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Harvard University Program in Neuroscience History of Enrollment in The Program in Neuroscience November 2015 Updated each July

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Page 1: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Harvard University Program in Neuroscience

History of Enrollment in

The Program in Neuroscience

November 2015 Updated each July

Page 2: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Nicholas Spitzer, M.D./Ph.D.B.A., Harvard College

Entered 1966 * Defended May 14, 1969Advisor: David Po er

A Physiological and Histological Inves ga onof the Intercellular Transfer of Small Molecules

_____________Professor of Neurobiology

University of California at San Diego

Eric Frank, Ph.D.B.A., Reed College

Entered 1967 * Defended January 17, 1972Advisor: Edwin J. Furshpan

The Control of Facilita on at the Neuromuscular Junc on of the Lobster_______________

Professor Emeritus of PhysiologyTu s University School of Medicine

David Van Essen, Ph.D.B.S., California Ins tute of Technology

Entered 1967 * Defended October 22, 1971Advisor: John Nicholls

E ects of an Electronic Pump on Signaling by Leech Sensory Neurons______________

Professor of Anatomy and NeurobiologyWashington University

Albert Hudspeth, M.D./Ph.D.B.A., Harvard College

Entered 1967 * Defended April 30, 1973Advisor: David Po er

Intercellular Junc ons in Epithelia_______________

Professor of NeuroscienceRockefeller University

Page 3: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Richard Mains, Ph.D.Sc.B., M.S., Brown University

Entered 1968 * Defended April 24, 1973Advisor: David Po er

Tissue Culture of Dissociated Primary Rat Sympathe c Neurons:Studies of Growth, Neurotransmi er Metabolism, and Matura on

_______________Professor of Neuroscience

University of Connec cut Health Center

Peter MacLeish, Ph.D.B.E.Sc., University of Western Ontario

Entered 1969 * Defended December 29, 1976Advisor: David Po er

Synapse Forma on in Cultures of Dissociated Rat Sympathe c NeuronsGrown on Dissociated Rat Heart Cells

_______________Professor and Director of the Neuroscience Ins tute

Morehouse College of Medicine

Peter Sargent, Ph.D.B.A., Amherst College

Entered 1969* Defended March 21, 1975Advisor: Zachary Hall

Transmi ers in the Leech Central Nervous System: Analysis of Sensoryand Motor Cells_______________

Professor of Cell and Tissue BiologyUniversity of California at San Francisco School of Den stry

Bruce Wallace, Ph.D.B.A., Amherst College

Entered 1969 * Defended September 27, 1974Advisor: Edward Kravitz

The Biosynthesis of Octopamine in the Nervous System of the Lobster,Homarus americanus_______________Professor Emeritus

University of Colorado School of Medicine

Page 4: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Joshua Sanes, Ph.D.B.A., Yale University

Entered 1970 * Defended January 15, 1976Advisor: John Hildebrand

Origin and Di eren a on of Sensory Neurons in an Insect Antenna_______________

Professor of Molecular and Cellular BiologyHarvard University

Colin Nurse, Ph.D.B.Sc., University of Western Ontario

Entered 1971 * Defended May 11, 1977Advisor: David Po er

The forma on of cholinergic synapses between dissociated ratsympathe c neurons

and skeletal myotubes in cell culture_______________

Professor of BiologyMcMaster University

Carla Shatz, Ph.D.B.A., Radcli e College

M.Phil., University College, LondonEntered 1971 * Defended March 25, 1976

Advisor: David HubelA Study of Abnormal Connec ons in the Visual System of the Siamese

Cat, a Gene c Mutant_______________

Professor of NeurobiologyStanford University

King Wai Yau, M.D./Ph.D.A.B., Princeton University

Entered 1971 * Defended October 31, 1975Advisor: John Nicholls

Recep ve Fields, Geometry and Conduc on Block of Sensory Cells in theLeech Central Nervous System

_______________Professor of Neuroscience

Johns Hopkins University School of Medicine

Page 5: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Linda Lai Yue Chun, Ph.D.B.S., Illinois Ins tute of Technology

Entered 1972 * Defended May 9, 1977Advisor: Paul Pa erson

The Role of Nerve Growth Factor in the Development of RatSympathe c Neurons in vitro

_______________

Assistant in Molecular BiologySeed Lab, Massachuse s General Hospital

Charles Gilbert, M.D./Ph.D.B.A., Amherst College

Entered 1971 * Defended July 22, 1976Advisor: David Hubel

Laminar Organiza on of Cat Visual Cortex_______________

Professor of NeurobiologyRockefeller University

Duane Edgington, Ph.D.B.S., California Ins tute of TechnologyEntered 1973 * Defended July 28, 1978

Advisor: Anne StuartProper es and Distribu on of Ionic Channels in a Photoreceptor Neuron

_______________So ware Engineering Group Leader

Monterey Bay Aquarium Research Ins tute

Patrick Hogan, Ph.D.A.B., Harvard College

Entered 1973 * Defended September 29, 1983Advisor: David Po er

Expression of Markers for Pain Sensory Neurons in Cell Culture_______________

Professor, Division of Signaling and Gene ExpressionLa Jolla Ins tute for Allergy and Immunology

Page 6: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Crispin Weinberg, Ph.D.S.B., S.M., University of Chicago

Entered 1973 * Defended August 21, 1980Advisor: Zachary Hall

Molecular Specializa ons at Mature and Developing NeuromuscularJunc ons in Adult Rats_______________

President and FounderBiomedical Modeling, Inc.

Kathleen Buckley, Ph.D.A.B., Radcli e College

Entered 1974 * Defended October 30, 1980Advisor: Story Landis

Morphological and Func onal Studies of Synapses and Varicosi esin Cultures of Dissociated Rat Sympathe c Neurons

_______________Associate Provost for Science

Harvard University

David Ferster, Ph.D.Sc.B., Brown University

Entered 1973 * Defended October 23, 1979Advisor: David Hubel

Cor cal Mechanisms of Depth Vision in the Cat_______________Professor Emeritus

Northwestern University

Steve Hauger, Ph.D.B.s., Stanford University

Entered 1974 * Defended January 14, 1981Advisor: Edward Furshpan

The Spa al Distribu on of Sensi vity to Acetylcholine of RatSympathe c Neurons in Cell Culture

_______________Stained Glass Ar san

Page 7: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Margaret Livingstone, Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1974 * Defended February 20, 1980

Advisor: Edward KravitzMonoamines in the Lobster: Biochemistry, Anatomy, and Possible

Func onal Role_______________

Professor of NeurobiologyHarvard Medical School

Patricia Walicke, M.D./Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1975 * Defended October 11, 1979

Advisor: Paul Pa ersonRole of Calcium and Cyclic AMP in the Mechanism of the Development

E ects of Neuronal Ac vity_______________

Medical O cerNa onal Ins tute of Neurological Disorders and Stroke

Sue Schneps, A.M.B.S., Massachuse s Ins tute of Technology

Entered 1976 * Le June 1, 1980_______________

Withdrew with Master’s degreeJewelry Designer and Owner,

Sea and Star Designs

Leslie Timpe, Ph.D.A.B., Princeton University

Entered 1976 * Defended September 8, 1982Advisor: Ann Stuart

The Roles of Acetylcholine and Gamma aminobutyrin Acid in the VisualSystem of the Barnacle

_______________Adjunct Faculty

San Mateo Community College District

Page 8: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Paul Desan, Ph.D.A.B., Princeton University

Entered 1977 * Defended June 27, 1984Advisor: Simon LeVay

The Organiza on of the Cerebral Cortex of the Pond Turtle, Pseudemysscripta elegans

_______________Assistant Professor of Psychiatry

Yale University School of Medicine

Thomas Schwarz, Ph.D.A.B., Harvard College

Entered 1977 * Defended June 27, 1983Advisor: Edward Kravitz

Proctolin in the Lobster and Protein Phosphoryla onin a Lobster Muscle_______________

Professor of NeurobiologyHarvard Medical School

Allison Doupe, M.D./Ph.D.DCS, B.Sc., McGill University

Entered 1978 * Defended April 26, 1984Advisor: Paul Pa erson

Role of Glucocor coids and Growth Factors in the Development ofNeural Crest Deriva ves

_______________Deceased October 24, 2014

(Formerly Professor at University of California, San Francisco)

Jonathan Horton, M.D./Ph.D.A.B., Stanford University

Entered 1978 * Defended February 16, 1983Advisor: Simon LeVay

Cytochrome Oxidase Patches: A New Cytoarchitectonic Feature_______________

Professor of OphthalmologyUniversity of California at San Francisco

Page 9: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kathleen King Siwicki, Ph.D.Sc.B., Brown University

M.Phil., Cambridge UniversityEntered 1782 * Defended May 7, 1985

Advisor: Edward KravitzThe Neuropep de, Proctolin, in the Lobster Nervous System:

Cellular Distribu on of Proctolin Like An gens and Biochemical Studies ofIden ed Neurons_______________

