa have also been checked against INA Database. Those which are not also found in INA Da-tabase probably represent invalid names. Furthermore, some taxa are classified under “Preliminary Entry” in Al-gaeBase. These are taxa where detailed studies on their taxonomic status are not available. These are all listed in the respective Tables for record purposes and are not included in the total count of species.

Among the greens, 15 taxa have their type localities in the Philippines. This number is 40 for the browns and 33 for the reds. Proportionally, this is 6.5% of the total for the greens, 23.4% for the browns and 5.9% for the reds. The high proportion of type species (taxa) in browns is mainly due to the relatively high number of Sargassum taxa reported as being new to science by Grunow (1915, 1916) (16 taxa) and Trono (1994) (12 species). Two taxa (Sargassum cinctum var. jagorii Grunow and S. crassi-folium var. subduplicatum (‘subduplicata’) Grunow) de-scribed by Grunow (1915) based on materials from the Philippines are not recorded in AlgaeBase (see Table 2). To include these would increase the proportion of brown algae with type or syntype locality in the Philippines to 24.3%. on the other hand, the green alga Cladophora

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Table 1. Listing of green (Chlorophyta) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed.

List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality

1 Anadyomenaceae Anadyomene eseptata Gilbert 1961Dalupiri Island, Babuyan Islands, Cagayan, Luzon, Philippines (Gilbert 1961: 425).

2 Anadyomene leclancheri Decaisne 1844 Microdictyon clathratum G. Martens

Syntype localities: Zamboanga, Min-danao, Philippines; Seram and Flores, Indonesia (Martens 1866: 66-67).

3 Anadyomene plicata C. Agardh 1823

4 Anadyomene stellata (Wulfen) C. Agardh 1823

Anadyomene brownii Schroder

5 Anadyomene wrightii Harvey ex J. e. Gray 1866

6 Microdictyon boergesenii Setchell 1925

7 Microdictyon japonicum Setchell 1925

8 Microdictyon okamurae Setchell 1925

9 Microdictyon tenuius J. e. Gray 1866 Microdictyon agardhianum Decaisne

10 Boodleaceae Boodlea coacta (Dickie) G. Murray & De Toni 1889

11 Boodlea composita (Harvey) F. Brand 1904

12 Boodlea montagnei (Harvey ex J. e. Gray) egerod 1952

13 Boodlea struveoide M. A. Howe 1918

14 Cladophora herpestica (Montagne) Kützing 1849

Cladophoropsis zollingeri (Kützing) Reinbold

15 Cladophoropsis fasciculata (Kjellman) Wille 1910

Cladophoropsis sundanensis Reinbold

16 Cladophoropsis gracillima e. Y. Dawson 1950

17 Cladophoropsis membranacea (Hofman Bang ex C. Agardh) Børgesen 1905

18 Cladophoropsis philippinensis W. R. Taylor 1961

Little Santa Cruz Island, opposite Zamboanga, The Philippines (Taylor 1961: 58).

19 Cladophoropsis vaucheriiformis (Areschoug) Papenfuss 1958

Cladophoropsis dichotoma (Zanardini) Papenfuss; Cladophoropsis neocaledon-ica (Grunow ex G. Murray & Boodle) Papenfuss

20 Phyllodictyon anastomosans (Harvey) Kraft & M. J. Wynne 1996

Struvea anastomosans (Harvey) Piccone & Grunow ex Piccone

21 Phyllodictyon pulcherrimum J. e. Gray 1866

Microdictyon curtissiae W. R. Taylor; Struvea ramosa Dickie

22 Struvea okamurae Leliaert 2007 Chamaedoris orientalis okamura & Higashi

23 Bryopsidaceae Bryopsis corticulans Setchell 1899

24 Bryopsis indica A. Gepp & e. S. Gepp 1908

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25 Bryopsis pennata J. V. Lamouroux 1809

26 Bryopsis pennata var. secunda (Harvey) Collins & Hervey 1917

27 Bryopsis plumosa (Hudson) C. Agardh 1823

28 Caulerpaceae Caulerpa antoensis Yamada 1940 Caulerpa arenicola W. R. Taylor

29 Caulerpa brachypus Harvey 1860

30 Caulerpa brachypus f. parvifolia (Harvey) A. B. Cribb 1958 Caulerpa parvifolia Harvey

31 Caulerpa cupressoides (M. Vahl) C. Agardh 1817

32 Caulerpa cupressoides f. amicorum (Harvey) Weber-van Bosse 1898

33 Caulerpa cupressoides f. disticha Weber-van Bosse 1898

34Caulerpa cupressoides var. [lycopodium] f. elegans (P. L. Crouan & H. M. Crouan) Weber-van Bosse 1898

Caulerpa cupressoides var. lycopodium f. elegans (P. L. & H. M. Crouan) Weber-van Bosse

35 Caulerpa cupressoides var. ericifolia (Turner) Weber-van Bosse 1898

36 Caulerpa cupressoides var. lycopodium Weber-van Bosse 1898

37Caulerpa cupressoides var. urvilleana (Montagne) L. J. Hodgson, Pham Huu Tri, Lewmanomont & McDermid 2004

Caulerpa urvilleana Mon-tagne

38 Caulerpa elongata Weber-van Bosse 1898

39 Caulerpa fastigiata Montagne 1837

40 Caulerpa fergusonii G. Murray 1891

41 Caulerpa filicoides Yamada 1936

42 Caulerpa lentillifera J. Agardh 1837

43 Caulerpa lentillifera var. compacta G. C. Trono & Ang 1982

Bugsuk, Palawan, Philippines (Trono and Ang 1982: 7)

44 Caulerpa lessonii Bory de Saint-Vincent 1828

45 Caulerpa mexicana Sonder ex Kützing 1849

46 Caulerpa mexicana var. pluriseriata W. R. Taylor 1975

47 Caulerpa microphysa (Weber-van Bosse) Feldmann 1955

48 Caulerpa peltata J. V. Lamouroux 1809

Caulerpa racemosa var. laetevirens (Montagne) Weber-van Bosse; Caulerpa racemosa var. peltata (J. V. Lamouroux) eubank

49 Caulerpa peltata var. macrodisca (Decaisne) Weber-van Bosse 1898

Caulerpa macrodisca Decaisne

50 Caulerpa prolifera (Forsskål) J. V. Lamouroux 1809

51 Caulerpa racemosa (Forsskål) J. Agardh 1873

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52 Caulerpa racemosa var. lamourouxii (Turner) Weber-van Bosse 1898

53 Caulerpa racemosa var. macra Weber-van Bosse 1913

54 Caulerpa racemosa var. macrophysa (Sonder ex Kützing) W. R. Taylor 1928

55 Caulerpa racemosa var. occidentalis (J. Agardh) Børgesen 1907

56 Caulerpa racemosa var. turbinata (J. Agardh) eubank 1946

57 Caulerpa reyesii Menez & H. P. Calumpong 1982

58 Caulerpa selago (Turner) C. Agardh 1817

59 Caulerpa serrulata (Forsskål) J. Agardh 1837

60 Caulerpa serrulata f. lata (Weber-van Bosse) C. K. Tseng 1936

61 Caulerpa serrulata f. spiralis (Weber-van Bosse) Gilbert 1942

62 Caulerpa serrulata var. boryana (J. Agardh) Gilbert 1942

63 Caulerpa serrulata var. boryana f. longifolia Gilbert 1942

64Caulerpa serrulata var. boryana f. oc-cidentalis (Weber-van Bosse) Yamada & Tanaka 1938

65 Caulerpa serrulata var. pectinata (Weber-van Bosse) W. R. Taylor 1960

66 Caulerpa sertularioides (S. G. Gmelin) M. A. Howe 1905

67 Caulerpa sertularioides f. brevipes (J. Agradh) Svedelius 1906

68 Caulerpa sertularioides f. farlowii (Weber-van Bosse) Børgesen 1907

69 Caulerpa sertularioides f. longipes (J. Agardh) Collins 1909

70 Caulerpa sertularioides f. longiseta (Bory de Saint-Vincent) Svedelius 1906

71 Caulerpa subserrata okamura 1897

72 Caulerpa taxifolia (M. Vahl) C. Agardh 1817

73 Caulerpa urvilleana var. vitiensis Weber-van Bosse 1898

74 Caulerpa verticillata J. Agardh 1847

75 Caulerpa vesiculifera (Harvey) Harvey 1863

76 Caulerpa webbiana Montagne 1837

77 Caulerpa webbiana var. pickeringii (Har-vey & Bailey) eubank 1946

78 Caulerpella ambigua (okamura) Prud'homme van Reine & Lokhorst 1992 Caulerpa ambigua okamura

79 Cladophoraceae Chaetomorpha aerea (Dillwyn) Kützing 1849

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80 Chaetomorpha antennina (Bory de Saint-Vincent) Kützing 1847

81 Chaetomorpha brachygona Harvey 1858

82 Chaetomorpha crassa (C. Agardh) Kützing 1845

83 Chaetomorpha gracilis Kützing 1845

84 Chaetomorpha inflata Kützing 1849

85 Chaetomorpha kellersii M. A. Howe 1932 Panay Island, Philippines (Howe 1932: 169)

86 Chaetomorpha ligustica (Kützing) Kützing 1849

87 Chaetomorpha linum (o. F. Müller) Kützing 1845

88 Chaetomorpha philippinensisLeliaert 2011

Dapdap, Siquijor,Philippines (Leliaert etal. 2011: 385)

89 Chaetomorpha spiralis okamura 1903

90 Chaetomorpha vieillardii(Kützing) M. J. Wynne 2011

91 Cladophora albida (Nees) Kützing 1843

92 Cladophora aokii Yamada 1925

93 Cladophora catenata (Linnaeus) Kützing 1843

94 Cladophora conferta P. L. Crouan & H. M. Crouan 1865

95 Cladophora crispula Vickers 1905

96 Cladophora crucigera Grunow 1867

97 Cladophora cymopoliae Børgesen 1925

98 Cladophora dalmatica Kützing 1843

99 Cladophora japonica Yamada 1931

100 Cladophora laetevirens (Dillwyn) Kützing 1843

101 Cladophora liebetruthii Grunow 1884

102 Cladophora pellucida (Hudson) Kützing 1843

103 Cladophora prolifera (Roth) Kützing 1843

104 Cladophora quisumbingii Manza 1939 Batan Island, Batanes, Luzon, Philip-pines (Manza 1939: 109)

105 Cladophora rupestris (Linnaeus) Kützing 1843

106 Cladophora sakaii I. A. Abbott 1972

107 Cladophora sericea (Hudson) Kützing 1843

Cladophora crystallina (Roth) Kützing

108 Cladophora sibogae Reinbold 1905

109 Cladophora vagabunda (Linnaeus) Hoek 1963 Cladophora inserta Dickie

110 Cladophora wrightiana Harvey 1860

111 Rhizoclonium africanum Kützing 1853

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112 Rhizoclonium crassipellitum var. robustum G. S. West 1904

113 Rhizoclonium grande Børgesen 1935

114 Rhizoclonium riparium (Roth) Harvey 1849 Rhizoclonium kerneri Stock-mayer

115 Rhizoclonium riparium var. implexum (Dillwyn) Rosenvinge 1893

Rhizoclonium kochianum Kützing

116 Rhizoclonium setaceum Kützing 1847

117 Codiaceae Codium arabicum Kützing 1856

118 Codium bartlettii C. K. Tseng & W. J. Gilbert 1942

Puerto Galera Bay, Mindoro oriental, Philippines (Tseng and Gilbert 1942: 291-293).

119 Codium contractum Kjellman 1857

120 Codium cylindricum Holmes 1896

121 Codium decorticatum (Woodward) M. A. Howe 1911

Codium elongatum Aucto-rum

122 Codium edule P. C. Silva 1952

123 Codium fragile (Suringar) Hariot 1889

124 Codium geppiorum o. C. Schmidt 1923 Codium geppii o. C. Schmidt

125 Codium intricatum okamura 1913

126 Codium muelleri Kützing 1856

127 Codium ovale Zanardini 1878

128 Codium papillatum C. K. Tseng & W. J. Gilbert 1942

Puerto Galera, Mindoro oriental, Philippines (Tseng and Gilbert 1942: 293-295).

