alternative mechanisms of carbon fixation (5.4). rubisco rubisco is the most abundant protein on...
TRANSCRIPT
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Alternative Mechanisms of Carbon Fixation
(5.4)
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RuBisCO
• RuBisCO is the most abundant protein on Earth
• catalyzes the first reaction of the Calvin cycle
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Photorespiration
• in the first reaction of the Calvin cycle, CO2 is “fixed”, forming 2 molecules of 3-PGA
• O2 competes with CO2 for the active site of RuBisCO (ribulose-1,5-bisphosphate carboxylase oxidase)
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Photorespiration
• RuBP is combined with O2
• produces only 1 molecule of 3-PGA
• also produces a molecule that is toxic to the cell that must be removed in a series of reactions, which use ATP release 1 CO2
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Photorespiration
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In hot & dry climates…
• stomata close during the day to reduce water loss
• so less CO2 is present, less is fixed into G3P, and more photorespiration occurs
• How have plants in warmer climates evolved to cope with RuBisCO’s affinity for O2?
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C4 Photosynthesis
• CO2 is combined with PEP to form 4-carbon oxaloacetate (enzyme PEP carboxylase) in mesophyll cells
• oxaloacetate is converted to malate, then transported into bundle-sheath cells
• malate is decarboxylated forming pyruvate, and CO2 enters the Calvin cycle
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C4 Photosynthesis
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C4 Photosynthesis
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C4 Plants
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CAM Plants
• crassulacean acid metabolism (CAM), first discovered in Crassulacea family of plants
• stomata are closed during the day, so CO2 is only absorbed at night
• C4 pathway at night, C3 pathway (Calvin cycle) during the day
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CAM Plants
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CAM plants
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Animation
Narrated animation of C4 photosynthesis
and CAM plants:
http://www.uic.edu/classes/bios/bios100/lectures/c4.htm