Professor of NeurobiologySwarthmore College

Karla Kent, Ph.D.B.S., University of California at Santa Barbara

M.A., Boston UniversityEntered 1979 * Defended October 11, 1985

Advisor: John HildebrandMetamorphosis of the antennal center and the in uence of sensory innerva on

on the forma on of glomeruli in the hawk moth Manduca sexta_______________

Associate ProfessorOregon Health Sciences University

Anne Schneiderman, Ph.D.B.S., Stanford University

Entered 1979 * Defended July 16, 1984Advisor: John Hildebrand

Postembryonic Development of a Sexually Dimorphic Sensory Pathwayin the Central Nervous System of the Sphinx Moth, Manduca Sexta

_______________Patent A orneyHarris Beach LLC

Leslie Stevens, Ph.D.A.B., University of California at BerkeleyEntered 1979 * Defended March 27, 1987

Advisor: Story LandisPostembryonic Development of a Sexually Dimorphic Sensory Pathway

in the Central Nervous System of the Sphinx Moth, Manduca Sexta_______________

Research Assistant ProfessorUniversity of Texas at Aus n

Page 10: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

James Hue ner, Ph.D.B.S., B.A., Indiana University of PennsylvaniaEntered 1981 * Defended January 15, 1987

Advisor: Robert BaughmanProper es of Iden ed Cor cocollicular Neurons in Primary Cultures of

Rat Visual Cortex_______________

Professor of Cell BiologyWashington University School of Medicine

Susan McConnell, Ph.D.B.A., Radcli e College

Entered 1981 * Defended March 16, 1987Advisor: Simon LeVay

Fates of Transplanted Visual Cor cal Neurons_______________

Professor of BiologyStanford University School of Medicine

Dinah Sah, Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1981 * Defended February 12, 1987

Advisor: David Po erSerotonin and Neuropep des in Cultured Principal Neurons of the Rat

Superior Cervical Ganglion_______________

Senior Vice President of NeuroscienceVoyager Therapeu cs

Daniel Tso, Ph.D.A.B., Harvard College

Entered 1980 * Defended May 7, 1987Advisor: Torsten Wiesel

Connec vity and Func onal Organiza on in the Mammalian VisualCortex

_______________Associate Professor

Upstate Medical University of the State University of New York

Page 11: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Leonard Levin, M.D./Ph.D.A.B., Harvard College

Entered 1982 * Defended April 28, 1988Advisor: Howard Weiner

Immune Func ons of Astrocytes_______________

Professor of NeurologyUniversity of Wisconsin—Madison

Michael Shepley, M.D./Ph.D.B.S., Northeastern University

M.S.B.E., Massachuse s Ins tute of TechnologyEntered 1982 * Defended November 21, 1988

Advisor: Edward KravitzMonoclonal an body iden ca on of a 100 kDa membrane protein in

HeLa cells and human spinal cord involved in poliovirus a achment_______________

Deceased October 15, 2000(formerly Assistant Professor at SUNY Stony Brook)

David Turner, Ph.D.B.A., Northwestern University

Entered 1982 * Defended March 21, 1989Advisor: Connie Cepko

Cell Lineage in the Rat and Mouse Re nas_______________

Associate Research ProfessorUniversity of Michigan Medical School

Alfred Sandrock, M.D./Ph.D.B.A., Stanford University

Entered 1981 * Defended February 12, 1987Advisor: William Ma hew

The Immunologic Iden ca on of Molecules which Promote theRegenera on of Neuronal Axons

_______________Senior Vice President and Medical O cer

Biogen IDEC, Inc.

Page 12: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Mary Womack, Ph.D.B.A., Northwestern University

Entered 1982 * Defended April 16, 1990Advisor: Thomas Jessell

Mechanisms Underlying Modula on of Dorsal Horn Neuronsby Substance

_______________Instructor in Math and Biology

Nashville State Community College

Junying Yuan, Ph.D.B.S., Fudan University

Entered 1981 * Defended May 9, 1989Advisor: Robert Horvitz

Gene c and Molecular Studies of Two Genes that ControlProgrammed Cell Death in the Nematode C. Elegans

_______________Professor of Cell BiologyHarvard Medical School

Ben Barres, Ph.D.S.B., Massachuse s Ins tute of Technology

M.D., Dartmouth Medical SchoolEntered 1983 * Defended March 1, 1990

Advisor: David CoreyElectrophysiology of White Ma er Glia

_______________Professor of Neurobiology

Stanford University School of Medicine

O o Von FranqueB.A., Yale University

Entered 1982 * Le August 24, 1984_______________

Withdrew without taking degreeEnrolled in M.D. program at Stanford University

Physician

Page 13: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

David Frim, M.D./Ph.D.A.B., Harvard CollegeB.J.Ed., Hebrew College

Entered 1983 * Defended September 1, 1988Advisor: Mar n Moore Ede

Studies in the Structure and Regula on of the mRNA SpeciesEncoding Cor cotroping Releasing Hormone and Vasopressin

_______________Professor of Surgery and Neurology

University of Chicago School of Medicine

Timothy Laman, A.M.B.S., Hope CollegeEntered 1983

_______________

Transferred to OEB at Harvard CollegeWildlife Photojournalist, Na onal Geographic

Peggy Mason, Ph.D.B.A., Harvard College

Entered 1983 * Defended May 14, 1987Advisor: Edward Kravitz

The Intracellularyl Recorded Ac vity of Medullary Raphe NeuronsDuring a Nocicep ve Trigeminial Re ex

_______________Professor of Neurobiology

University of Chicago

Anthony Monaco, M.D./Ph.D.A.B., Princeton University

Entered 1983 * Defended May 13, 1987Advisor: Edward Kravitz

Iden ca on and Par al Cloning of the Locus Responsefor Duchene and Becker Muscular Dystrophy

_______________President

Tu s University

Page 14: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

E. Houston Warren, Ph.D.A.B., Harvard College

Entered 1983 * Defended September 19, 1988Advisor: Charles Liberman

E ect of Sound Evoked Ac vity in Olivocochlear E erent Neuronson Auditory Nerve Response

_______________Associate Professor of Medical OncologyUniversity of Washington Medical School

Daniela BonafedeB.S., California Ins tute of Technology

Entered 1984 * Le July 1, 1986_______________

Withdrew without taking degreeTransferred to Yale University

Steven McIn re, M.D./Ph.D.B.S., M.S., Stanford University

Entered 1984 * Defended March 1, 1990Advisor: Robert Horvitz

A Gene c Characteriza on of GABAergic Neurotransmission and AxonalOutgrowth in the Nematode Caenorhabdi s elegans

_______________Consul ng Professor, Neurology

Stanford University School of Medicine

Laura Regan, Ph.D.B.A., Swarthmore College

Entered 1983 * Defended January 10, 1990Advisor: Bruce Bean

Calcium Currents in Freshly Dissociated Rat Purkinje and Dorsal RootGanglion Cells

_______________Dean, School of Classical Chinese Medicine

Na onal College of Chinese Natural Medicine

Page 15: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Carmela Abraham, Ph.D.B.Sc., Tel Aviv University

Entered 1985 * Defended January 12, 1988Advisor: Hun ngton Po er

The protease inhibitor, 1 an chymotrypsin, in brain aging and disease_______________

Professor of BiochemistryBoston University School of Medicine

John Assad, Ph.D.B.A., State University of New York at Bu aloEntered 1985 * Defended June 20, 1991

Advisor: David CoreySensory Adapta on of Vertebrate Hair Cells

_______________Professor of Neurobiology

Harvard Medical School

David Lopes Cardozo, Ph.D.B.A., Concordia UniversityB.Sc., Dalhousie University

Entered 1985 * Defended June 1, 1993Advisor: David Po er

Pharmacology of Synap c Transmission in the Gold sh Sacculus_______________

Associate Dean, Graduate Student A airsDivision of Medical Sciences, Harvard Medical School

Anna Wang RoeA.B., Harvard College

Entered 1984 * Le May 13, 1985_______________

Withdrew without taking degreeEnrolled in MIT and earned Ph.D. in Neuroscience

Professor of Neuroscience, Oregon Health & Sciences University

Page 16: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Chinfei Chen, M.D./Ph.D.B.A.S., University of Pennsylvania

Entered 1985 * Defended September 21, 1990Advisor: Peter Hess

Voltage Dependent Calcium Channels in 3T3 Fibroblasts_______________

Associate Professor of NeurologyHarvard Medical School

Sheila Nirenberg, Ph.D.B.A., State University of New York at Albany

Entered 1985 * Defended June 1, 1992Advisor: Connie Cepko

Targeted Abla on of Diverse Cell Types in the Nervous System:A Method for Blocking Cell Cell Interac ons in vivo

_______________Professor of Physiology

Cornell Weill School of Medicine

Philip Starr, M.D./Ph.D.B.A., Princeton University

Entered 1985 * Defended March 20, 1989Advisor: William Sewell

Pharmacology of Synap c Transmission in the Gold sh Sacculus_______________

Professor of Neurological SurgeryUniversity of California at San Francisco Medical Center

Rudolph Tanzi, Ph.D.B.S., University of Rochester

Entered 1985 * Defended March 19, 1990Advisor: James Gusella

The Amyloid Beta Protein Precursor Gene in Familial and SporadicAlzheimer’s Disease_______________