129 Codium platyclados R. Jones & Kraft 1984

130 Codium pungniforme okamura 1915

131 Codium tenue (Kützing) Kützing 1856

132 Codium tomentosum Stackhouse 1797

133 Dasycladaceae Bornetella nitida Munier-Chalmas ex Sonder 1880

134 Bornetella oligospora Solms-Laubach 1892

135 Bornetella sphaerica (Zanardini) Solms-Laubach 1892 Bornetella ovalis Yamada

136 Chloroclados australasicus Sonder 1871

Chloroclados philippinensis Gilbert; Chlorocladus australasicus Sonder; Chlorocladus philippinensis Gilbert

137 Cymopolia van-bosseae Solms-Laubach 1892

Cymopolia vanbosseae Solms-Laubach

138 Dasycladus vermicularis (Scopoli) Krasser 1898

139 Neomeris annulata Dickie 1874

140 Neomeris bilimbata Koster 1937

141 Neomeris vanbosseae M. A. Howe 1909

142 Derbesiaceae Derbesia attenuata Funk 1955

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143 Derbesia marina (Lyngbye) Solier 1846

144 Derbesia tenuissima (Moris & De Notaris) P. L. Crouan & H. M. Crouan 1867

145 Pedobesia ryukyuensis (Yamada & T. Tanaka) Kobara & Chihara 1984

146 Dichotomosiphona-ceae

Avrainvillea erecta (Berkeley) A. Gepp & e. S. Gepp 1911

Philippine Islands (Berkeley 1842: 157).

147 Avrainvillea lacerata Harvey ex J. Agardh 1887

148 Avrainvillea longicaulis (Kützing) G. Murray & Boodle 1889

149 Avrainvillea nigricans Decaisne 1842

150 Avrainvillea obscura (C. Agardh) J. Agardh 1887

Avrainvillea capituliformis Tanaka

151 Gayraliaceae Gayralia oxysperma (Kützing) K. L. Vinogradova ex Scagel 1989

Monostroma oxyspermum (Kützing) Doty

152 Gomontiaceae Monostroma latissimum Wittrock 1866

153 Monostroma nitidum Wittrock 1866 Porphyra crispata Kjellman

154 Halimedaceae Halimeda copiosa Goreau & e. A. Graham 1967

155 Halimeda cuneata Hering 1846

156 Halimeda cylindracea Decaisne 1842

157 Halimeda discoidea Decaisne 1842

158 Halimeda fragilis W. R. Taylor 1950

159 Halimeda gigas W. R. Taylor 1950

160 Halimeda goreauii W. R. Taylor 1962

161 Halimeda gracilis Harvey ex J. Agardh 1887

162 Halimeda incrassata (J. ellis) J. V. Lamouroux 1816

163 Halimeda lacunalis W. R. Taylor 1950

164 Halimeda macroloba Decaisne 1841

165 Halimeda macrophysa Askenasy 1888

166 Halimeda micronesica Yamada 1941 Halimeda orientalis Gilbert

167 Halimeda opuntia (Linnaeus) J. V. Lamouroux 1816

168 Halimeda renschii Hauck 1886 Halimeda batanensis W. R. Taylor

169 Halimeda simulans M. A. Howe 1907

170 Halimeda stuposa W. R. Taylor 1950

171 Halimeda taenicola W. R. Taylor 1950

172 Halimeda tuna (J. ellis & Solander) J. V. Lamouroux 1816

173 Halimeda velasquezii W. R. Taylor 1962 Sta. Ana, Cagayan, Luzon Island, Philippines (Taylor 1962: 177).

174 Halimeda velasquezii var. robusta P. A. Cordero 1974

Cadadalman, Camiguin Island, Cagayan, Philippines (Cordero 1974: 115).

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175 Ostreobiaceae Ostreobium quekettii Bornet & Flahault 1889

176 Phaeophilaceae Phaeophila dendroides (P. L. Crouan & H. M. Crouan) Batters 1902

177 Polyphysaceae Acetabularia caliculus J. V. Lamouroux 1824

178 Acetabularia crenulata J. V. Lamouroux 1816

179 Acetabularia dentata Solms-Laubach 1895

180 Acetabularia major G. Martens 1866

181 Acetabularia ryukyuensis okamura & Yamada 1932

Acetabularia philippinensis Gilbert Acetabularia roxasii G. C. Trono, Santiago & Ganzon-Fortes Acetabularia ryukyuensis var. philippinensis (Gilbert) Valet

182 Halicoryne wrightii Harvey 1860

183

Parvocaulis clavatus (Yamada) S. Berger, U. Fettweiss, S.Gleissberg, L. B. Liddle, U.Richter, H. Sawitzky & G. C.Zuccarello 2003

Acetabularia clavata Yamada

184

Parvocaulis parvulus (Solms-Laubach) S. Berger, U. Fettweiss, S. Gleissberg, L. B. Liddle, U. Richter, H. Sawitzky & G. C. Zuccarello 2003

Acetabularia minutissima okamura; Acetabularia moebii Solms-Laubach; Acetabularia parvula Solms-Laubach; Acetabularia velasquezii G. C. Trono, Santiago & Ganzon-Fortes

185

Parvocaulis polyphysoides (P. L. Crouan & H. M. Crouan) S. Berger, U. Fettweiss, S. Gleissberg, L. B. Liddle, U. Richter, H. Sawitzky & G.C. Zuccarello 2003

Acetabularia exigua Solms-Laubach

186 Rhipiliaceae Rhipilia crassa Millar & Kraft 2001

187 Rhipilia orientalis A. Gepp & e. S. Gepp 1911

188 Rhipilia tomentosa Kützing 1858

189 Rhipiliopsis carolyniae Kraft 1986 Bulusan, Sorsogon, Philippines (Kraft 1986: 51)

190 Siphonocladaceae Boergesenia forbesii (Harvey) Feldmann 1938

191 Dictyosphaeria cavernosa (Forsskål) Børgesen 1932

192 Dictyosphaeria cavernosa var. bullata Børgesen 1952

193 Dictyosphaeria intermedia Weber-van Bosse 1905

194 Dictyosphaeria ocellata (M. A. Howe) olsen-Stojkovich 1985

195 Dictyosphaeria versluysii Weber-van Bosse 1905

196 Ernodesmis verticillata (Kützing) Børgesen 1912

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197 Udoteaceae Boodleopsis pusilla (F. S. Collins) W. R. Taylor, A. B. Joly & Bernatowicz 1953

198 Boodleopsis verticillata e. Y. Dawson 1960

199 Chlorodesmis caespitosa J. Agardh 1887

200 Chlorodesmis fastigiata (C. Agardh) S. C. Ducker 1969

Chlorodesmis comosa Har-vey & Bailey

201 Chlorodesmis hildebrandtii A. Gepp & e. S. Gepp 1911

202 Chlorodesmis major Zanardini 1874

203 Rhipidosiphon javensis Montagne 1842 Udotea javensis (Montagne) A. Gepp & e. S. Gepp

204 Tydemania expeditionis Weber-van Bosse 1901

205 Udotea argentea Zanardini 1858

206 Udotea argentea var. spumosa A. Gepp & e. S. Gepp 1911

Syntype localities: Tanah Djampeah and Salayer Islands, Indonesia; Pearl Bank, Tawitawi, Sulu Archipelago, Philippines (Coppejans et al. 2001: 429)

207 Udotea flabellum (J. ellis & Solander) M. A. Howe 1904

208 Udotea geppiorum Yamada 1930 Udotea geppii Yamada

209 Udotea glaucescens Harvey ex J. Agardh 1887

210 Udotea indica A. Gepp & e. S. Gepp 1911

211 Udotea occidentalis A. Gepp & e. S. Gepp 1911

212 Udotea orientalis A. Gepp & e. S. Gepp 1911

Syntype localities: various in Indian and Pacific oceans; Indonesia; Philip-pine Islands (Gepp and Gepp 1911: 119-120).

213 Ulvaceae Ulva australis Areschoug 1854 Ulva pertusa Kjellman

214Ulva chaetomorphoides (Børgesen) Hayden, Blomster, Maggs, P. C. Silva, M. J. Stanhope & J. R. Waaland 2003

Enteromorpha chaetomor-phoides Børgesen

215 Ulva clathrata (Roth) C. Agardh 1811

Enteromorpha clathrata (Roth) Greville; Enteromorpha ramulosa (Smith) Carmichael

216 Ulva compressa Linnaeus 1753 Enteromorpha compressa (Linnaeus) Nees

217 Ulva flexuosa Wulfen 1803

Enteromorpha flexuosa (Wulfen) J. Agardh; Enteromorpha lingulata J. Agardh; Enteromorpha tubulosa (Kützing) Kützing

218 Ulva flexuosa f. submarina (F. S. Collins & Hervey) M. J. Wynne 2005

Enteromorpha flexuosa f. submarina Collins & Hervey

219 Ulva flexuosa subsp. paradoxa (C.Agardh) M. J. Wynne 2005

Enteromorpha flexuosa subsp. paradoxa (C. Agardh) Bliding

220 Ulva intestinalis Linnaeus 1753 Enteromorpha intestinalis (Linnaeus) Nees

221Ulva kylinii (Bliding) Hayden, Blomster, Maggs, P. C. Silva, M. J. Stanhope & J. R.Waaland 2003

Enteromorpha kylinii Bliding

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222 Ulva lactuca Linnaeus 1753 Ulva fasciata Delile

223 Ulva prolifera o. F. Müller 1778 Enteromorpha prolifera (o. F. Müller) J. Agardh

224 Ulva reticulata Forsskål 1775

225 Ulva rigida C. Agardh 1823

226 Valoniaceae Valonia aegagropila C. Agardh 1823

227 Valonia fastigiata Harvey ex J. Agardh 1887

228 Valonia macrophysa Kützing 1843

229 Valonia utricularis (Roth) C. Agardh 1823

230 Valonia ventricosa J. Agardh 1887 Ventricaria ventricosa (J. Agardh) olsen & J. A. West

231 Valoniopsis pachynema (G. Martens) Børgesen 1934

Valonia confervoides Za-nardini

Invalid Species Name

Family Taxon Recorded Remarks Reference

Ulvaceae Enteromorpha aragoensis Bliding 1960, nom. illeg.

*Enteromopha has now been included under Ulva. This species is currently consid-ered illegitimate because no type material was designated.

Silva et al. 1987;Silva 1996-to date

Species with No Matching Record in AlgaeBase

Family Taxon Recorded Remarks Reference

Caulerpaceae Caulerpa cupressoides var. cupressoides Not listed also in INA Database

Belleza and Liao 2007

Caulerpa cupressoides var. lycopodium f. disticha Weber-van Bosse

Same as Caulerpa cupres-soides f. disticha Weber-van Bosse 1898

Belleza and Liao 2007

Caulerpa mexicana var. mexicana Not listed also in INA Database Belleza and Liao 2007

Caulerpa racemosa var. racemosa (Forsskål) J. Agardh

Listed in INA Database as Caulerpa racemosa var. racemosa J. Agardh

Caulerpa racemosa var. serrulata (Fors-skål) J. Agardh

Not listed also in INA Database

Caulerpa serrulata var. serrulata f. lata (Weber-van Bosse) C. K. Tseng

Same as Caulerpa serrulata f. lata (Weber-van Bosse) C. K. Tseng

Belleza and Liao 2007

Caulerpa sertularioides var. sertularioides f. brevipes (J. Agardh) Svedelius

Same as Caulerpa sertulari-oides f. brevipes (J. Agardh) Svedelius

Belleza and Liao 2007

Cladophoraceae Cladophora filipendula No authority of taxon listed;Not listed also in INA Database

Ganzon-Fortes 2012

Species Listed as Preliminary Entry in AlgaeBase

Family Taxon Recorded Remarks Reference

Caulerpaceae Caulerpa sertularioides f. flagellata (Weber-van Bosse) Weber-van Bosse

Listed in INA Database Silva et al. 1987

Cladophoraceae Cladophora luzoniensis V. Martens Listed in INA Database; Type locality: San Mateo River, Luzon, Philippines (Martens 1866: 20)

Guiry and Guiry 2013;Silva 1996-to date

Halimedaceae Halimeda bikinensis W. R. Taylor Listed in INA Database Ganzon-Fortes 2012

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Table 2. Listing of brown (ochrophyta, Phaeophyceae) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed.