Professor of Child NeurologyHarvard Medical School

Page 17: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Margaret Thompson, Ph.D.B.S., University of Michigan

Entered 1985 * Defended August 2, 1993Advisor: Michael E. Greenberg

Serotonergic Neurons in the Lobster: Excitatory and Inhibitory Inputs_______________

Director, IDDRC CoreBoston Children’s Hospital

Chris an Titlow, Ph.D.B.S., California Ins tute of TechnologyEntered 1985 * Defended June 8, 1992

Advisor: Marge LivingstoneAssembly of Human Monoamine Oxidase Type B (MAO B) Into the

Mitochondrial Outer Membrane:Support for a New Class of Leader Sequences

_______________

Whereabouts Unknown

Margot U erbackB.S., University of California at Los AngelesEntered 1985 * Le September 1, 1992

Advisor: Lydia Villa Komaro_______________

Withdrew without taking degreeNot employed

Wendy Weiger, M.D./Ph.D.A.B., Harvard College

Entered 1985 * Defended February 14, 1995Advisor: Edward Kravitz

Serotonergic Neurons in the Lobster: Excitatory and Inhibitory Inputs_______________

Physician, author, wilderness guide,and environmental ac vist

Page 18: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Mel Feany, Ph.D.A.B., Harvard College

Entered 1986 * Defended April 29, 1993Advisor: Kathleen Buckley and Margaret Livingstone

Molecular Analyses of Synap c Vesicle Proteins/Gene c and Molecular Studies of Drosophila Learning Mutants

_______________Professor of PathologyHarvard Medical School

Carl Fulwiler, Ph.D.B.A., Hofstra University

M.D., Washington UniversityEntered 1986 * Defended May 31, 1995

Advisor: John DowlingGene c Analysis of Re nal Development in Zebra sh

_______________Associate Professor of Psychiatry

University of Massachuse s Medical School

Linda Kobierski, Ph.D.B.S., Simmons College

Entered 1986 * Defended May 5, 1993Advisor: Michael Comb

Proenkephalin Gene Expression: Mechanisms of cAMP Regula on_______________

High School Science TeacherDedham Public Schools

Julie Nardone, Ph.D.A.B., Harvard College

Entered 1986 * Defended April 20, 1993Advisor: David Po er

Site Directed Mutagenesis of a B2 Bradykinin Receptor_______________

Senior Research Scien st, Bioinforma csMetamark Gene cs, Inc.

Page 19: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Bradford Navia, Ph.D.B.A., M.D., Columbia University

Entered 1986 * Defended August 12, 1995Advisor: Gail Bruns

Molecular Analysis of the Region Surroundingthe X Linked Re ni s Pigmentosa (RP3) Locus in Xp21.1

_______________Whereabouts unknown

Formerly Associate Professor at Tu s University School of Medicine

Pamela Prosser FriedmanB.A., Dartmouth College

Entered 1986 * Le June 1, 1987_______________

Withdrew without taking degreeWhereabouts unknown

Allan RosenbergB.S., University of Michigan

Entered 1986 * Le September 1, 1990_______________

Withdrew without taking degreeEarned J.D. at Stanford Law School

Assistant Vice President, State Street Global Exchange

Hung Ming Chu, Ph.D.B.M., Na onal Taiwan University

Entered 1987 * Defended June 1, 1992Advisor: Michael Comb

Proenkephalin Gene Regula on: S mulus Transcrip on Coupling_______________Medical Director

Chinatown/North Beach Mental Health Clinic (San Francisco)

Page 20: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Elizabeth Finch, Ph.D.B.A., University of California at Santa Barbara

Entered 1987 * Defended May 31, 1994Advisor: Stan Goldin

Inositol Trisphosphate Mediated Calcium Release: Kine cs andRegula on by IP, Calcium, and ATP

_______________Whereabouts unknown

Formerly Assistant Professor of Cell Biology at Emory University

Michael Gille e, M.D./Ph.D.Oxford University

Entered 1987 * Defended May 4, 1995Advisor: Raymon Dacheux

Components of Adapta on in Saccular Hair Cellsof the Bullfrog, Rana catesbeiana

_______________Scien st, Broad Ins tute

Instructor, Harvard Medical SchoolPhysician, Pulmonary Care Unit, MGH

Susan Lewis, Ph.D.A.B., Harvard College

Entered 1987 * Defended June 7, 1994Advisor: Steven Hyman

Cytokine Regula n of Neuronal Gene Expression in Sympathe cNeurons and a Neuroblastoma Cell Line

_______________Grant writer and editor

Tu s University

Emily Liman, Ph.D.A.B., Princeton University

Entered 1987 * Defended March 31, 1992Advisor: Peter Hess

Structure and Func on of Mammalian Potassium Channels_______________

Professor of Biological SciencesUniversity of Southern California

Page 21: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Gordon Shepherd, M.D./Ph.D.A.B., Vassar College

Entered 1987 * Defended July 27, 1993Advisor: David Corey

Components of Adapta on in Saccular Hair Cellsof the Bullfrog, Rana catesbeiana

_______________Associate Professor in Physiology

Northwestern University School of Medicine

Doreen Valen ne, Ph.D.B.S., State University of New York at Stony BrookEntered 1987 * Defended February 27, 1995

Advisor: Cynthia MossSpa al Percep on and Sensorimotor Integra on: The Role of the

Superior Colliculus in the Echoloca ng Bat, Eptesicus fuscus_______________

Associate Scien c DirectorClinical Thinking/Nucleus Global

Dayao Zhao, Ph.D.M.D., Beijing Medical University

Entered 1987 * Defended October 31 1984Advisor: William Chin

Re noic Acid: Biosynthesis and Receptors_______________

Vice PresidentJanssen Pharmaceu cals (China)

Marina Chicurel, Ph.D.Universidad Nacional Autónoma de MexicoEntered 1988 * Defended May 12, 1995

Advisor: Hun ngton Po erRNA Localiza on and RNA Cytoskeletal Interac ons in Neurons

_______________Associate Director for Science

Hereditary Disease Founda on

Page 22: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Robin Cloues, A.M.B.A., Barnard College

Entered 1988 * Le November 1, 1991_______________

Withdrew with Master’s degreeTransferred to University of London PhD program

Adjunct Faculty, Regis University

Shing Chuan Hooi, Ph.D.M.B.B.S., Na onal University of SingaporeEntered 1988 * Defended March 23, 1992

Advisor: Lee KaplanNeuroendocrine Regula on of Galanin Expression

_______________Professor and Vice Dean of Educa on

Yong Loo Lin School of MedicineNa onal University of Singapore

Mark Johnson, M.D./Ph.D.A.B., Amherst College

Entered 1988 * Defended July 27, 1993Advisor: David Po er

Electrophysiological and Cytochemical Proper esof Rat Mesopon ne Serotonergic Neurons in Culture

_______________Associate Professor of Neurosugery

Harvard Medical School

Tejvir Khurana, Ph.D.M.B.Bs., Delhi University

Entered 1988 * Defended August 12, 1991Advisor: Louis Kunkel

Iden ca on and Characteriza on of Dystrophin Related Protein_______________

Professor of PhysiologyUniversity of Pennsylvania School of Medicine

Page 23: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Chung Chin Kuo, Ph.D.B.M., Na onal Taiwan University

Entered 1988 * Defended May 1, 1992Advisor: Peter Hess

Ion Permea on through L Type Calcium Channel_______________

Professor of NeurophysiologyNa onal Taiwan University, College of Medicine

Nicholas Marsh Armstrong, Ph.D.B.A., Haverford College

Entered 1988 * Defended June 1, 1994Advisor: Walter Gilbert

Re noic Acid and the Pa erning of the Zebra sh Re na and Trunk_______________Research Scien st

Kennedy Krieger Ins tute

Bruce Peters, Ph.D.B.S., Rensselaer Polytechnic Ins tute

Entered 1988 * Defended April 12, 1996Advisor: Richard Masland

Responses to Light of Starburst Amacrine Cellsin the Re na of the Rabbit

_______________Senior Scien stINDEC Systems

Sonja Potrebic, M.D./Ph.D.B.S., University of Pennsylvania

Entered 1988 * Defended February 28, 1994Advisor: Howard Fields

The Role of Serotonin in the Modula on of Nocicep onby the Rostral Ventromedial Medulla of the Rat

_______________Neurologist

Southern California Kaiser Permanente Medical Group

Page 24: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

David Welsh, M.D./Ph.D.B.S., Stanford University

Entered 1988 * Defended April 1, 1995Advisor: Steven Reppert

Circadian Clock Neurons in Culture_______________

Associate Professor of PsychiatryUniversity of California at San Diego School of Medicine

Cynthia WetmoreA.B., Harvard College

Entered 1988 * Le June 1, 1990_______________

Withdrew without taking degreeTransferred to Karolinska Ins tuteAssociate Professor of Pediatrics

Emory University School of Medicine

Andrew Ahn, M.D./Ph.D.B.S., Columbia University

Entered 1989 * Defended May 8, 1995Advisor: Louis Kunkel

The Membrane Cytoskeleton in Muscular Dystrophy_______________

Senior Medical Advisor and Scien c LeadEli Lilly & Co.