List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality

1 Acinetosporaceae Asteronema breviarticulatum (J. Agardh) ouriques & Bouzon 2000

Hincksia breviarticulata (J. Agardh) P. C. Silva

2 Ectocarpus rallsiae Vickers 1905 Hincksia rallsiae (Vickers) P. C. Silva

3 Feldmannia columellaris (Børgesen) Islam 1976

4 Feldmannia formosana (Yamada) Itono 1973

5 Feldmannia indica (Sonder) Womers-ley & A. Bailey 1970

6 Feldmannia irregularis (Kützing) G. Hamel 1939

7 Feldmannia mitchelliae (Harvey) H. S. Kim 2010

Hincksia mitchelliae (Harvey) P. C. Silva

8 Chordariaceae Leathesia marina (Lyngbye) Decaisne 1842 Leathesia difformis Areschoug

9 Cutleriaceae Mutimo cylindricus (okamura) H. Kawai & T. Kitayama 2012 Cutleria cylindrica okamura

10 Dictyotaceae Canistrocarpus cervicornis (Kützing) De Paula & De Clerck 2006

Dictyota cervicornis Kützing; Dictyota indica Sonder ex Kützing

11 Dictyopteris camiguinensis Tanaka 1967

San Pio Quinto, Camiguin Island, Ba-buyan Islands, Cagayan, Luzon, The Philippines (Tanaka 1967: 16)

12 Dictyopteris delicatula J. V. Lamour-oux 1809

13 Dictyopteris divaricata (okamura) okamura 1932

14 Dictyopteris jamaicensis W. R. Taylor 1960

15 Dictyopteris pacifica (Yendo) I. K. Hwang, H. S. Kim & W. J. Lee 2004 Spatoglossum pacificum Yendo

16 Dictyopteris polypodioides (A. P. De Candolle) J. V. Lamouroux 1809

17 Dictyopteris repens (okamura) Børge-sen 1924

18 Dictyopteris undulata Holmes 1896

19 Dictyota bartayresiana J. V. Lamour-oux 1809 Dictyota patens J. Agardh

20 Dictyota ceylanica Kützing 1859

21 Dictyota ciliolata Sonder ex Kützing 1859

22 Dictyota crenulata J. Agardh 1847 Dictyota bartayresiana var. denticulata Kützing

23 Dictyota crispata J. V. Lamouroux 1809 Dictyota bidentata Harvey & Bailey

24 Dictyota dichotoma (Hudson) J. V. Lamouroux 1809

25 Dictyota friabilis Setchell 1926

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26 Dictyota implexa (Desfontaines) J. V. Lamouroux 1809

Dictyota divaricata J. V. Lamouroux Dictyota linearis (C. Agardh) Greville

27 Dictyota lata J. V. Lamouroux 1809

28 Dictyota major W. R. Taylor 1945

29 Dictyota mertensii (Martius) Kützing 1859 Dictyota dentata J. V. Lamouroux

30Lobophora variegata (J. V. Lamou-roux) Womersley ex e. C. oliveira 1977

Pocockiella variegata (J. V. Lamou-roux) Papenfuss

31 Padina antillarum (Kützing) Piccone 1886

32 Padina arborescens Holmes 1896

33 Padina australis Hauck 1887

34 Padina boryana Thivy ex W. R. Taylor 1966

35 Padina distromatica Hauck 1887

36 Padina fernandeziana Skottsberg & Levring 1941

37 Padina fraseri (Greville) Greville 1830

38 Padina gymnospora (Kützing) Sonder 1871 Padina crassa Yamada

39 Padina japonica Yamada 1931

40 Padina jonesii Tsuda 1972

41 Padina minor Yamada 1925

42 Padina moffittiana I. A. Abbott & Huisman 2003

43 Padina pavonica (Linnaeus) Thivy 1960

44 Padina sanctae-crucis Børgesen 1914

45 Padina tetrastromatica Hauck 1887

46Rugulopteryx okamurae (e. Y. Daw-son) I. K. Hwang, W. J. Lee & H. S. Kim 2009

Dilophus okamurae e. Y. Dawson

47 Spatoglossum asperum J. Agardh 1894

48 Spatoglossum variabile Figari & De Notaris 1853

49 Stypopodium flabelliforme Weber-van Bosse 1913

50 Stypopodium zonale (J. V. Lamouroux) Papenfuss 1940

Stypopodium lobatum (C. Agardh) Kützing

51 Zonaria diesingiana J. Agardh 1841

52 Zonaria flabellata (okamura) Papen-fuss 1944

53 LaminariaceaeSaccharina latissima (Linnaeus) C. e. Lane, C. Mayes, Druehl & G. W. Saunders 2006

Ulva latissima Linnaeus

54 Mesosporaceae Mesospora negrosensis J. A. West & H. P. Calumpong 1996

Lalaan, San Jose, Negros oriental, Phil-ippines (West and Calumpong 1996: 5).

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55 Ralfsiaceae Ralfsia fungiformis (Gunnerus) Setch-ell & N. L. Gardner 1924

56 Ralfsia verrucosa (Areschoug) Are-schoug 1845

57 Sargassaceae Hormophysa cuneiformis (J. F. Gme-lin) P. C. Silva 1987

Hormophysa triquetra (C.Agardh) Kützing

58 Myagropsis myagroides (Mertens ex Turner) Fensholt 1955

59 Sargassum abbottiae G. C. Trono 1994 Pulong Bakaw, Calatagan, Batangas, Philippines (Trono 1994: 5).

60 Sargassum agardhianum Farlow 1889

61 Sargassum angii L. M. Liao 1998 Sargassum yamadae G. C. Trono Barrio Pangil, Currimao, Ilocos Norte, Philippines (Trono 1994: 4).

62 Sargassum aquifolium (Turner) C. Agardh 1820

Sargassum binderi Sonder ex J. Agardh; Sargassum biserrula J.Agardh; Sargassum crassifolium J. Agardh; Sargassum heterocystum (Kützing) Montagne; Sargassum oocyste J. Agardh

63 Sargassum baccularia (Mertens) C. Agardh 1824

64 Sargassum balingasayense G. C. Trono 1994

Barrio Balingasay, Bolinao, Pangas-inan, Philippines (Trono 1994: 5).

65 Sargassum bataanense G. C. Trono 1994

Pulong Bato, orion, Bataan, Philippines (Trono 1994: 6).

66 Sargassum belangeri Bory de Saint-Vincent 1834

Syntype localities: Sunda Strait and Java, Indonesia; Philippine Islands (Silva et al. 1996: 659).

67 Sargassum cervicorne Greville 1849 Sargassum binderi var. angustifolium Sonder

68 Sargassum cinctum J. Agardh 1848

69 Sargassum cinctum var. gracilentum Grunow 1915

Sargassum cinctum var. gracilenta Grunow

Syntype localities: Coast of Australia: Goodie Isl., Balina, Rockinham; near Manila, Philippines (Grunow 1915: 419).

70 Sargassum cinctum var. mixtum Grunow 1915 Sargassum cinctum var. mixta Grunow Near Manila, Luzon, Philippines (Grun-

ow 1915: 419).

71 Sargassum confusum C. Agardh 1824

72 Sargassum corderoi R. B. Modelo Jr., I. Umezaki & L. M. Liao 1998

Polo, New Washington, Aklan, Panay Is., Philippines (Modelo et al. 1998: 79).

73 Sargassum crispifolium Yamada 1931

74 Sargassum currimaoense G. C. Trono 1994

Pangil, Currimao, Ilocos Norte, Philip-pines (Trono 1994: 6).

75 Sargassum cystocarpum C. Agardh 1820

76 Sargassum cystophyllum Montagne 1842

Manila, Luzon, Philippines (Montagne 1842: 249).

77 Sargassum distichum Sonder 1845 Sargassum aemulum Sonder

78 Sargassum dotyi G. C. Trono 1994 Honduras, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 3).

79Sargassum droserifolium var. spathu-latum Grunow 1915

Sargassum droserifolium var. spathu-lata Grunow

Strait Bernardino, Philippines (Grunow 1915: 411).

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80 Sargassum duplicatum var. rotunda-tum Grunow 1915

Sargassum duplicatum var. rotundata Grunow

Syntype localities: Philippine Islands; Java, Indonesia; Seychelles (Grunow 1915: 392-393).

81 Sargassum esperi C. Agardh 1820

82 Sargassum feldmannii Pham-Hoang Ho 1967

83 Sargassum filifolium (C. Agardh) C. Agardh 1824

84 Sargassum filiforme Montagne 1844 "in insulis Philippinensibus” (Montagne 1844: 660) (Silva et al. 1996: 672).

85 Sargassum fluitans (Børgesen) Børge-sen 1914

86 Sargassum fulvellum (Turner) C. Agardh 1820

87 Sargassum furcatum Kützing 1843

88 Sargassum gaudichaudii Montagne 1842

Syntype localities: Mauritius; Philip-pine Islands (Montagne 1842: 249) (Silva et al. 1996: 673).

89 Sargassum giganteifolium Yamada 1925

90 Sargassum gracile J. Agardh 1848

91 Sargassum gracillimum Reinbold 1913

92 Sargassum granuliferum C. Agardh 1820

93 Sargassum hemiphyllum (Turner) C. Agardh 1820

94 Sargassum hombronianum var. ma-nilense (Grunow) Grunow 1916

Sargassum polycystum var. manilense Grunow

Cavite, Luzon, Philippines (Grunow 1916: 8)

95 Sargassum horneri (Turner) C. Agardh 1820 Sargassum filicinum Harvey

96 Sargassum ilicifolium (Turner) C. Agardh 1820

Sargassum berberifolium J. Agardh; Sargassum cristaefolium C. Agardh; Sargassum droserifolium Bory de Saint-Vincent; Sargassum duplicatum Bory de Saint-Vincent; Sargassum duplicatum (J. Agardh) J. Agardh; Sargassum sandei Reinbold; Sargassum turbinatifolium C.K.Tseng & Lu

97 Sargassum ilicifolium f. benkulense Grunow 1915

Sargassum ilicifolium f. benkulensis Grunow

Syntype localities: Pulau Tikus, near Bengkulu, Sumatra, Indonesia; Philip-pine Islands (Grunow 1915: 407).

98 Sargassum ilicifolium var. compactum (Bory de Saint-Vincent) Grunow 1915

99 Sargassum ilicifolium var. pseudospi-nulosum Grunow 1915

100 Sargassum kushimotense Yendo 1907

101 Sargassum latifolium (Turner) C. Agardh 1820

102 Sargassum microcystum J. Agardh 1848Syntype localities: "in marfi Indico"; Moluccas, Indonesia; Manila, Philip-pines; Singapore (Silva et al. 1996: 685).

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103 Sargassum microcystum f. luzonense Grunow 1915

Sargassum microcystum f. luzonensis Grunow

Near Manila, Luzon, Philippines (Grun-ow 1915: 413)

104 Sargassum microcystum var. euryphyl-lum (Grunow) Grunow 1915

Sargassum myriocystum var. eury-phylla (Grunow) Grunow; Sargassum myriocystum var. euryphyl-lum (Grunow) Grunow; Sargassum polycystum var. euryphyl-lum Grunow

Ticao Island, Masbate, Philippines (Grunow 1915: 440)

105 Sargassum microcystum var. microtis Grunow 1915

Syntype localities: Java, Indonesia; Basilan Strait, Philippines (Grunow 1915: 414).

106 Sargassum microphyllum C. Agardh 1820

107 Sargassum miyabei Yendo 1907

108 Sargassum natans (Linnaeus) Gaillon 1828

109 Sargassum nigrifolium Yendo 1907

110 Sargassum notarisii Zanardini 1858

111 Sargassum ohnoi G. C. Trono 1994 Big Balatero, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 7).

112 Sargassum oligocystum Montagne 1845

113 Sargassum oligocystum var. bernar-dium Grunow 1915

"in freto Bernardino" (San Bernardino Strait between Sorsogon, Luzon, and Northern Samar, Philippines) (Grunow 1915: 386)

114 Sargassum oligocystum var. subflexuo-sum Grunow 1915

Sargassum oligocystum var. subflex-uosa Grunow

“Fretum Bernardium, Macassar” (San Bernardino Strait, Philippines; Makas-sar, Indonesia) (Grunow 1915: 385)

115 Sargassum oocyste var. bernardinum Grunow 1915

Sargassum oocyste var. bernardina Grunow 1915

"in freto Bernardino" (San Bernardino Strait, Philippines)(Grunow 1915: 437).