Belinda Chang, Ph.D.A.B., Princeton University

Entered 1989 * Defended September 26, 1995Advisor: Naomi Pierce

Opsin Phylogeny and Evolu on_______________

Associate Professor of Ecology and Evolu onary BiologyUniversity of Toronto

Page 25: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Richard Chung, M.D./Ph.D.A.B., Harvard College

Entered 1989 * Defended August 11, 1992Advisor: Bernd R. Sseizinger

P53: A Bifunc onal Transcrip onal Regulatory ProteinInac vated in Human Astrocy c Tumors

_______________Neurosurgeon

Neurosurgical Associates of Santa Barbara

Lise Heginbotham Thomas, Ph.D.B.A., Swarthmore College

M.S., University of Colorado Health Sciences CenterEntered 1989 * Defended May 12, 1993

Advisor: Roderick MacKinnonIden ca on and Molecular Characteriza on

of a Potassium Channel Pore_______________

Associate Professor of BiologyQuinnipiac University

Albert Hung, M.D./Ph.D.A.B., Harvard College

Entered 1989 * Defended March 31, 1994Advisor: Dennis Selkoe

Regula on of Expression and Processing of the b Amyloid PrecursorProtein of Alzheimer’s Disease

_______________Assistant Professor of Neurology

Harvard Medical School

Raul Krauss, Ph.D.M.Sc., Catholic University of Chile

Entered 1989 * Defended May 19, 1997Advisor: Gerald Fishbach

Studies of Nico nic Acetylcholine Receptor Gene Expressionin the Central Nervous System

_______________Co Head North America Neurology Disease Team

Vertex Pharmaceu cals

Page 26: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Camilo Mar nez, M.D./Ph.D.B.S., University of Southern CaliforniaEntered 1989 * Defended May 1, 1994

Advisor: Joseph MajzoubMolecular Regula on of Cor cotropin Releasing Hormone GeneExpression by the Protein Kinase A Pathway and Glucocor coids

_______________Partner

QM VEntures

Kenton Swartz, Ph.D.B.Sc., Eastern Manitoba College

M.D., Boston University School of MedicineEntered 1989 * Defended March 22, 1993

Advisor: Bruce BeanRegula on of Neuronal Ca Channels by Neurotransmi ers,

G Proteins and Kinase C_______________Senior Inves gator

Na onal Ins tute of Neurological Disorders and Stroke

Anne Sydor, Ph.D.B.A., University of Wisconsin

Entered 1989 * Defended June 20, 1995Advisor: Daniel Jay

The Role of Cytoskeletal Proteins Talin and Vinculinin Neuronal Growth Cone Mo lity

_______________Editorial Director, Educa onal Products

Thieme Medical Publishers

Azad Bonni, Ph.D.M.D., Queen’s University

Entered 1990 * Defended May 9, 1996Advisor: Michael Greenberg

Neurotrophic Factor Regula on of Immediate Early Gene Expression_______________

Professor of NeurobiologyWashington University School of Medicine

Page 27: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Rebecca Leah Cole, Ph.D.B.A., Macalester College

Entered 1990 * Defended July 17, 1996Advisor: Steven Hyman

Neurotransmi er Regula on of CRE– and AP 1 MediatedGene Expression in Rat Striatum

_______________Research Scien st

Merck Research Laboratories

Paul Jackson, M.D./Ph.D.B.S., Stanford University

Entered 1990 * Defended June 29 1995Advisor: Larry Benowitz

VSOAC: A Volume Sensi ve Anion Channel with a Novel Func on_______________

NeurosurgeonPalo Alto, California

Roger Knowles, Ph.D.B.S., United States Military Academy

Entered 1990 * Defended December 19, 1995Advisor: Kenneth Kosik

Transloca on of RNA Granules in Living Neurons_______________

Professor of NeuroscienceDrew University

Inge KnudsonB.A., Smith College

M.S., University of New HampshireEntered 1990 * Le May 6, 1993

_______________Withdrew without taking degree

Associate Scien st, Massachuse s Eye and Ear In rmary

Page 28: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Mary Morris Saba ni, Ph.D.B.A., Agnes Sco CollegeM.D., Columbia University

Entered 1990 * Defended July 31, 1997Advisor: Chuck Weitz

Entertainment of the Mammalian Circadian Clock: Iden ca onof Light Induced Genes in the SCN

_______________Obstetrician

Massachuse s General Hospital

Caroline Overly, Ph.D.A.B., Harvard College

Entered 1990 * Defended May 7, 1996Advisor: Peter Hollenbeck

Organelle Mo lity and Endocy c Pathways in Cultured Neurons_______________

Community VolunteerSea le, Washington

Alfred Roca, Ph.D.B.A., Cornell University

B.S., University of WyomingEntered 1990 * Defended August 7, 1996

Advisor: Steven ReppertThe Mouse Mel1a Melatonin Receptor Gene

_______________Associate Professor of Animal Sciences

University of Illinois at Urbana Champaign

Karin Sorra, Ph.D.B.Sc., Dalhousie University

Entered 1990 * Defended November 26, 1996Advisor: Kristen Harris

Synapse Structure, Func on and Plas city in Hippocampal Area CA1_______________

President and Chief Scien c O cerArroscience, Inc.

Page 29: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ashley Bruce, Ph.D.A.B., Brown University

Entered 1991 * Defended August 22, 1997Advisor: Mar n Shankland

Characteriza on of an orthoden cle Homolog in the Leechand Inves ga on of Ectopic Gene Express Techniques

_______________Associate Professor of Cell Biology

University of Toronto

Anne e Jenner, Ph.D.B.A., Mt. Holyoke College

Entered 1991 * Defended April 28, 1998Advisor: Gordon Sherman

The Connec vity of Ectopic Neurons in the Neocortex and their E ectson Neuronal Size in the Thalamus: Implica ons for Developmental

Dyslexia_______________

Adjunct Assistant ProfessorSUNY Upstate Medical Center

Stephen Macknik, Ph.D.B.A., University of California at Santa Cruz

Entered 1991 * Defended September 20, 1996Advisor: Marge Livingstone

The Physiology and Percep on of Visibility and Visual Masking_______________

Professor of Ophthalmology, SUNY Downstate Medical CenterColumnist for Scien c American

Phillip Schwartz, Ph.D.B.S., Columbia University

M.Sc., The Weizmann Ins tuteEntered 1991 * Defended June 17, 1998

Advisor: Rosalind SegalNeurotrophins in the Regula on of Cerebellar Development

_______________Chief Execu ve O cer

Entera Bio Ltd.

Page 30: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Patrick Senatus, M.D./Ph.D.A.B., Harvard College

Entered 1991 * Defended June 16, 1997Advisor: David Po er

Electrophysiological and Cytochemical Proper es ofRat Neostriatal Neurons in Microculture

_______________

NeurosurgeonConnec cut

Lawrence Sincich, Ph.D.B.S., Stanford University

Entered 1991 * Defended May 13, 1999Advisor: Gary Blasdel

Local Circuits in Primary Visual Cortex_______________

Assistant Professor of Vision ServicesUniversity of Alabama at Birmingham School of Optometry

Deborah Sodickson Cohen, Ph.D.B.A., Wesleyan University

Entered 1991 * Defended January 8, 1997Advisor: Bruce Bean

Control of Inwardly Rec fying Potassium Channels and VoltageDependent Calcium Channels

by G Protein Coupled Transmi er Receptors in Hippocampal Neurons_______________

Homemaker

Mark Tramo, Ph.D.B.A., Yale University

M.D., Cornell UniversityEntered 1991 * Defended November 10, 1998

_______________Neurologist

Lecturer, U.C.L.A. School of Music

Page 31: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Cathy WaldmanB.A., Arcadia University

Entered 1991 * Withdrew without degree August 31, 1992_______________Speech Pathologist

Deborah Watson, Ph.D.B.A., Mills College

Entered 1991 * Defended April 11 1997Advisor: Dennis Selkoe

E ects of the HCHWA D Muta on on bAPP Processing, Heterogeneity ofAb Deposits, and Heparin/HSPG Binding: Implica ons for Alzheimer’s

Disease_______________

Research Assistant Professor of NeurosurgeryUniversity of Pennsylvania School of Medicine

Anne West, M.D./Ph.D.B.A., Cornell University

Entered 1991 * Defended December 2, 1996Advisor: Kathleen Buckley

Protein Targe ng in Neurons_______________

Associate Professor of NeurobiologyDuke University

Robert WhorfB.S., University of New Hampshire

Entered 1991 * Le September 1, 1993_______________

Enrolled in University of Rochester School of MedicineOncologist, Florida

Page 32: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Sanjay Aggarwal, M.D./Ph.D.B.S., Johns Hopkins University

Entered 1992 * Defended June 3, 1996Advisor: Roderick MacKinnon

Iden ca on of the Voltage Sensor in the Voltage Ac vated PotassiumChannel

_______________Pediatrician

Bal more, Maryland

Marc Awobuluyi, M.D./Ph.D.A.B., Harvard College

Entered 1992 * Defended June 10, 1997Advisor: Stuart Lipton

Transla onal Control of N Methyl D Aspartate Subunit 1 (NMDAR1)Expression

_______________Assistant Clinical Professor in Neuroradiology

University of California, San Francisco Medical Center

Kris na Wietasch Becker, Ph.D.B.A., M.D., Ruhr University

Entered 1992 * Defended March 21, 1997Advisor: Gerald Fischbach

Regula on of Neuronal Nico nic Acetylcholine Receptorsin the Interpeduncular Nucleus

_______________Patent A orney

Germany

Michael Belliveau, Ph.D.B.S., University of Michigan

Entered 1992 * Defended September 9, 1998Advisor: Connie Cepko

Intrinsic and Extrinsic Regula on of Cell Fate Choices in the VertebrateNeural Re na

_______________Patent A orney and Partner

Clark & Elbing LLP

Page 33: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Solange Brown, M.D./Ph.D.Princeton University

Entered 1992 * Defended June 7, 1999Advisor: Richard Masland

Local Circuits in the Re na: Spa al Sensi vity, Contrast Adapta onand Direc on Selec vity of Re nal Ganglion Cells

_______________Assistant Professor of Neuroscience

Johns Hopkins University School of Medicine

Yu Ju ChenB.S., Na onal Taiwan University

Entered 1992* Le January 1, 1995_______________

Withdrew without taking degreeAdjunct Professor of Proteomics and Mass Spectrometry

Na onal Taiwan University

Donnella Green Comeau, M.D./Ph.D.B.A., Amherst College

Entered 1992 * Defended September 19, 1997Advisor: Stephen Cannon

Func onal Consequences of Sodium Channel Muta onsin Human Neuromuscular Disease

and a Novel Mechanism for Acetazolamide Therapy_______________

RadiologistCumberland, Rhode Island

Amy Malick, Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1992 * Defended May 4, 2000

Advisor: Rami BursteinOn Trigeminohypothalamic Pain Pathways and How They Contribute to

Anorexic Behavior_______________

Managing Editor and Group PublisherQuan aMD

Page 34: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Michael Olivares, M.D.B.S., University of Texas at Aus n

Entered 1992 * Le February 1, 1993_______________

Withdrew without taking degreeReturned to Harvard Medical School to complete M.D. degree

Whereabouts unknown

Christopher Reid, Ph.D.B.S., United States Military Academy

Entered 1992 * Defended January 9, 1997Advisor: Christopher A.. Walsh

Cell Lineage in Mammalian Cerebral Cortex_______________

Chief Execu ve O cerPDC Capital Group

Kerry Ressler, M.D./Ph.D.B.S., Massachuse s Ins tute of Technology

Entered 1992 * Defended December 18, 1995Advisor: Linda Buck

Odorant Receptor Organiza on and Informa on Codingin the Mouse Olfactory System

_______________Professor of PsychiatryHarvard Medical School

Volney Sheen, M.D./Ph.D.Johns Hopkins University

Entered 1992 * Defended May 6, 1997Advisor: Je rey Macklis

Transplanta on Strategies for Reconstruc onof Complex Neocor cal Circuitry

_______________Assistant Professor of Neurology

Harvard Medical School

Page 35: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Paul Tolen no, M.D./Ph.D.Sc.B., Brown University

Entered 1992 * Defended May 27, 1995Advisor: Lydia Villa Komaro

Transcrip onal and Post Transcrip onal Regula onof VIP mRNA Expression

_______________Neurosurgeon

Cape Girardeau, Missouri

Kelly Van Koughnet, Ph.D.B.Sc., University of Toronto

Entered 1992 * Defended March 12, 1999Advisor: Steve Hyman

Proenkephalin Transgene Regula on in the Paraventricular Nucleus ofthe Hypothalamus by Lipopolysaccharide and Interleukin 1ß: a Model

for the Study of Neuro Immune Communica on_______________

Associate Vice President of Research and Knowledge Transla oinCanadian Ins tutes of Health Research

Brenda Allwardt, Ph.D.B.A., Lawrence University

Entered 1993 * Defended May 6, 1999Advisor: John Dowling

Ultrastructural Analysis of Photoreceptorsin Wildtype and Mutant Zebra sh Larvae

_______________Not currently employed

Joshua Berke, Ph.D.B.A., Cambridge University

Entered 1993 * Defended September 10, 1998Advisor: Steve Hyman

A Complex Program of Dopamine Induced Gene Expression Iden edUsing Di eren al Display PCR: Discovery of a Novel Inducible Cyclin

_______________Associate Professor of Psychology and Neuroscience

University of Michigan

Page 36: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Steven Chao, M.D./Ph.D.A.B., Harvard College

Entered 1993 * Defended August 24, 1999Advisor: Larry Benowitz

GAP 43: Regula on and Func on_______________

PartnerClearView Healthcare Partners

Katrin Chua, M.D./Ph.D.A.B., Harvard College

Entered 1993 * Defended May 7, 1999Advisor: Robin Reed

Remodeling of the Spliceosome for Catalysis of Exon Liga on_______________

Associate Professor of MedicineStanford University School of Medicine

Saumya Das, M.D./Ph.D.A.B., Harvard College

Entered 1993 * Defended September 9, 1997Advisor: Nobuki Nakanishi

Iden ca on and Characteriza on of a Novel N Methyl D AspartateReceptor Subunit NR3A

_______________Researcher and Electrophysiologist

Beth Israel Deaconess Medical Center

Michael Bianche a, Ph.D.B.A., Brown University

Entered 1993 * Defended July 23, 2004Advisor: Jonathan Cohen

Self Associa on of the Muscle Speci c Receptor Tyrosine Kinase MuSK_______________

Whereabouts unknown

Page 37: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jorge Luis Gonzalez, Ph.D.B.S., Tu s University

Entered 1993 * Defended May 10, 1999Advisor: Christopher A. Walsh

Cellular and Biochemical Analysis of Muta ons that A ect CerebralCor cal Development_______________

Execu ve Vice President of Systems and Technology Innova onEF Educa onal Tours

Heather HindsA.B., B.S., Stanford University

M.Sc., Oxford UniversityEntered 1993 * Le May 1, 1994

_______________Transferred to Massachuse s Ins tute of Technology– Tonegawa Lab

Neurologist — San Francisco, California

Lisa Horowitz, M.D./Ph.D.B.S., University of California at Berkeley

Entered 1993 * Defended September 14, 1998Advisor: Linda Buck

Connec vity in the Mouse Olfactory System_______________

PhysicianBoston, Massachuse s

Holly Engelman, A.M.B.S., Columbia University

Entered 1993 * Le January 1, 1996_______________

Transferred to Columbia UniversityReceived Ph.D. in 2002

Research support sta at Pennsylvania State College of Medicine

Page 38: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jerri Mar n, A.M.B.S., California Ins tute of TechnologyEntered 1993 * Le June 1, 1997

_______________Withdrew with Master’s degree

Elementary school volunteer; formerly scien st at Theravance, Inc.San Francisco, California

Mar n Niethammer, Ph.D.Diploma, Universitat Tubingen

Entered 1993 * Defended September 3, 1998Advisor: Morgan Sheng

Organiza on of an NMDA Receptor Associated Complex via the PDZDomains of PSD 95_______________

Adjunct Clinical Assistant ProfessorAlbert Einstein College of Medicine

Lianna Orlando, Ph.D.B.S., Syracuse University

Entered 1993 * Defended April 25, 2001Advisor: Anne Young

Cellular and Molecular Mechanisms of Neuronal Chain Migra on_______________

Scien c Program O cerMuscular Dystrophy Associa on

Thomas Jordan, Ph.D.Bucknell University

Entered 1993 * Defended December 8, 1997Advisor: John Dowling

Network Dynamics and Their Neuromodula on: Dopaminergic E ectson Re nal Recep ve Fields

_______________Founder and Chief Execu ve O cer

Aldexa Therapeu cs

Page 39: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Heather Rie , Ph.D.B.A., Williams College

Entered 1993 * Defended August 13, 1999Advisor: Gabriel Corfas

The Role of Neuregulin and erB Receptors in Cerebellar Development_______________

Senior Science Policy AnalystNa onal Ins tute of Neurological Disorders and Strokes

Bernardo Saba ni, M.D./Ph.D.A.B., Harvard College

Entered 1993 * Defended September 16, 1997Advisor: Wade Regehr

Calcium Control of Neurotransmi er Release at a Cerebellar Synapse_______________

Professor of NeurobiologyHarvard Medical School

Maurice Sholas, M.D./Ph.D.B.S., Southern University

Entered 1993 * Defended June 28, 1996Advisor: Daniel Kenshalo

Changes in Primary Somatosensory Cortex Processing of Nocicep veInforma on

Induced by In amma on_______________

Senior Medical DirectorChildren’s Hospitals and Clinics of Minnesota

Megan Benson Pra , Ph.D.B.S., Massachuse s Ins tute of Technology

M.A., Harvard UniversityEntered 1993 * Defended September 13, 1999

Advisor: Jonathan CohenMapping Noncompe ve Antagonist Binding Sites in the Nico nic