116 Sargassum paniculatum J. Agardh 1848

117 Sargassum parvifolium (Turner) C. Agardh 1820

118 Sargassum patens C. Agardh 1820

119 Sargassum patens var. schizophyllum (Kützing) Yendo 1905

120 Sargassum philippinense Grunow 1916

San Bernardino Strait, Philippines (Grunow 1916: 166)

121 Sargassum piluliferum (Turner) C. Agardh 1820

122 Sargassum polyceratium Montagne 1837

123 Sargassum polycystum C. Agardh 1824 Sargassum myriocystum J. Agardh

124 Sargassum pteropleuron Grunow 1868 Sargassum pteropleron Grunow

125 Sargassum pulchellum Grunow 1888

126 Sargassum sagamianum Yendo 1907

127 Sargassum samarense G. C. Trono 1994

Borangan, eastern Samar, Philippines (Trono 1994: 4)

128 Sargassum serratifolium (C. Agardh) C. Agardh 1820

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129 Sargassum siliquosum J. Agardh 1848 Sargassum siloquosum J. Agardh

130 Sargassum siliquosum var. basilani-cum Grunow 1916

Sargassum siliquosum var. basilanica Grunow

Basilan Strait (between Mindanao and Basilan Island) (Grunow 1916: 175)

131 Sargassum siliquosum var. bicornutum Grunow 1916

Sargassum siliquosum var. bicornuta Grunow

China Sea; Ticao Island, Masbate, Phil-ippines (Grunow 1916: 175)

132 Sargassum siliquosum var. manipaense Grunow 1916

Sargassum siliquosum var. mani-paensis Grunow;Spelled in AlgaeBase as Sargassum siliquosum var. manipense Grunow

"ad insulas Philippinas, Manipa" (Grun-ow 1916: 175)

133 Sargassum spinifex C. Agardh 1820

134 Sargassum subspathulatum (Grunow) Grunow 1916

135 Sargassum sullivanii G. C. Trono 1994 Palawig, Zambales, Philippines (Trono 1994: 6)

136 Sargassum tenerrimum J. Agardh 1848

137 Sargassum tristichum Sonder 1845 Sargassum biforme Sonder

138 Sargassum turbinarioides Grunow 1915

139 Sargassum umezakii G. C. Trono 1994 Medio Island, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 7)

140 Sargassum velasquezii G. C. Trono 1994

San Isidro, Puerto Galera, Mindoro oriental, Philippines (Trono 1994: 4)

141 Sargassum vulgare C. Agardh 1820

142 Sargassum vulgare var. indicum C. Agardh 1820

143 Sargassum yendoi okamura & Ya-mada 1938

144 Sargassum yoshidae G. C. Trono 1994 Cabugao Bay, Virac, Catanduanes, Philippines (Trono 1994: 5)

145 Sirophysalis trinodis (Forsskål) Kütz-ing 1849

Cystoseira trinodis (Forsskål) C.Agardh

146Stephanocystis hakodatensis (Yendo) Draisma, Ballesteros, F. Rousseau & T. Thibaut 2010

Cystoseira hakodatensis (Yendo) Fensholt

147 Turbinaria condensata Sonder 1860

148 Turbinaria conoides (J. Agardh) Kütz-ing 1860

149 Turbinaria conoides f. laticuspidata W. R. Taylor 1964 Cebu, Philippines (Taylor 1964: 481)

150 Turbinaria conoides f. retroflexa W. R. Taylor 1964

Tandayong Island, Luzon, Philippines (Taylor 1964: 481-482)

151 Turbinaria decurrens Bory de Saint-Vincent 1828

152 Turbinaria denudata Bory de Saint-Vincent 1828

153 Turbinaria filamentosa Yamada 1925

154 Turbinaria luzonensis W. R. Taylor 1964

Mulanay, Luzon, Philippines (Taylor 1964: 482).

155 Turbinaria murrayana e. S. Barton 1891

156 Turbinaria ornata (Turner) J. Agardh 1848

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157 Turbinaria ornata f. evesiculosa (e. S. Barton) W. R. Taylor 1964

158 Turbinaria trialata var. capensis Kütz-ing 1860

159 Turbinaria turbinata (Linnaeus) Kuntze 1898 Turbinaria trialata (J. Agardh) Kützing

160 Turbinaria vulgaris J. Agardh 1848 nom. illeg.

161 Scytosiphonaceae Chnoospora implexa J. Agardh 1848

162 Chnoospora minima (Hering) Papen-fuss 1956

163 Chnoospora pannosa J. Agardh 1848

164 Colpomenia sinuosa (Mertens ex Roth) Derbes & Solier 1851

165 Hydroclathrus clathratus (C. Agardh) M. A. Howe 1920

166 Hydroclathrus tenuis C. K. Tseng & Lu 1983

167 Rosenvingea intricata (J. Agardh) Børgesen 1914

168 Rosenvingea orientalis (J. Agardh) Børgesen 1914

Hab. in mari Indico ad Manillam (Hb. Binder!) [Manila, the Philippines] (Agardh 1848: 78).

169 Sphacelariaceae Sphacelaria novae-hollandiae G. Sonder 1845

170 Sphacelaria rigidula Kützing 1843

171 Sphacelaria tribuloides Meneghini 1840

Species with No Matching Record in AlgaeBase

Family Taxon Recorded Remarks Reference

Sphacelariaceae Sphacelaria mucifera No authority of taxon listed; Also not listed in INA Database Ganzon-Fortes 2012

Sargassaceae Sargassum cinctum var. jagorii Grunow

Type locality: Luzon, Philippines (Grunow 1915: 421)

Grunow 1915; Silva 1996-to date

Sargassum crassifolium var. subdupli-catum ('subduplicata') Grunow

Syntype localities: Makassar, Indo-nesia; Strait Bernardino, Philippines (Grunow 1915: 421)

Grunow 1915; Silva 1996-to date

Species Listed as Preliminary Entry in AlgaeBase

Family Taxon Recorded Remarks Reference

Sargassaceae Sargassum elongatum No authority of taxon listed; Not listed in INA Database

Phillips 1995; Ganzon-Fortes 2012

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Table 3. Listing of red (Rhodophyta) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed.

List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality

1 Acrochaetiaceae Acrochaetium hancockii (e. Y. Daw-son) Papenfuss 1845

2 Acrochaetium liagorae Børgesen 1915

3 Acrochaetium nitidulum I. A. Abbott 1962

Balabac Island, Gnat Reef, Pala-wan, Philippines (Abbott 1962: 95)

4 Acrochaetium papenfussii I. A. Abbott 1962

Cagayan Sulu Island, Palawan, Philippines (Abbott 1962: 97)

5 Acrochaetium robustum Børgesen 1915

6 Acrochaetium sinicola (e. Y. Dawson) Papenfuss 1945

7 Acrochaetium trichogloeae Børgesen 1952

8 Acrochaetium tuticorinense Børgesen 1937

9Colaconema gracile (Børgesen) Ateweberhan & Prud'homme van Reine 2005

Acrochaetium gracile Børgesen

10Colaconema hallandicum (Kylin) Afonso-Carillo, Sanson, Sangil & Diaz-Villa 2007

Acrochaetium hallandicum (Kylin) Hamel

11 Colaconema hypneae (Børgesen) A. A. Santos & C. W. N. Moura 2010 Acrochaetium seriatum Børgesen

12 Ahnfeltiaceae Ahnfeltiopsis concinna (J.Agardh) P. C. Silva & DeCew 1992 Ahnfeltia concinna J. Agardh

13 Bangiaceae Bangia atropurpurea (Mertens ex Roth) C. Agardh 1824

14 Bangia yamadae Tanaka 1944

15 Porphyra atropurpurea (olivi) De Toni 1897

16 Porphyra marcosii P. A. Cordero 1976

Dirique Bay, Burgos, Ilocos Norte, Philippines (Cordero 1976: 15)

17 Porphyra umbilicalis Kützing 1843 Ulva umbilicalis Linneaus

18 Pyropia denticulata (Levring) J. A. Philips & J. e. Sutherland 2011

Porphyra denticulata Levring

19Pyropia suborbiculata (Kjellman) J.e.Sutherland, H. G. Choi, M. S. Hwang & W. A. Nelson 2011

Porphyra suborbiculata Kjellman

20Pyropia vietnamensis (Tak. Tanaka & P.H. Ho) J. e. Sutherland & Monotilla 2011

Porphyra vietnamensis T. Tanaka & P. H. Ho

21 Wildemania variegata (Kjellman) De Toni 1890

Porphyra variegata (Kjellman) Kjellman

22 Bonnemaisoniaceae Asparagopsis taxiformis (Delile) Trevisan de Saint-Leon 1845

Falkenbergia hillebrandii (Bornet) Falkenberg

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23 Callithamniaceae Crouania attenuata (C. Agardh) J. Agardh 1842 Crouania minutissima Yamada

24 Caulacanthaceae Catenella caespitosa (Withering) L. M. Irvine 1976

25 Catenella impudica (Montagne) J. Agardh 1852

26 Catenella nipae Zanardini 1872

27 Caulacanthus ustulatus (Mertens ex Turner) Kützing 1843

28 Ceramiaceae Acrothamnion preissii (Sonder) e. M. Wollaston 1968

29 Antithamnion antillanum Børgesen 1917

Antithamnion lherminieri (P. L. Crouan & H. M. Crouan) Bornet ex Nasr

30 Balliella subcorticata (Itono) Itono & T. Tanaka 1973

31 Centroceras clavulatum (C. Agardh) Montagne 1846

32 Centroceras minutum Yamada 1944

33 Ceramium affine Setchell & N. L. Gardner 1930

34 Ceramium cimbricum H. e. Petersen 1924 Ceramium fastigiatum Harvey

35 Ceramium cruciatum F. S. Collins & Hervey 1917

36 Ceramium diaphanum (Lightfoot) Roth 1806

Ceramium gracillimum (Kützing) Griffiths & Harvey; Ceramium tenuissimum (Roth) Areschoug

37 Ceramium equisetoides e. Y. Dawson 1944

38 Ceramium loureiri C. Agardh 1824

39 Ceramium luetzelburgii o. C. Schmidt 1924

40 Ceramium marshallense e. Y. Daw-son 1957

41 Ceramium maryae Weber-van Bosse 1923

42 Ceramium mazatlanense e. Y. Daw-son 1950

43 Ceramium multijugum Jaasund 1970

44 Ceramium nitens C. Agardh (J. Agardh) 1851

45 Ceramium personatum Setchell & N. L. Gardner 1930

46 Ceramium procumbens Setchell & N. L. Gardner 1924

Ceramiella procumbens (Setchell & N. L. Gardner) Diaz-Piferrer

47 Ceramium serpens Setchell & N. L. Gardner 1924

48 Ceramium sinicola Setchell & N. L. Gardner 1924

49 Ceramium tenerrimum (G. Martens) okamura 1921

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50 Ceramium vagans P. C. Silva 1987

51 Ceramium zacae Setchell & N. L. Gardner 1937

52 Corallophila kleiwegii Weber-van Bosse 1923 Centroceras apiculatum Yamada

53 Gayliella flaccida (Harvey ex Kütz-ing) T. o. Cho & L. J. McIvor 2008

Ceramium flaccidum (Harvey ex Kützing) Ardissone

54 Gayliella taylorii (e. Y. Dawson) T. o. Cho & S. M. Boo 2008 Ceramium taylorii e. Y. Dawson

55 Herpochondria elegans (okamura) Itono 1977 Microcladia elegans okamura

56 Microcladia glandulosa (Solander ex Turner) Greville 1830

57 Champiaceae Champia bifida okamura 1901

58 Champia caespitosa e. Y. Dawson 1944

59 Champia compressa Harvey 1838

60 Champia disticha e. Y. Dawson 1944

61 Champia japonica okamura 1931

62 Champia parvula (C. Agardh) Harvey 1853

63 Champia salicornoides Harvey 1853

64 Champia spathulata Weber-van Bosse 1928

65 Champia vieillardii Kützing 1866

66 Coelothrix irregularis (Harvey) Børgesen 1920

67 Corallinaceae Amphiroa anastomosans Weber-van Bosse 1904

68 Amphiroa anceps (Lamarck) Decaisne 1842

69 Amphiroa annulata M. Lemoine 1929

70 Amphiroa beauvoisii J. V. Lamouroux 1816 Amphiroa zonata Yendo

71 Amphiroa crassa J. V. Lamouroux 1824

72 Amphiroa cumingii Montagne 1844

73 Amphiroa dimorpha M. Lemoine 1929

74 Amphiroa ephedraea (Lamarck) Decaisne 1842

75 Amphiroa foliacea J. V. Lamouroux 1824

76 Amphiroa foliacea f. erecta Weber- van Bosse 1904

77 Amphiroa fragilissima (Linnaeus) J. V. Lamouroux 1816

78Amphiroa fragilissima f. cyathifera (J. V. Lamouroux) Weber-van Bosse 1904

79 Amphiroa hancockii W. R. Taylor 1942

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80 Amphiroa pacifica Kützing 1858