Acetylcholine Receptor_______________

Educa onal Outreach CoordinatorIns tute for Human and Machine Cogni on

Page 40: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jamie Zeitzer, Ph.D.B.A., Vassar College

Entered 1993 * Defended March 26, 1999Advisor: Charles Czeisler

Physiology and Anatomy of Human Circadian Photorecep onand Melatonin Regula on

_______________Assistant Professor of Psychiatry

Stanford University School of Medicine

Pradeep Atluri, M.D./Ph.D.A.B., Harvard College

Entered 1994 * Defended January 27, 2000Advisor: Wade Regehr

The Role of Presynap c Calcium in Short Term Synap c Plas city_______________

PsychiatristNew York City

Stacie Weninger, Ph.D.University of North Carolina

Entered 1994 * Defended September 9, 1999Advisor: Joseph Majzoub

Role of Cor cotropin Releasing Hormone (CRH) and CRH Like Moleculesin Stress Induced Behavior

_______________Execu ve Director

Fidelity Biosciences Research Ini a ve

Fiona Watson, Ph.D.B.Sc., M.S., Dalhousie University

Entered 1993 * Defended May 18, 1999Advisor: Rosalind Segal

Retrograde Signaling by Neutrophins_______________

Associate Professor of BiologyWashington and Lee University

Page 41: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Michelle Day, Ph.D.B.S., University of California, Los AngelesEntered 1994 * Defended June 2, 2000

Advisor: Steve HymanThe E ects of Serotonin 5 HT2A Agonists and Antagonists on High

Voltage Ac vated Calcium Currents in Prefrontal Pyramidal Neurons_______________

Research Assistant ProfessorNorthwestern University Feinberg School of Medicine

Jeremy Di man, M.D./Ph.D.B.S., Stanford University

Entered 1994 * Defended June 2, 1998Advisor: Wade Regehr

Presynap c Modula on of Transmi er Releasein the Central Nervous System

_______________Associate Professor of Biochemistry and Neuroscience

Weill Cornell Medical College

Stephanie Gaus, Ph.D.B.S., Stanford University

Entered 1994 * Defended April 5, 2002Advisor: Cli ord Saper

Neuroanatomic Substrates for Circadian Control of Sleep_______________Research Scien st

University of California, San FranciscoSeeley Selec ve Vulnerability Research Laboratory

Paul Borghesani, M.D./Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1994 * Defended December 20, 1999

Advisor: Rosalind SegalNeuronal Migra on in Development and Degenera on

_______________Psychiatrist

Sea le, Washington

Page 42: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

James Morrill, M.D./Ph.D.A.B., Harvard College

Entered 1994 * Defended July 26, 1999Advisor: Stephen Cannon

Biophysical Proper es of the Skeletal Muscle L Type Calcium Channelin Health and Disease_______________

Instructor in MedicineMassachuse s General Hospital

Paula Smith, PH.D.B.S., Cornell University

Entered 1994 * Defended September 7, 1999Advisor: Gary Yellen

Voltage dependent Ga ng of HERG Potassium Channels_______________

Medical WriterNovo Nordisk

Mark Zylka, Ph.D.B.X., Virginia Polytechnic Ins tute

Entered 1994 * Defended September 17, 1999Advisor: Steven Reppert

Characteriza on of a Mammalian Circadian Feedback Loop_______________

Associate Professor of Cell BiologyUniversity of North Carolina, Chapel Hill

Lisa Genova, Ph.D.B.S., Bates College

Entered 1994 * Defended April 30, 1998Advisor: Steve Hyman

Amphetamine Induced Adapta ons in Striatal Neurons: Roles ofSerotonin and Condi oned S muli

_______________Best selling author

“S ll Alice”, “Le Neglected” and “Inside the O’Briens”

Page 43: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

A.B., Yun Beom Choi, M.D./Ph.D.Harvard College

Entered 1995 * Defended December 21, 1998Advisor: Stuart Lipton

Molecular Analysis of Modulatory Sites on the NMDA Receptor:Redox, Zn2+, and Nitric Oxide Related Species

_______________Neurologist

Englewood, New Jersey

Hui Zhou, A.M.B.S., Huazhong University of Science and Technology

M.S., Beijing UniversityEntered 1994 * Le March 1, 1997

_______________

Withdrew with Master’s degreeTransferred to Washington University

Richard Chris e, M.D./Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1995 * Defended December 21, 1999

Advisor: Bradley HymanIn vivo Imaging of Amyloid Deposi on in a Transgenic Mouse Model

of Alzheimer’s Disease_______________

Independent Consultant (Pharmaceu cals)Obex Enterprises LLC

Manish Aghi, M.D./Ph.D.B.S., Stanford University

Entered 1995 * Defended November 20, 1998Advisor: Xandra Breake eld

Improving the E cacy of Chemosensi zingBrain Tumor Gene Therapies

_______________Associate Professor of Neurosurgery

University of California, San Francisco

Page 44: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jerry Lin, M.D./Ph.D.A.B., Harvard College

Entered 1995 * Defended June 13, 2000Advisor: Morgan Sheng

I. Characteriza on of an NMDA receptor associated complexJ. centered around NRI;

II. The role of AMPA receptor tra cking in synap c plas city_______________OtolaryngologistFairfax, Virginia

Sanjay Magavi, Ph.D.Sc.B., Brown University

Entered 1995 * Defended September 28, 2001Advisor: Je rey Macklis

Induc on of Neurogenesis in the Neocortex of Adult Mice andAssessment of the Func onal Integra on of Adult born Neurons

_______________Senior Research Scien stVertex Pharmaceu cals

Heather Mason Chris e, Ph.D.A.B., Amherst College

Entered 1995 * Defended April 27, 2001Advisor: Gabriel Corfas

Cellular and Molecular Mechanisms of Neuronal Chain Migra on_______________

Homemaker

Jesus Garcia, M.D./Ph.D.B.A., Loyola Marymount University

Entered 1995 * Defended September 10, 1999Advisor: Margaret Livingstone

Mechanisms of Color and Mo on in Monkey Visual Cortex_______________

Plas c SurgeonSacramento, California

Page 45: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Philip Nadeau, Ph.D.B.S., Massachuse s Ins tute of Technology

Entered 1995 * Defended September 15, 2000Advisor: Bruce Yankner

An Inves ga on of the Role of Apolipoproteins, Lipoproteins and thePresenilin Proteins in Amyloid Beta Metabolism and Alzheimer’s Disease

_______________Managing Director and Senior Research Analyst

Cowen and Company

Sco Naisbi , M.D./Ph.D.B.S., University of Utah

Entered 1995 * Defended September 29, 2000Advisor: Morgan Sheng

The Molecular Organiza on of the Postsynap c Densityby GKAP and SHANK_______________

Vice President of Scien c and Medical A airsLutonix, Inc.

Asha Elizabeth Nigh, Ph.D.B.S., Stanford University

Entered 1995 * Defended March 7, 2003Advisor: Michael Greenberg

The Control of Protein Expression During Adap ve Neuronal Responses_______________

Chief Scien c Editor and FounderScience Editors Network

Mireya Nadal Vicens, M.D./Ph.D.A.B., Harvard College

Entered 1995 * Defended September 14, 2001Advisor: Michael Greenberg

Signaling mechanisms that govern the switchfrom neuronal to astrocyte di eren a on in neocor cal progenitors

_______________Psychiatrist and Instructor of Psychiatry

Massachuse s General Hospital

Page 46: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Marcus Ware, M.D./Ph.D.B.S., Tougaloo College

Entered 1995 * Defended November 16, 1998Advisor: Christopher A. Walsh

Cellular and Molecular Events in the Developmentof the Cerebral Cortex_______________

Clinical Assistant Professor of Neurological SurgeryTulane University

Vasanth Vedantham, M.D./Ph.D.B.S., Yale University

Entered 1995 * Defended June 2, 2000Advisor: Stephen Cannon

Moving Parts of Voltage Gated Sodium Channels_______________

Assistant Professor of MedicineUniversity of California, San Francisco Medical Center

John Reppas, M.D./Ph.D.B.A., Oxford University

Entered 1995 * Defended September 22, 2000Advisor: R. Clay Reid

Saccadic Eye Movements and Early Vision_______________

Director of Public PolicyNeurotechnology Industry Organiza on

Jeremiah Scharf, M.D./Ph.D.B.S., Yale University

Entered 1995 * Defended September 8, 1999Advisor: Louis Kunkel

Compara ve Genomic Analysis of the Spinal Muscular Atrophy (SMA)Region in Mouse and Man: Applica ons for Understanding Disease

Pathogenesis_______________

Assistant Professor of NeurologyMassachuse s General Hospital

Page 47: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Alex Carter, M.D./Ph.D.B.A., Brandeis University

Entered 1996 * Defended December 17, 2001Advisor: Rosalind Segal

Neurotrophin Modula on of Synap c Plas city and CerebellarDevelopment

_______________Assistant Professor of Neurology

Washington University School of Medicine

Lisa Catapano, M.D./Ph.D.B.A., Dartmouth College

Entered 1996 * Defended September 21, 2001Advisor: Je rey Macklis

Stage Speci c Control over Neocor cal Callosal Projec on NeuronSurvival and Di eren a on

_______________Assistant Professor of PsychiatryGeorge Washington University

Thomas Chou, Ph.D.B.S., M.S., Massachuse s Ins tute of TechnologyEntered 1996 * Defended February 10, 2003

Advisor: Cli ord SaperRegula on of sleep wake ming: Circadian rhythms

and bistability of sleep wake behavior_______________

Whereabouts UnknownFormerly: Assistant Professor—Medical University of South Carolina

Bevil Conway, Ph.D.B.Sc., McGill University

Entered 1996 * Defended April 18, 2001Advisor: Margaret Livingstone

Mechanisms of Color and Mo on in Monkey Visual Cortex_______________

Associate Professor of NeuroscienceWellesley College

Page 48: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Michael Deans, Ph.D.B.A., Michigan State University

Entered 1996 * Defended April 26, 2002Advisor: David Paul

Connexin 36 Localiza on and Func on Within the Rod Pathwaysof the Mouse Re na_______________

Assistant Professor of OtolaryngologyUniversity of Utah School of Medicine

Shin Jung Kang, Ph.D.B.S., M.S., Seoul Na onal University

Entered 1996 * Defended September 25, 2001Advisor: Junying Yuan

A Dual Role of Capase 11 under Pathologicaland Physiological Condi ons

_______________Assistant Professor of Molecular Biology

Sejong University

Amy McMichaelB.S., Kalamazoo College

Entered 1996 * Le June 1, 1997_______________

Withdrew without taking degreeWhereabouts unknown

Magdalene Moran, Ph.D.B.A., Williams College

Entered 1996 * Defended October 7, 2002Advisor: David Clapham

TLF is a Func onal Regulator or Transcrip on with Targetsthat Are Dis nct from the TATA Box Binding Protein (TBP)

_______________Senior Vice President, Biology

Hydra Biosciences

Page 49: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kim Seth, Ph.D.A.B., Harvard College

Entered 1996 * Defended September 25, 2002Advisor: Joseph Majzoub

A Molecular Gene c Inves ga on of the Role of Re 1/REST In theTranscrip onal Regula on

of the Cor cotropin Releasing Hormone (CRH) Gene_______________

AdvisorResilience Therapeu cs

Abraha Taddese, M.D./Ph.D.B.A., Washington University

Entered 1996 * Defended June 6, 2001Advisor: Bruce Bean

Electrophysiology of Spontaneously Ac veTuberomammillary Nucleus Neurons

_______________Physician

Miami, Florida

Jonathan Trinidad, Ph.D.B.S., William and Mary College

Entered 1996 * Defended October 10, 2002Advisor: Jonathan Cohen

The Localiza on and Func on of Neuregulin and the ErbB Receptorsat the Neuromuscular Junc on

_______________Director, Biological Mass Spectrometry

Indiana University

Doris Tsao, Ph.D.B.S., California Ins tute of TechnologyEntered 1996 * Defended May 20, 2002

Advisor: Margaret LivingstoneStereopsis

_______________Professor of Biology

California Ins tute of Technology

Page 50: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kevin Welch, A.M.B.S., Beloit College

Entered 1996 * Le June 1, 2007_______________

Withdrew with Master’s degreeWhereabouts unknown

Alo Basu, Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1997 * Defended December 20, 2004

Advisor: Ed KravitzCharacteriza on of an Amine Modulated Stress Neuropep de System in

Homarus americanus_______________

Assistant ProfessorCollege of the Holy Cross

Adam Carter, Ph.D.B.A., Cambridge University

ESntered 1997 * Defended April 29, 2002Advisor: Wade Regehr

Synap c Transmission at Cerebellar Stellate Cells_______________

Associate Professor of Neural ScienceNew York University

Dennis Chang, Ph.D.B.Sc.H., Queen’s University

Entered 1997 * Defended July 16, 2003Advisor: Steven Reppert

Circadian Clock Molecules of a Lepidopteran Insect, Antheraea pernyi,and Comparison with the Fruit Fly, Drosophila melanogaster

_______________Principal

Clarion Healthcare

Page 51: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Mark Emerson, Ph.D.B.A., Oberlin College

Entered 1997 * Defended January 26, 2005Advisor: Davie Van Vactor

A Misexpression Gene c Screen in Drosophila Iden esCandidate Axon Guidance Factors with Di eren al Expression

_______________Assistant Professor of Biology

City College of New York

Heather Heerssen, Ph.D.B.A., Kenyon College

Entered 1997 * Defended December 3, 2003Advisor: Rosalind Segal

Mechanisms of Trk Receptor Retrograde Transport and Signaling_______________

Senior Medical WriterComplete Healthcare Communica ons

Jane Ko, Ph.D.B.A., Tulane University

Entered 1997 * Defended May 13, 2004Advisor: Li Huei Tsai

CdK5 Ac vators in Development and Poten al Regula on of Kinesins_______________

Associate Director, Global Marke ngKellogg Company

Genevieve Konopka, Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1997 * Defended May 14, 2004

Advisor: Azad BonniRegula on of STAT3 Signaling in Astrocyte Transforma on

_______________Associate Professor of Neuroscience

University of Texas Southwestern Medical Center

Page 52: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Anatol Kreitzer, Ph.D.B.S., University of California at BerkeleyEntered 1997 * Defended April 25, 2001

Advisor: Wade RegehrAc vity Dependent Modula on of Synap c Strength in the Central

Nervous System_______________

Associate Professor of NeuroscienceUniversity of California, San Francisco

Robert Moy, A.M.B.A., Oberlin College

Entered September 1997 * Withdrew June 1, 2004———————

Withdrew with a Master’s degreePrincipal

Clarion Healthcare

Lisa Simon NelsonB.S., Rutgers University

Entered 1997 * Le January 1, 2000_______________

Withdrew without taking degreeWhereabouts unknown

Christov Roberson, Ph.D.B.A., Northern Arizona University

Entered 1997 * Defended August 5, 2004Advisor: David Clapham

G Protein Regula on of Inwardly Rec fying Potassium Channels_______________

LecturerJohns Hopkins University

Page 53: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Gisela Sandoval, M.D./Ph.D.B.S., California Ins tute of TechnologyEntered 1997 * Defended July 9, 2002

Advisor: Gary RuvkunMolecular Analysis of an Interac on between the Vesicular

Acetylcholine Transporter and Synaptobrevin in Caenorhabdi s Elegans_______________

PsychiatristStanford University Hospital and Clinics

Claire Sauvageot, Ph.D.B.A., University of California, Berkeley

Entered 1997 * Defended October 9, 2003Advisor: Charles S les

Genomic Analysis of Fate Choice in Cor cal Progenitor Cells_______________

Senior Manager, Scien c Strategy and Publica onsTakeda Oncology (Dana Farber Cancer Ins tute)

Sohail Tavazoie, M.D./Ph.D.B.S., University of California, BerkeleyEntered 1997 * Defended June 15, 2001

Advisor: R. Clay ReidThe Development of Neural Connec ons from Re na to Thalamus:

Matura on and Plas city of Recep ve Fields_______________

Associate Professor of Cancer BiologyThe Rockefeller University

Sarah Webster, Ph.D.B.S., University of Connec cut

Entered 1997 * Defended December 12, 2003Advisor: Gary Yellen

Structures and mechanisms of ga ng inShaker voltage dependent potassium channels

_______________Assistant Professor

College of the Holy Cross

Page 54: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ying Zhang, Ph.D.B.S., University of Science and Technology (China)

M.S., Duke UniversityEntered 1997 * Defended May 5, 2003

Advisor: R. Clay ReidWhere Is It? Deriving S mulus Loca on from the Responses of Visual

Neurons_______________

Whereabouts unknown

Bruce Altevogt, Ph.D.B.A., University of Virginia

Entered 1998 * Defended November 20, 2003Advisor: David Paul

Analysis of Connexins in Myela on_______________

Director of Science Policy and Science AdvocacyP zer, Inc.

Teresa Chae, M.D./Ph.D.A.B., Harvard College

Entered 1998 * Defended May 15, 2002Advisor: Christopher A. Walsh

Mechanisms of Dorsal Pa erning in the Developing Forebrain_______________OphthalmologistHouston, Texas

Michael Tri Do, Ph.D.B.S., Georgetown University

Entered 1998 * Defended December 15, 2003Advisor: Bruce Bean

Genera on and Regula on of Pacemaking in Subthalamic Neurons_______________

Assistant Professor of NeurologyHarvard Medical School

Page 55: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

John Elfar, M.D.Johns Hopkins University

Entered 1998 * Withdrew September 1, 2000

_______________

Withdrew without taking a Ph.D.Returned to Harvard Medical School to complete his M.D.