81 Amphiroa rigida J. V. Lamouroux 1816

82 Amphiroa subcylindrica e. Y. Dawson 1953

83 Amphiroa tribulus (J. ellis & Solan-der) J. V. Lamouroux 1816

84 Amphiroa valonioides Yendo 1902

85 Arthrocardia palmata (J. ellis & Solander) Areschoug 1852

86 Bossiella frondescens (Postels & Ruprecht) e. Y. Dawson 1964

Corallina frondescens Postels & Ruprecht

87Hydrolithon farinosum (J. V. Lamour-oux) D. Penrose & Y. M. Chamberlain 1993

Fosliella farinosa (J. V. Lamouroux) M. A. Howe

88 Hydrolithon reinboldii (Weber-van Bosse & Foslie) Foslie 1909

89 Jania acutiloba (Decaisne) J. H. Kim, Guiry & H. G. Choi 2007

Cheilosporum jungermannioides Ruprecht ex Areschoug

90 Jania adhaerens J. V. Lamouroux 1816

91 Jania capillacea Harvey 1853

92 Jania cubensis Montagne ex Kützing 1849

Haliptilon cubense (Montagne ex Kützing) Garbary & H. W. Johansen

93 Jania cultrata (Harvey) J. H. Kim, Guiry & H. G. Choi 2007

Cheilosporum cultratum (Harvey) Areschoug

94 Jania decussato-dichotoma (Yendo) Yendo 1905

95 Jania longiarthra e. Y. Dawson 1953

96 Jania micrarthrodia J. V. Lamouroux 1816

97 Jania pacifica Areschoug 1852

98 Jania pumila J. V. Lamouroux 1816

99 Jania rubens (Linnaeus) J. V. Lamouroux 1816

100 Jania sagittata (J. V. Lamouroux) Blainville 1834

Cheilosporum sagittatum (J. V. Lamouroux) Areschoug

101 Jania spectabilis (Harvey ex Grunow) J. H. Kim, Guiry & H. G. Choi 2007

Cheilosporum spectabile Harvey ex Grunow

102 Jania tenella (Kützing) Grunow 1874

103 Jania tenella var. zacae e. Y. Dawson 1953

104 Jania ungulata (Yendo) Yendo 1905

105 Jania ungulata f. brevior (Yendo) Yendo 1905

106 Lithophyllum byssoides (Lamarck) Foslie 1900

107 Lithophyllum pallescens (Foslie) Foslie 1900

108 Lithophyllum pygmaeum (Heydrich) Heydrich 1897

Lithophyllum moluccense (Foslie) Foslie

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109 Lithoporella indica (Foslie) W. H. Adey 1970

110 Lithoporella melobesioides (Foslie) Foslie 1909

111 Mastophora rosea (C. Agardh) Setchell 1943

112 Mesophyllum simulans (Foslie) Me. Lemoine 1928

113 Neogoniolithon brassica-florida (Har-vey) Setchell & L. R. Mason 1943

Neogoniolithon frutescens (Foslie) Setchell & L. R. Mason

114Neogoniolithon megalocystum (Weber-van Bosse & Foslie) Setchell & L. R. Mason 1943

115 Porolithon onkodes (Heydrich) Foslie 1909

116 Corynocystaceae Corynocystis prostrata Kraft 1999 Bulusan, Sorsogon, Philippines (Kraft et al. 1999: 28).

117 Cubiculosporaceae Cubiculosporum koronicarpis Kraft 1973

Bulusan, Sorsogon, Philippines. (Kraft 1973: 881).

118 Cystocloniaceae Hypnea boergesenii T. Tanaka 1941

119 Hypnea caespitosa Geraldino & S. M. Boo 2010

Intertidal zone off Panglao Is-land, Bohol, Philippines (Ger-aldino et al. 2010: 338)

120 Hypnea cenomyce J. Agardh 1851

121 Hypnea charoides J. V. Lamouroux 1813

122 Hypnea cornuta (Kützing) J. Agardh 1851

123 Hypnea divaricata (C. Agardh) Greville 1830

124 Hypnea musciformis (Wulfen) J. V. Lamouroux 1813

125 Hypnea musciformis var. esperi J. Agardh 1851

126 Hypnea musciformis var. hippuroides (Kützing) Weber-van Bosse 1928

127 Hypnea nidulans Setchell 1924

128 Hypnea pannosa J. Agardh 1847

129 Hypnea saidana Holmes 1896

130 Hypnea spinella (C. Agardh) Kützing 1847 Hypnea cervicornis J. Agardh

131 Hypnea stellulifera (J. Agardh) Yama-gishi & M. Masuda 2003

Hypnea cornuta var. stellulifera J. Agardh

Lectotype locality: Manila, Phi-lippines (Yamagishi et al. 2003: 535).

132 Hypnea valentiae (Turner) Montagne 1841

133 Rhodophyllis peltata var. lacunosa Grunow 1874

134 Dasyaceae Dasya anastomosans (Weber-van Bosse) M. J. Wynne 2002

Dasya adhaerens Yamada; Eupogodon pilosus (Weber-van Bosse) P. C. Silva

135 Dasya antillarum (M. A. Howe) A.J.K .Millar 1996

Eupogodon antillarum (M. A. Howe) P. C. Silva

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136 Dasya baillouviana (S. G. Gmelin) Montagne 1841

137 Dasya caraibica Børgesen 1919

138 Dasya mollis Harvey 1853

139 Dasya ocellata (Grateloup) Harvey 1833

140 Dasya punicea (Zanardini) Meneghini ex Zanardini 1841

141 Dasya sessilis Yamada 1928

142 Dasya villosa Harvey 1844

143 Eupogodon planus (C. Agardh) Kütz-ing 1845

144 Heterosiphonia crispella (C. Agardh) M. J. Wynne 1985

Heterosiphonia crispella var. laxa (Børgesen) M. J. Wynne

145 Heterosiphonia muelleri (Sonder) De Toni 1903

146 Tapeinodasya bornetii Weber-van Bosse 1904

147 Delesseriaceae Apoglossum spathulatum (Sonder) Womersley & Shepley 1982

Hypoglossum spathulatum (Sonder) Kützing

148 Botryocarpa prolifera Greville 1830

149 Caloglossa bengalensis (G. Martens) R. J. King & Puttock 1994

Caloglossa adnata (Zanardini) De Toni

150 Caloglossa leprieurii (Montagne) G. Martens 1869

151 Caloglossa ogasawaraensis okamura 1897

152 Caloglossa vieillardii (Kützing) Setchell 1924

Caloglossa leprieurii var. hookeri Post

153 Claudea batanensis Tanaka 1967Basco, Batan Island, Batanes, Luzon, Philippines (Tanaka 1967: 18).

154 Claudea multifida Harvey 1854

155 Hypoglossum attenuatum N. L. Gardner 1927

156 Hypoglossum harveyanum (J. Agardh) Womersley & Shepley 1982 Hypoglossum serrulatum J. Agardh

157 Hypoglossum heterocystideum (J. Agardh) J. Agardh 1898

158 Martensia australis Harvey 1855 Martensia speciosa Zanardini

159 Martensia flabelliformis Harvey ex J. Agardh 1863

160 Martensia fragilis Harvey 1854

161 Martensia martensii (F. Schmitz) S. M. Lin, Fredericq & L. M. Liao 2001 Opephyllum martensii F. Schmitz

an der küste der Philippinen’ [Philippines] (Schmitz and Hauptfleisch 1897: 410).

162 Taenioma perpusillum (J. Agardh) J. Agardh 1863

163 Vanvoorstia spectabilis Harvey 1854

164 Zellera tawallina G. Martens 1866

165 Dicranemataceae Dicranema revolutum (C. Agardh) J. Agardh 1852

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166 Dumontiaceae Gibsmithia hawaiiensis Doty 1963

167 Rhodopeltis borealis Yamada 1931

168 Stenopeltis gracilis (Yamada & Tanaka) Itono & Yoshizaki 1992

Rhodopeltis gracilis Yamada & Tanaka

169 Endocladiaceae Gloiopeltis complanata (Harvey) Yamada 1932

170 Gloiopeltis tenax (Turner) Decaisne 1842

171 Erythropeltidaceae Erythrocladia irregularis Rosenvinge 1909

172 Erythrocladia pinnata W. R. Taylor 1942

173 Erythrotrichiaceae Erythrotrichia bangioides Levring 1955

174 Erythrotrichia biseriata Tanaka 1944

175 Erythrotrichia parietalis Tanaka 1952

176 Faucheaceae Gloiocladia ramellifera Hauck 1886

177

Leptofauchea leptophylla (Segawa) Mas. Suzuki, Nozaki, R. Terada, Kita-yama, Tetsu. Hashimoto & Yoshizaki 2012

Fauchea leptophylla Segawa

178 Galaxauraceae Actinotrichia fragilis (Forsskål) Børgesen 1932

179 Dichotomaria apiculata (Kjellman) A. Kurihara & Masuda 2005 Galaxaura apiculata Kjellman

180 Dichotomaria falcata (Kjellman) A. Kurihara & Masuda 2005 Galaxaura falcata Kjellman

181 Dichotomaria marginata (J. ellis & Solander) Lamarck 1816

Galaxaura marginata (J. ellis & Solander) J. V. Lamouroux

182 Dichotomaria obtusata (J. ellis & Solander) Lamarck 1816

Galaxaura obtusata (J. ellis & Solander) J. V. Lamouroux

183 Dichotomaria spathulata (Kjellman) A.Kurihara & Huisman 2006 Galaxaura arborea Kjellman

184 Galaxaura contigua Kjellman 1900

185 Galaxaura divaricata (Linnaeus) Huisman & R.A.Townsend 1993 Galaxaura fasciculata Kjellman

186 Galaxaura filamentosa R. C. Y. Chou 1945

187 Galaxaura kjellmanii Weber-van Bosse 1921

188 Galaxaura rugosa (J. ellis & Solan-der) J. V. Lamouroux 1816

Galaxaura subfruticulosa R. C. Y. Chou; Galaxaura subverticillata Kjellman

189 Galaxaura striata Kjellman 1900

190Tricleocarpa cylindrica (J. ellis & Solander) Huisman & Borowitzka 1990

Galaxaura fastigiata DecaisneLectotype (Galaxaura fastigata Decaisne): Manila, Luzon, Phil-ippines (Huisman 2006: 25).

191 Tricleocarpa fragilis (Linnaeus) Huis-man & R. A. Townsend 1993

Galaxaura oblongata (J. ellis & Solander) J. V. Lamouroux

192 Gelidiaceae Gelidium amansii (J. V. Lamouroux) J. V. Lamouroux 1813

193 Gelidium capense (S. G. Gmelin) P. C. Silva 1987

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194 Gelidium coulteri Harvey 1853

195 Gelidium crinale (Hare ex Turner) Gaillon 1828

196 Gelidium crinale var. perpusillum Piccone & Grunow 1884

197 Gelidium divaricatum G. Martens 1866

198 Gelidium isabelae W. R. Taylor 1945

199 Gelidium kintaroi Yamada 1941

200 Gelidium pulchellum (Turner) Kütz-ing 1868

201 Gelidium pusillum (Stackhouse) Le Jolis 1863

202 Gelidium pusillum var. pacificumW. R. Taylor 1945

203 Gelidium rigens (C. Agardh) Greville ex Kützing 1849

204 Ptilophora scalaramosa (Kraft) R. e. Norris 1987 Beckerella scalaramosa Kraft Bulusan, Sorsogon, Luzon, Phil-

ippines (Kraft 1976: 85).

205 Gelidiellaceae Gelidiella acerosa (Forsskål) Feld-mann & G. Hamel 1934

206 Parviphycus adnatus (e. Y. Dawson) B. Santelices 2004 Gelidiella adnata e. Y. Dawson

207 Gigartinaceae Chondracanthus tenellus (Harvey) Hommersand 1993 Gigartina tenella Harvey

208 Gracilariaceae Gracilaria arcuata Zanardini 1858

209 Gracilaria arcuata var. snackeyi Weber-van Bosse 1928

210 Gracilaria blodgettii Harvey 1853

211 Gracilaria bursa-pastoris (S. G. Gmelin) P. C. Silva 1952

Gracilaria compressa (C.Agardh) Greville

212 Gracilaria canaliculata Sonder 1871 Gracilaria crassa Harvey ex J.Agardh

213Gracilaria changii (B. M. Xia & I. A. Abbott) I. A. Abbott, J. Zhang & B. M. Xia 1991

214 Gracilaria coronopifolia J. Agardh 1852

215 Gracilaria cylindrica Børgesen 1920

216 Gracilaria damaecornis J. Agardh 1852

217 Gracilaria debilis (Forsskål) Børge-sen 1932

Polycavernosa debilis (Forsskål) Fredericq & J. N. Norris

218 Gracilaria disticha (J. Agardh) J. Agardh 1852

219 Gracilaria firma C. F. Chang & B. M. Xia 1976

220 Gracilaria foliifera (Forsskål) Børge-sen 1932

221 Gracilaria foliifera f. aeruginosa Børgesen 1938

222 Gracilaria gigas Harvey 1860

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223 Gracilaria incurvata okamura 1931

224 Gracilaria isabellana Gurgel, Fred-ericq & J. N. Norris 2004

Gracilaria lacinulata (M. Vahl) M. A. Howe

225 Gracilaria manilaensis Yamamoto & G. C. Trono 1994

Hali Beach, Parañaque, Manila Bay, Philippines (Yamamoto and Trono 1994: 95).

226 Gracilaria minor (Sonder) Durairat-nam 1961

227 Gracilaria papenfussii I. A. Abbott 1983

228 Gracilaria punctata (okamura) Yamada 1941

229 Gracilaria salicornia (C. Agardh) e. Y. Dawson 1954

Manila, Luzon, Philippines (Dawson 1954: 4; Millar and Xia 1997; see also Ganzon-Fortes 2012).

230 Gracilaria spinigera e. Y. Dawson 1949

231 Gracilaria spinulosa (okamura) C. F. Chang & B. M. Xia 1976

232Gracilaria srilankia (C. F. Chang & B. M. Xia) Withell, A. J. K. Millar & Kraft 1994

233 Gracilaria sullivanii Yamamoto & G. C. Trono 1994

Matabungkay, Batangas, Luzon Island, Philippines (Yamamoto and Trono 1994: 99).

234 Gracilaria tenuistipitata C. F. Chang & B. M. Xia 1976

235 Gracilaria tenuistipitata var. liui J. Zhang & B. M. Xia 1988

236 Gracilaria textorii (Suringar) De Toni 1895

237 Gracilaria turgida e. Y. Dawson 1949

238 Gracilaria vanbosseae (I. A. Abbott) I. A. Abbott 1991

Polycavernosa vanbosseae I. A. Abbott

239 Gracilaria venezuelensis W. R. Taylor 1942

240 Gracilaria vieillardii P. C. Silva 1987

241 Gracilariopsis bailiniae J. Zhang & B.M.Xia 1991

Gracilaria heteroclada J. Zhang & B. M. Xia 1988 nom. Illeg.

242 Gracilariopsis heteroclada J. Zhang & B. M. Xia 1991

243Gracilariopsis lemaneiformis (Bory de Saint-Vincent) e. Y. Dawson, Acleto & Foldvik 1964

Gracilaria lemaneiformis (Bory de Saint-Vincent) Greville

244Gracilariopsis longissima (S. G. Gmelin) M. Steentoft, L. M. Irvine & W. F. Farnham 1995

Gracilaria verrucosa (Hudson) Papenfuss

245 Hydropuntia edulis (S. G. Gmelin) Gurgel & Fredericq 2004

Gracilaria edulis (S. G. Gmelin) P. C. Silva; Gracilaria lichenoides Greville; Polycavernosa fastigiata C.F.Chang & B.M.Xia

246 Hydropuntia eucheumatoides (Har-vey) Gurgel & Fredericq 2004

Gracilaria eucheumatoides Harvey; Gracilaria eucheumoides Harvey

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247 Hydropuntia urvillei Montagne 1842

248 Halymeniaceae Carpopeltis capitellata (Sonder) F. Schmitz 1897

249 Carpopeltis maillardii (Montagne & Millardet) Chiang 1970

250 Cryptonemia crenulata (J. Agardh) J. Agardh 1851

251 Cryptonemia denticulata J. Agardh 1851

252 Cryptonemia seminervis (C. Agardh) J. Agardh 1846

Cryptonemia luxurians (C. Agardh) J. Agardh

253 Grateloupia acuminata Holmes 1896 Halymenia acuminata (Holmes) okamura

254 Grateloupia angusta (okamura) S. Kawaguchi & H. W. Wang 2001

Carpopeltis angusta (Harvey) okamura;Prionitis angusta (okamura) okamura

255 Grateloupia articulata (okamura) S. Kawaguchi & H. W. Wang 2001

Carpopeltis articulata (okamura) okamura

256 Grateloupia chiangii S. Kawaguchi & H. W. Wang 2001 Carpopeltis divaricata okamura

257 Grateloupia conferta (Kützing) Kütz-ing 1867

258 Grateloupia cornea okamura 1913 Prionitis cornea (okamura) e. Y. Dawson

259 Grateloupia crispata (okamura) Y. P. Lee 2008 Carpopeltis crispata okamura

260 Grateloupia dichotoma J. Agardh 1842

261 Grateloupia divaricata okamura 1895

262 Grateloupia doryphora (Montagne) M. A. Howe 1914

263 Grateloupia filicina (J. V. Lamour-oux) C. Agardh 1822

264 Grateloupia ramosissima okamura 1913

265 Grateloupia schmitziana (okamura) S. Kawaguchi & H. W. Wang 2001

Cryptonemia schmitziana (oka-mura) okamura

266 Halymenia dilatata Zanardini 1851

267 Halymenia durvillei Bory de Saint-Vincent 1828

268 Halymenia elongata C. Agardh 1822

269 Halymenia floresii (Clemente) C. Agardh 1817

270 Halymenia formosa Harvey ex Kütz-ing 1866

271 Halymenia harveyana J. Agardh 1892

272 Halymenia maculata J. Agardh 1885

273 Halymenia microcarpa (Montagne) P. C. Silva 1987

274 Halymenia stipitata I. A. Abbott 1998

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275 Polyopes affinis (Harvey) S. Kawagu-chi & H. W. Wang 2002

Carpopeltis affinis (Harvey) oka-mura

276 Polyopes prolifer (Hariot) S. Kawa-guchi & H. W. Wang 2002

Carpopeltis prolifera (Hariot) S. Kawaguchi & Masuda

277

Spongophloea procumbens (Weber-van Bosse) Huisman, De Clerck, Prud'homme van Reine & Borowitzka 2011

Thamnoclonium procumbens Weber-van Bosse

278

Spongophloea treubii (Weber-van Bosse) Huisman, De Clerck, Prud'homme van Reine & Borowitzka 2011

Thamnoclonium treubii Weber-van Bosse

279 Yonagunia formosana (okamura) S. Kawaguchi & Masuda 2004

Carpopeltis formosana okamura; Prionitis formosana (okamura) S. Kawaguchi & Nguyen

280 Hapalidiaceae Lithothamnion australe f. brachiatum Foslie 1904

281 Lithothamnion australe f. minutulum Foslie 1904

282 Lithothamnion australe f. ubianum Foslie 1904

283 Lithothamnion indicum f. subtile Foslie 1907

284 Mesophyllum erubescens (Foslie) Me. Lemoine 1928

285 Mesophyllum imbricatum (Dickie) W. H. Adey 1970

286 Mesophyllum pulchrum (Weber-van Bosse & Foslie) Lemoine 1928

287 Mesophyllum siamense (Foslie) W. H. Adey 1970

288 Phymatolithon calcareum (Pallas) W. H. Adey & D. L. McKibbin 1970

289Phymatolithon purpureum (P. L. Crouan & H. M. Crouan) Woelkerling & L. M. Irvine 1986

290 HymenocladiaceaeAsteromenia exanimans G. W. Saun-ders, C. e. Lane, C. W. Schneider & Kraft 2006

291 Hymenocladia dactyloides (Sonder) J. Agardh 1870 Gracilaria dactyloides Sonder 1845

292 Kallymeniaceae Callophyllis adhaerens Yamada 1932

293 Callophyllis adnata okamura 1932

294 Callophyllis fastigiata (J. Agardh) J. Agardh 1876 Gracilaria fastigiata J. Agardh

295 Kallymenia callophylloides okamura & Segawa 1935

296 Kallymenia pacifica Kylin 1956

297 Kallymenia perforata J. Agardh 1872

298 Kallymenia rosea Womersley & R. e. Norris 1971

299 Kallymenia rubra Womersley & R. e. Norris 1971

300 Kallymenia sessilis okamura 1934

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301 Kallymenia tasmanica Harvey 1860

302 Pugetia japonica Kylin 1941 Callophyllis okamurae P. C. Silva

303 Liagoraceae Dermonema virens (J. Agardh) Pedro-che & Ávila ortíz 1996

Dermonema frappieri (Montagne & Millardet) Børgesen

304 Ganonema farinosum (J. V. Lamour-oux) K. C. Fan & Yung C. Wang 1974 Liagora farinosa J. V. Lamouroux

305 Helminthocladia australis Harvey 1863

306 Izziella orientalis (J. Agardh) Huis-man & Schils 2002 Liagora orientalis J. Agardh

307 Liagora boergesenii Yamada 1938

308 Liagora canariensis Børgesen 1927

309 Liagora ceranoides J. V. Lamouroux 1816

Liagora ceranoides f. leprosa (J. Agardh) Yamada

310 Liagora fragilis Zanardini 1851

311 Liagora hawaiiana Butters 1911

312 Liagora japonica Yamada 1938

313 Liagora robusta Yamada 1938

314 Liagora segawae Yamada 1938

315Liagoropsis schrammii (P. L. Crouan & H. M. Crouan) Doty & I. A. Abbott 1964

316 Neoizziella divaricata (C. K. Tseng) S. M. Lin, S. Y.Yang & Huisman 2011 Liagora divaricata C. K. Tseng

317 Titanophycus setchellii (Yamada) S. M. Lin, S. Y. Yang & Huisman 2011 Liagora setchellii Yamada

318Titanophycus validus (Harvey) Huisman, G. W. Saunders & A. R. Sherwood 2006

Liagora tenuis J. Agardh; Liagora valida Harvey

319 Trichogloea requienii (Montagne) Kützing 1847

320 Yamadaella caenomyce (Decaisne) I. A. Abbott 1970 Liagora caenomyce Decaisne Manila, Luzon, Philippines (De-

caisne 1842: 119).

321 Lomentariaceae Ceratodictyon intricatum (C. Agardh) R. e. Norris 1987

Gelidiopsis intricata (C. Agardh) Vickers

322 Ceratodictyon repens (Kützing) R. e. Norris 1987

Gelidiopsis repens (Kützing) Weber-van Bosse

323 Ceratodictyon spongiosum Zanardini 1878

324 Ceratodictyon variabile (J.Agardh) R. e. Norris 1987

Gelidiopsis variabilis (Greville ex J. Agardh) F. Schmitz

325 Lomentaria articulata (Hudson) Lyngbye 1819

326 Lomentaria baileyana (Harvey) Farlow 1876

327 Lomentaria hakodatensis Yendo 1920

328 Lomentaria pinnata Segawa 1938

329 Palmariaceae Palmaria palmata (Linnaeus) Weber & Mohr 1805

330 Peyssonneliaceae Peyssonnelia conchicola Piccone & Grunow 1884

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331 Peyssonnelia distenta (Harvey) Yamada 1930

332 Peyssonnelia evae Weber-van Bosse 1921

Pearl Bank, Tawitawi, Sulu Archipelago, Philippines (Weber-van Bosse 1921: 279)

333 Peyssonnelia foveolata (Weber-van Bosse) Denizot 1968

334 Peyssonnelia indica (Weber-van Bosse) Denizot 1968

Syntype localities: Philippine Islands; various in Indonesia, including Semau and Solor (Silva et al. 1996: 212).

335 Peyssonnelia involvens Zanardini 1858

336 Peyssonnelia luzonensis P. A. Cordero 1977

Quezon Is., Hundred Is., Pan-gasinan, Philippines (Cordero 1977: 80).

337 Peyssonnelia mariti (Weber-van Bosse) Denizot 1968

338 Peyssonnelia obscura Weber-van Bosse 1921

Syntype localities: Philippine Islands; various in Indonesia, including Semau and Solor (We-ber-van Bosse 1921: 274-275).

339 Peyssonnelia orientalis (Weber-van Bosse) Cormaci & G. Furnari 1987

Peyssonnelia rubra f. orientalis Weber-van Bosse

Syntype localities: Various lo-calities in Indonesia and Philip-pine Islands (Weber-van Bosse 1921: 272).

340 Peyssonnelia rubra (Greville) J. Agardh 1851

341 Peyssonnelia squamaria (S. G. Gme-lin) Decaisne 1842

342 Polystrata dura Heydrich 1905

343 Ramicrusta calcea (Heydrich) K. R. Dixon 2013 Peyssonnelia calcea Heydrich

344 Phyllophoraceae Ahnfeltiopsis divaricata (Holmes) Masuda 1993 Gymnogongrus divaricatus Holmes

345 Ahnfeltiopsis flabelliformis (Harvey) Masuda 1993

Gymnogongrus flabelliformis Harvey

346 Ahnfeltiopsis okamurae P. C. Silva & DeCew 1992 Ahnfeltia furcellata okamura

347 Ahnfeltiopsis pygmaea (J. Agardh) P. C. Silva & DeCew 1992 Gymnogongrus pygmaeus J. Agardh

348 Gymnogongrus dilatatus (Turner) J. Agardh 1851

349 Phyllophora submaritima e. Y. Dawson 1949

350 Plocamiaceae Plocamium cirrhosum (Turner) M. J. Wynne 2002

Plocamium costatum (C. Agardh) J. D. Hooker & Harvey

351 Plocamium patens G. Martens 1868

Syntype localities: Timor, Indo-nesia; Zamboanga, Mindanao, Philippines (Silva et al. 1996: 343).

352 Plocamium telfairiae (W. J. Hooker & Harvey) Harvey ex Kützing 1849

353 Plocamium serrulatum var. pectina-tum P. A. Cordero 1977

Tajojora, Batan Is. Batanes, Philippines (Cordero 1977: 140).

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354 Pterocladiaceae Pterocladiella caloglossoides (M. A. Howe) Santelices 1998

Pterocladia caloglossoides (M. A. Howe) e. Y. Dawson

355 Pterocladiella capillacea (S. G. Gme-lin) Santelices & Hommersand 1997

Pterocladia capillacea (S. G. Gme-lin) Bornet; Pterocladia densa okamura

356 Pterocladiella nana (okamura) Shi-mada, Horiguchi & Masuda 2000 Pterocladia nana okamura

357 Pterocladiella taylorii (Joly) Santeli-ces 2007 Gelidiella taylori Joly

358 Rhizophyllidaceae Portieria hornemannii (Lyngbye) P. C. Silva 1987

Chondrococcus hornemannii (Lyng-bye) F. Schmitz Desmia hornemannii Lyngbye

359 Portieria japonica (Harvey) P. C. Silva 1987

360 Rhodogorgonaceae Renouxia antillana Fredericq & J. N. Norris 1995

361 Rhodomelaceae Acanthophora aokii okamura 1934

362 Acanthophora muscoides (Linnaeus) Bory de Saint-Vincent 1828

363 Acanthophora spicifera (M. Vahl) Børgesen 1910

Acanthophora orientalis J. Agardh; Acanthophora thierryi J. V. Lamour-oux

364 Acrocystis nana Zanardini 1872

365 Alsidium pusillum e. Y. Dawson 1963

366 Amansia pumila (Sonder) J.Agardh 1892

Melanamansia pumila (Sonder) R. Norris

367 Aneurianna lorentzii (Weber-van Bosse) L. e. Phillips 2006

Lenormandiopsis lorentzii (Weber-van Bosse) Papenfuss

368 Bostrychia calliptera (Montagne) Montagne 1842

369 Bostrychia intricata (Bory de Saint-Vincent) Montagne 1852

370 Bostrychia kelanensis Grunow 1936

371 Bostrychia moritziana (Sonder ex Kützing) J. Agardh 1863

372 Bostrychia radicans (Montagne) Montagne 1842

373 Bostrychia simpliciuscula Harvey ex J. Agardh 1863

374 Bostrychia tenella (J. V. Lamouroux) J. Agardh 1863 Bostrychia binderi Harvey

375 Chondria armata (Kützing) okamura 1907

376 Chondria crassicaulis Harvey 1860

377 Chondria curvilineata F. S. Collins & Hervey 1917

378 Chondria dasyphylla (Woodward) C. Agardh 1817

379 Chondria polyrhiza F. S. Collins & Hervey 1917

380 Chondria repens Børgesen 1924

381 Chondria sedifolia Harvey 1853

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382 Chondria seticulosa (Forsskål) C. Agardh 1822

383 Chondria sibogae Weber-van Bosse 1923

Syntype localities: Borneo Bank and Lombok, Indonesia; Sulu Archipelago, Philippines (Silva et al. 1996: 485).

384 Chondrophycus carolinensis (Y. Saito) K. W. Nam 2007 Laurencia carolinensis Y. Saito

385 Chondrophycus cartilagineus (Ya-mada) Garbary & J. T. Harper 1998 Laurencia cartilaginea Yamada

386Chondrophycus ceylanicus (J. Agardh) M. J. Wynne, Serio, Cormaci & G. Furnari 2005

Laurencia ceylanica J. Agardh

387Chondrophycus columellaris (Børge-sen) e. Coppejans & A. J. K. Millar 2000

Laurencia columellaris Børgesen

388 Chondrophycus glandulifer (Kützing) Lipkin & P. C. Silva 2002

Laurencia glandulifera (Kützing) Kützing

389 Chondrophycus intermedius (Ya-mada) Garbary & J. T. Harper 1994 Laurencia intermedia Yamada

390Chondrophycus parvipapillatus (C.K.Tseng) Garbary & J. T. Harper 1994

Laurencia parvipapillata C.K. Tseng

391 Chondrophycus tronoi (Ganzon-Fortes) K. W. Nam 1999 Laurencia tronoi Ganzon-Fortes

North of Pulong Bakaw, Ca-latagan, Batangas, Philippines (Ganzon-Fortes 1982: 404).

392 Chondrophycus undulatus (Yamada) Garbary & J. T. Harper 1998 Laurencia undulata Yamada

393 Digenea simplex (Wulfen) C. Agardh 1822

394 Enantiocladia okamurae Yamada 1930

395 Endosiphonia spinulosa (Harvey) Womersley & Parsons 2003 Endosiphonia spinuligera Zanardini

396 Erythrocolon podagricum J. Agardh 1874

397 Exophyllum wentii Weber-van Bosse 1911

Syntype localities: Stat. 58. Ile de Savu récif; Stat. 80. Banc de Bornéo 40-50 m; Stat. 99. Ile Ubian du nord, Archipel de Sulu 12-33 m [North Ubian Island, Sulu Archipelago, Philippine Islands; Indonesia: Borneo Bank and Sawu] (Weber-van Bosse 1911: 29).

398 Herposiphonia delicatula Hollenberg 1968

399 Herposiphonia dendroidea Hollen-berg 1968

400 Herposiphonia dendroidea var. minor Hollenberg 1968

401 Herposiphonia nuda Hollenberg 1968

402 Herposiphonia obscura Hollenberg 1968

403 Herposiphonia pacifica Hollenberg 1968

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404 Herposiphonia parca Setchell 1926

405 Herposiphonia plumula (J. Agardh) Falkenberg 1901

406 Herposiphonia secunda (C. Agardh) Ambronn 1880

407 Herposiphonia secunda f. tenella (C. Agardh) M. J. Wynne 1985

408 Herposiphonia subdisticha okamura 1899

409 Herposiphonia tenella (C. Agardh) Ambronn 1880

410 Herposiphonia trichia Hollenberg 1968

411 Laurencia brongniartii J. Agardh 1841

412 Laurencia caraibica P. C. Silva 1972

413 Laurencia chondrioides Børgesen 1918

414 Laurencia clavata Sonder 1853

415 Laurencia composita Yamada 1931

416 Laurencia decumbens Kützing 1863

417 Laurencia dendroidea J. Agardh 1852 Laurencia obtusa var. dendroidea (J. Agardh) Yamada

418 Laurencia distichophylla J. Agardh 1852

419 Laurencia flexilis Setchell 1926

420 Laurencia forsteri (Mertens ex Turner) Greville 1830

421 Laurencia intricata J. V. Lamouroux 1813 Laurencia implicata J. Agardh

422 Laurencia majuscula (Harvey) A. H. S. Lucas 1935

423 Laurencia mariannensis Yamada 1931

424 Laurencia nidifica J. Agardh 1852

425 Laurencia nipponica Yamada 1931

426 Laurencia obtusa (Hudson) J. V. Lamouroux 1813

427 Laurencia obtusa var. densa Yamada 1931

428 Laurencia okamurae Yamada 1931 Laurencia japonica Yamada

429 Laurencia pinnata Yamada 1931

430 Laurencia similis K. W. Nam & Saito 1991

431 Laurencia snackeyi (Weber-van Bosse) M. Masuda 1997

Laurencia obtusa var. snackeyi (Weber-van Bosse) Yamada

432 Laurencia subsimplex C. K. Tseng 1943

433 Laurencia tropica Yamada 1931

434 Laurencia venusta Yamada 1931

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435 Leveillea jungermannioides (Hering & G. Martens) Harvey 1855

436 Lophocladia lallemandi (Montagne) F. Schmitz 1893

437 Lophosiphonia cristata Falkenberg 1901

438 Melanamansia glomerata (C. Agardh) R. e. Norris 1995 Amansia glomerata C. Agardh

439 Murrayella periclados (C. Agardh) F. Schmitz 1893

440 Neosiphonia apiculata (Hollenberg) Masuda & Kogame 2003 Polysiphonia apiculata Hollenberg

441 Neosiphonia beaudettei (Hollenberg) M. S. Kim & I. A. Abbott 2006 Polysiphonia beaudettei Hollenberg

442Neosiphonia ferulacea (Suhr ex J. Agardh) S. M. Guimarães & M. T. Fujii 2004

Polysiphonia ferulacea Suhr ex J. Agardh

443 Neosiphonia gorgoniae (Harvey) S. M. Guimarães & M. T. Fujii 2004 Polysiphonia gorgoniae Harvey

444 Neosiphonia hawaiiensis (Hollen-berg) M. S. Kim & I. A. Abbott 2006

Polysiphonia hawaiiensis Hol-lenberg

445 Neosiphonia poko (Hollenberg) I. A. Abbott 2002 Polysiphonia poko Hollenberg

446 Neosiphonia sertularioides (Grate-loup) K. W. Nam & P. J. Kang 2012

Neosiphonia flaccidissima (Hollen-berg) M. S. Kim & I. K. Lee

447 Neosiphonia sphaerocarpa (Børge-sen) M. S. Kim & I. K. Lee 1999

Polysiphonia sphaerocarpa Børge-sen

448 Neosiphonia tepida (Hollenberg) S. M. Guimarães & M. T. Fujii 2004 Polysiphonia tepida Hollenberg

449 Neosiphonia tongatensis (Harvey ex Kützing) M. S. Kim & I. K. Lee 1999

Polysiphonia mollis var. tongatensis (Harvey & Kützing) Hollenberg

450 Neosiphonia upolensis (Grunow) M. S. Kim & Boo 2008 Polysiphonia upolensis Grunow

451 Neurymenia fraxinifolia (Mertens ex Turner) J. Agardh 1863

452 Osmundaria obtusiloba (C. Agardh) R. e. Norris 1991

Vidalia obtusiloba (Mertens ex C. Agardh) J. Agardh

453 Osmundea pinnatifida (Hudson) Stackhouse 1809

Laurencia pinnatifida (Hudson) J. V. Lamouroux

454 Palisada capituliformis (Yamada) K. W. Nam 2007 Laurencia capituliformis Yamada

455 Palisada corallopsis (Montagne) Sentíez, M. T. Fujii & Díaz 2008

Laurencia corallopsis (Montagne) M. A. Howe

456 Palisada flagellifera (J. Agardh) K. W. Nam 2007 Laurencia flagellifera J. Agardh

457 Palisada parvipapillata (C. K. Tseng) K. W. Nam 2007

Chondrophycus parvipapillatus (C. K. Tseng) Garbary & J. T. Harper

458Palisada patentiramea (Montagne) Cassano, Sentíes, Gil-Rodríguez & M. T. Fujii 2009

Chondrophycus patentiramea (Mon-tagne) Nam; Laurencia patentiramea (Montagne) Kützing

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459 Palisada perforata (Bory de Saint-Vincent) K. W. Nam 2007

Chondrophycus papillosus (C. Agardh) D. J. Garbary & J. T. Harper; Laurencia papillosa (C. Agardh) Greville, Setchell & N. L. Gardner

460 Palisada robusta K. W. Nam 2007 Laurencia palisada Yamada

461 Palisada surculigera (C. K. Tseng) K. W. Nam 2007 Laurencia surculigera C. K. Tseng

462Palisada thuyoides (Kützing) Cas-sano, Sentíes, Gil-Rodríguez & M. T. Fujii 2009

Laurencia paniculata (C. Agardh) J. Agardh

463 Palisada yamadana (M. A. Howe) K. W. Nam 2007 Laurencia yamadana M. A. Howe

464 Polysiphonia flabelluta Harvey 1860

465 Polysiphonia forfex Harvey 1859

466 Polysiphonia fragilis Suringar 1867

467 Polysiphonia howei Hollenberg 1945

468 Polysiphonia japonica var. savatieri (Hariot) Yoon 1986 Polysiphonia savatieri Hariot

469 Polysiphonia mollis J. D. Hooker & Harvey 1847

470 Polysiphonia pacifica Hollenberg 1942

471 Polysiphonia pacifica var. delicatula Hollenberg 1942

472 Polysiphonia scopulorum Harvey 1855

473 Polysiphonia scopulorum var. villum (J. Agardh) Hollenberg 1968

474 Polysiphonia sparsa (Setchell) Hol-lenberg 1968

475 Polysiphonia subtillisima Montagne 1840

476 Polysiphonia triton P. C. Silva 1987

477 Polysiphonia tsudana Hollenberg 1968

478 Tolypiocladia calodictyon (Harvey ex Kützing) P. C. Silva 1952

479 Tolypiocladia condensata (Weber-van Bosse) P. C. Silva 1952

480 Tolypiocladia glomerulata (C. Agardh) F. Schmitz 1897

481 Womersleyella setacea (Hollenberg) R. e. Norris 1992 Polysiphonia setacea Hollenberg

482 Yuzurua poiteaui (J. V. Lamouroux) Martin-Lescanne 2010

Laurencia poiteaui (J. V. Lamour-oux) M. A. Howe

483 Rhodymeniaceae Botryocladia botryoides (Wulfen) Feldmann 1941

484 Botryocladia leptopoda (J. Agardh) Kylin 1931

485 Botryocladia pyriformis (Børgesen) Kylin 1931

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486 Botryocladia skottsbergii (Børgesen) Levring 1941

487 Botryocladia uvarioides e. Y. Daw-son 1944

488 Chrysymenia kaernbachii Grunow 1898 Chrysemenia kaernbachii Grunow

489 Rhodymenia californica Kylin 1931

490 Rhodymenia coacta okamura & Segawa 1935

491 Rhodymenia decumbens W. R. Taylor 1945

492 Rhodymenia divaricata e. Y. Dawson 1941

493 Rhodymenia intricata (okamura) okamura 1930

494 Sarcodiaceae Sarcodia montagneana (J. D. Hooker & Harvey) J. Agardh 1852

495 Sarcomeniaceae Cottoniella filamentosa (M. A. Howe) Børgesen 1920

496 Schizymeniaceae Titanophora incrustans (J. Agardh) Børgesen 1949

497 Titanophora mauritiana Børgesen 1949

498 Titanophora weberae Børgesen 1943

499 Scinaiaceae Scinaia aborealis Huisman 1986

500 Scinaia hormoides Setchell 1914

501 Scinaia japonica Setchell 1914

502 Scinaia latifrons M. A. Howe 1911

503 Scinaia moniliformis J. Agardh 1885

504 Scinaia tsinglanensis C. K. Tseng 1941

505 SebdeniaceaeLesleigha yamadae (okamura & Segawa) G. W. Saunders & Kraft 2011

Sebdenia yamadae okamura & Segawa

506 Sebdenia flabellata (J. Agardh) P. G. Parkinson 1980

507 Sebdenia limensis (Kützing) M. A. Howe 1914

508 Solieriaceae Betaphycus gelatinus (esper) Doty ex P. C. Silva 1996

Eucheuma gelatinae (esper) J. Agardh; Eucheuma gelatinum (esper) Doty

509 Betaphycus philippinensis Doty 1995Betaphycus philippinense Doty; Betaphycus philippinensis var. philippinensis Doty

Holotype: eastern Sorsogon, Philippines (Doty 1995: 237).

510 Eucheuma alvarezii var. ajakii-assii Doty 1985

eucheuma farm, about 1 km southwest of the southern tip of Tumindao Island, Sulu Archipel-ago, Philippines (Doty 1985: 42).

511 Eucheuma amakusaense okamura 1902

512 Eucheuma arnoldii Weber-van Bosse 1928

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513 Eucheuma arnoldii var. alcyonida Kraft 1972

Bulusan, Sorsogon, Philippines (Kraft 1972: 323).

514 Eucheuma crassum Zanardini 1878

515 Eucheuma crustiforme Weber-van Bosse 1928

516 Eucheuma denticulatum (N. L. Bur-man) F. S. Collins & Hervey 1917

Eucheuma muricatum (S.G.Gmelin) Weber-van Bosse

517 Eucheuma denticulatum var. endong Trono & Ganzon-Fortes 2012

Dancalan, Sorsogon, Philippines (Ganzon-Fortes et al. 2012: 1110)

518 Eucheuma edule (Kützing) Weber-van Bosse 1926

519 Eucheuma horridum J. Agardh 1852

520 Eucheuma isiforme (C. Agardh) J. Agardh 1847

521 Eucheuma leeuwenii Weber-van Bosse 1928

522 Eucheuma perplexum Doty 1988

523 Eucheuma serra (J. Agardh) J. Agardh 1847

524 Kappaphycus alvarezii (Doty) Doty ex P. C. Silva 1996

Eucheuma alvarezii Doty; Eucheuma cottonii Weber-van Bosse; Eucheuma cottonii var. erecta ?

525 Kappaphycus alvarezii var. tamba-lang (Doty) nom. inval.

Eucheuma alvarezii var. tambalang Doty

Seaweed farm at Calatagan, Batangas, Philippines (Doty 1985: 41).

526 Kappaphycus cottonii (Weber-van Bosse) Doty ex P. C. Silva 1996 Eucheuma okamurae Yamada

527 Kappaphycus procrusteanus (Kraft) Doty 1988

Eucheuma procrusteanum Kraft; Kappaphycus procrusteanum Kraft

Semirara Islands of the northern Sulu Sea, Philippines (Kraft 1970: 215).

528 Kappaphycus striatus (F. Schmitz) Doty ex P. C. Silva 1996

Eucheuma striatum F. Schmitz; Kappaphycus striatum F. Schmitz

529 Meristotheca coacta okamura 1930

530 Meristotheca papulosa (Montagne) J. Agardh 1872

531 Sarconema filiforme (Sonder) Kylin 1932

Dicranema furcellatum (Montagne) J. Agardh

532 Solieria dura (Zanardini) F. Schmitz 1895

533 Solieria pacifica (Yamada) Yoshida 1989

534 Solieria robusta (Greville) Kylin 1932

535 Wurdemannia miniata (Sprengel) Feldmann & G. Hamel 1934

536 Sporolithaceae Sporolithon erythraeum (Rothpletz) Kylin 1956

537 Sporolithon schmidtii (Foslie) G. D. Gordon, Masaki & Akioka 1976

538 Sporolithon sibogae (Weber-van Bosse & Foslie) P. C. Silva 1987

Archaeolithothamnion sibogae Weber-van Bosse & Foslie

Pearl Bank, Sulu Archipelago, Philippines (Silva et al. 1996: 277)

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539 Sporolithon timorense (Foslie) P. C. Silva 1987

540 Spyridiaceae Spyridia filamentosa (Wulfen) Har-vey 1833

541 Spyridia velasquezii G. C. Trono 1972

542 Stylonemataceae Chroodactylon ornatum (C. Agardh) Basson 1979

543 Rhodosorus marinus Geitler 1930

544 Stylonema alsidii (Zanardini) K. M. Drew 1956

Goniotrichum alsidii (Zanardini) M. A. Howe

545 Wrangeliaceae Anotrichium tenue (C. Agardh) Nageli 1862

546 Dasyphila plumarioides Yendo 1920

547 Gordoniella yonakuniensis (Yamada & T. Tanaka) Itono 1977

548 Griffithsia ovalis Harvey 1855

549 Griffithsia rhizophora Grunow ex Weber-van Bosse 1923

550 Griffithsia subcylindrica okamura 1930

551 Gymnothamnion elegans (Schousboe ex C. Agardh) J. Agardh 1892

552 Haloplegma duperreyi Montagne 1842

553 Pleonosporium globuliferum Levring 1941

554Ptilothamnion cladophorae (Yamada & T. Tanaka) G. Feldmann-Mazoyer 1941

555 Wrangelia argus (Montagne) Mon-tagne 1856

556 Wrangelia bicuspidata Børgesen 1916

557 Wrangelia penicillata (C. Agardh) C. Agardh 1828

558 Wrangelia plumosa Harvey 1844

559 Wrangelia velutina (Sonder) Harvey 1855

560 Rhodophyta incertae sedis

Callophycus serratus (Harvey ex Kützing) P. C. Silva 1957

Taxon with Generic Record Only

List No. Family Taxon Recorded Remarks Reference

561 Callithamniaceae Callithamnion Lyngbye Kraft et al. 1999

562 Euptilota (Kützing) Kützing Kraft et al. 1999

563 Dumontiaceae Dudresnaya P. L. Crouan & H. M. Crouan Kraft et al. 1999

564 Rhabdoniaceae Rhabdonia J. D. Hooker & Harvey Kraft et al. 1999

Species with No Matching Record in AlgaeBase

Family Taxon Recorded Remarks Reference

Corallinaceae Amphiroa fragilissima f. fragilissima (Linaeus) (Lamoroux) Weber van Bosse

Also not listed in INA Database Trono 2004

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Jania tenella var. tenella e. Y. Dawson Also not listed in INA Database Trono 2004

Halymeniaceae Halymenia japonica? Domantay Also not listed in INA Database Silva et al. 1987, Ganzon-Fortes 2012

Kallymeniaceae Kallymenia denticulata No authority of taxon listed;Also not listed in INA Database

Ganzon-Fortes 2012

Species Listed as Preliminary Entry in AlgaeBase

Family Taxon Recorded Remarks Reference

Corallinaceae Jania constricta Kützing, nom. illeg. Also not listed in INA Database Trono 2004

Peyssonneliaceae Peyssonnelia rubra var. orientalis Weber-van Bosse

As Peyssonnelia rubra f. orientalis Weber-van Bosse in INA Database

Silva et al. 1987;Silva 1996-to date

luzoniensis Martens was also described based on Phil-ippine materials (Table 1). But this taxon is listed as a preliminary entry in AlgaeBase so its current taxonomic status remains unchecked.

Although the number of species (893) recorded from the Philippines compiled in the present work remains com-parable with that compiled by Ganzon-Fortez (2012) for records up to 1999 (888 species, excluding Cyanobacte-ria), the details of the species list are quite different. This is so mainly because of the large number of taxa that has been synonymized in recent years. on a purely nu-merical sense, there is a net increase of 21 species among the reds in the new list, compared to a drop of 1 and 15 species respectively among the browns and the greens. Payo et al. (2013) recently suggested naming 21 new cryptic species of the red alga Portieria from the Philip-pines based on three molecular markers they used. These “new” species have not been considered in the present compilation.

As in other attempts to compile a listing of algal records, no actual examination of the algal materials was done. This listing should therefore be viewed simply as a listed record with the validity of the published species names verified. It does not, however, verify the correct identifi-cation of each original record. Readers which find mis-takes in the list are most welcomed to inform the authors so an improved list can be made in the future. Further studies involving actual examination of the original algal materials are expected to verify the species identity.

ACKNoWLeDGeMeNTSThis paper is dedicated to the memory of Leonardo L. Co, a foremost Philippine plant taxonomist, conservationist and botanical activist whose fruitful life was cut short by the wanton gun fire in the forest of Leyte, Philippines on November 15, 2010. We thank Dr. Benito Tan of the

National University of Singapore for his comments and editorial suggestions on the manuscript. Special thanks are also given to Dr. Lawrence Liao of Hiroshima University, Japan, for his very valuable comments, corrections and additions on our earlier list of algal records from the Philippines. This work benefited from funding from the Research Grant Council of Hong Kong GRF Project No. CUHK460012.

ReFeReNCeSABBoTT IA. 1962. Some Liagora-inhabiting species

of Acrochaetium. occasional Papers Bernice Pauahi Bishop Museum 23: 77-120.

ABBoTT IA. 1985. Gracilaria from the Philippines: List and distribution of the species. Taxonomy of economic Seaweeds 1: 89-90.

ABBoTT IA. 1988. Some species of Gracilaria and Polycavernosa from the Siboga expedition. Taxonomy of economic Seaweeds 2: 151-156.

ABBoTT IA. 1994. New records and a reassessment of Gracilaria (Rhodophyta) from the Philippines. Taxonomy of economic Seaweeds 4: 111-118.

ABBoTT IA. 1999. Notes on some species of Halymenia in the southwestern Pacific. Taxonomy of economic Seaweeds 7: 163-172.

AGARDH JG. 1848. Species genera et ordines algarum, seu descriptiones succinctae specierum, generum et ordinum, quibus algarum regnum constituitur. Volumen Primum. Algas fucoideas complectens. Lundae [Lund]: C.W.K. Gleerup. pp. [i-vi], [i]-viii, [1]-363.

AJISAKA T, NoRo T, YoSHIDA T. 1995. Zygocarpic Sargassum species (Subgenus Sargassum) from Japan. Taxonomy of economic Seaweeds 5: 11-44.

ANG Po, TRoNo GC. 1987. The genus Sargassum

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