Associate Professor of OrthopaedicsUniversity of Rochester Medical Center

Edmund Gri n, Jr., M.D./Ph.D.B.S., Columbia University

Entered 1998 * Defended March 7, 2002Advisor: Chuck Weitz

Light, Cryptochrome, and Circadian Clocks_______________

Assistant Professor of Clinical PsychiatryColumbia University Medical Center

W. Taylor Kimberly, M.D./Ph.D.B.A., Princeton University

Entered 1998 * Defended May 10, 2002Advisor: Dennis Selkoe

Intramembrane Proteolysis and Signal Transduc on:The Gamma Secretase Complex and Its Connec on to Alzheimer’s

Disease_______________

Associate Director, Neuroscience Intensive Care UnitMassachuse s General Hospital

Gaby Maimon, Ph.D.B.S., Cornell University

Entered 1998 * Defended May 5, 2005Advisor: John Assad

Cortex, vision and ac on._______________

Assistant ProfessorThe Rockefeller University

Page 56: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Carolyn Rodriguez, M.D./Ph.D.A.B., Harvard College

Entered 1998 * Defended March 4, 2002Advisor: Susan Dymecki

Innova ng Gene c Technology to Map the Origin of the PrecerebellarSystem

_______________Assistant Professor of Psychiatry

Columbia University Medical Center

Erin Perry, A.M.B.A., Amherst College

Entered 1998 * Withdrew November 1, 2002_______________

Withdrew with Master’s degreeEnrolled in Harvard School of Educa on

Whereabouts unknown

Jun Shen, Ph.D.B.A., Dartmouth University

Entered 1998 * Defended December 13, 2004Advisor: Christopher A. Walsh

Using Forward and Reverse Gene cs to Study Genes that Control theSize of the Brain_______________

Instructor in PathologyHarvard Medical School

Susan St. Pierre, Ph.D.B.S., Stanford University

Entered 1998 * Defended March 2, 2004Advisor: Stefan Thor

Neuronal Speci ca on in Drosophila: The roles of rotund and squeeze,two related zinc nger genes

_______________Homemaker

Page 57: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Brian Whalen, Ph.D.B.S., Columbia University

Entered 1998 * Defended April 12, 2005Advisor: Dennis Selkoe

Early Events in Alzheimer’s Disease: An Examina on of Amyloid BetaProto bril Biology_______________

Vice President, Science and MedicineEvoke Health

Fu Chia Yang, Ph.D.A.B., Harvard College

Entered 1998 * Defended October 8, 2003Advisor: Chuck Weitz

The Poten al Role for the EGF Receptor System in the CircadianRegula on of Locomotor Ac vity

_______________Scien st I

Dana Farber Cancer Ins tute

Paul Arday o, Ph.D.Wayne State University

Entered 1999 * Defended May 4, 2006Advisor: Kwang Soo Kim

Behavioral Inves ga on of Cogni ve Func on in Mouse Modelsof Neurological and Psychiatric Disorders

_______________

Clinical Research Scien stEli Lilly and Company

Nathaniel Blair, Ph.D.B.A., University of Chicago

Entered 1999 * Defended January 15, 2004Advisor: Bruce Bean

Contribu on of Voltage Gated Sodium, Calcium and Potassium Currentsto the Excitability of Nocicep ve Sensory Neurons

_______________Scien st II

Hydra Biosciences

Page 58: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Laurie Burlingame, A.M.B.A., Wellesley College

Entered 1999 * Withdrew June 1, 2002_______________

Withdrew with Master’s DegreeAssociate A orney

Goodwin Procter LLC

Yoojin Choi, Ph.D.University of California, Irvine

Entered 1999 * Defended July 19, 2006Advisor: Rosalind Segal

Mitogenic Niches in the Developing Cerebellum_______________

Associate Professor of BiologyNorth Park University

Thomas Deuel, M.D./Ph.D.A.B., Princeton University

Entered 1999 * Defended August 4, 2004Advisor: Christopher A. Walsh

Doublecor n and the Doublecor n like Kinase in CNS Development andPlas city

_______________Neurologist and A liate Professor at Center for Arts/Experimental Media

University of Washington

Shelby Dietz, Ph.D.B.A., Cornell University

Entered 1999 * Defended November 29, 2005Advisor: Venkatesh Murthy

Plas city and Development of the Mitral Granule Reciprocal Synapsein the Mammalian Olfactory Bulb

_______________Research AssociateCornell University

Page 59: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Eleanore Edson, Ph.D.B.S., Stanford University

Entered 1999 * Defended August 3, 2005Advisor: Chinfei Chen

Entered September 1999Contribu ons of Glutamate Transporters and Metabotrophic Glutamate

Receptors to Synap c Transmission over Development at theRe nogeniculate Synapse

_______________Program O cer

Bill and Melinda Gates Founda on

Kelly Foster, Ph.D.B.S., University of Pi sburgh

Entered 1999 * Defended August 11, 2004Advisor: Wade Regehr

Mechanisms and Func onal Consequences of Depression at the ClimbingFiber to Purkinje Cell Synapse

_______________Associate Director of Biology

Ataxion Therapeu cs

Sanjiv Harpavat, M.D./Ph.D.B.S., Stanford University

Entered 1999 * Defended October 20, 2004Advisor: Connie Cepko

Roles of Thyroid Hormone in Chick Re nal Development_______________

Physician and researcherBaylor College of Medicine

Maria Leh nen, Ph.D.B.A., University of Pennsylvania

Entered 1999 * Defended December 15, 2005Advisor: Azad Bonni

Survival and Death Signaling Pathways: From Single Cells to Organisms_______________

Assistant Professor of PathologyHarvard Medical School

Page 60: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jacob Marcus, Ph.D.B.S., Massachuse s Ins tute of TechnologyEntered 1999 * Defended April 28, 2006

Advisor: Joel ElmquistOrexin Receptors and the Central Autonomic System

_______________Principal Scien st, Neuroscience Discovery

Merck Research Laboratories

Bence Ölveczky, Ph.D.M.Sc., Technical University of Budapest

M.Sc., Imperial UniversityEntered 1999 * Defended May 23, 2003

Advisor: Markus MeisterRe nal Processing of Natural Mo on

_______________Professor of Organismic and Evolu onary Biology

Harvard University

Hien Tran, M.D./Ph.D.Stanford University

Entered 1999 * Defended June 5, 2003Advisor: Michael Greenberg

A Role for Forkheads in the Response to Stress_______________

DermatologistSan Diego, California

Duo Wang, Ph.D.B.S., Beijing University

Entered 1999 * Defended May 2, 2005Advisor: Gary Ruvkun

Nega ve regula on of RNA interference in Caenorhabdi s elegans_______________

Senior Manager, Market Intelligence and Market DevelopmentMedtronic

Page 61: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Amy Ashbacher, A.M.B.S., Massachuse s Ins tute of TechnologyEntered September 2000 * Le June 1, 2005

Advisor: John Assad_______________

Withdrew with Master’s degreeWhereabouts unknown

Jay Biko , Ph.D.Sc.B., Brown University

Entered 2000 * Defended May 7, 2007Advisor: Michael Greenberg

Molecular Mechanisms Regula ng Cor cal Development: FromNeurons to Synapses_______________

Postdoctoral FellowColumbia University — Jessell Lab

Saskia de Vries, Ph.D.B.S., Yale University

Entered 2000 * Defended May 5, 2006Advisor: Markus Meister

Modula on of Re nal Responses by Single Amacrine Cells_______________

Scien st IIAllen Ins tute for Brain Science

Antoniu Fantana, Ph.D.Diploma, Heidelberg University

Entered 2000 * Defended May 1, 2006Advisor: Markus Meister

Odor Processing in the Olfactory Bulb: Structureof Mitral Cell Recep ve Fields

_______________Medical Liaison

Eli Lilly

Page 62: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Carson Alex Goddard, Ph.D.B.A., Oberlin College

Entered 2000 * Defended May 15, 2006Advisor: Carla Shatz

MHC Class I and the Homeosta c Regula on of CNS Synapses_______________

Inves gator IINovar s Ins tutes

Joshua Gooley, Ph.D.B.A., Wesleyan University

Entered 2000 * Defended October 31, 2005Advisor: Cli ord Saper

Entrainment of the Circadian Timing System by Light and Food_______________

Assistant Professor, Neuroscience and Behavioral Disorders ProgramDuke NUS Medical School

Anna Greka, M.D./Ph.D.A.B., Harvard College

Entered 2000 * Defended October 10, 2002Advisor: David Clapham

TRP Channels and the Cytoskeleton: Implica ons for Neuronal Growthand Cellular Mo lity_______________

Assistant Professor of MedicineHarvard Medical School

Kenichi Hartman, Ph.D.B.S., Brandeis University

Entered 2000 * Defended April 28, 2006Advisor: Venkatesh Murthy

Regula on of Neuronal Morphology and Synap c Developmentby Ac vity

_______________U.S. Patent Agent/Israeli Patent A orney Trainee

Allan C. En s, Ph.D. Intellectual Property Ltd.

Page 63: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ma hew Hass, Ph.D.B.S., University of Iowa

Entered 2000 * Defended November 16, 2006Advisor: Bruce Yankner

Iden ca on of Novel Roles of the Amyloid Precursor Protein in SignalTransduc on

_______________Research Instructor

Cincinna Children’s Hospital Medical Center

Rhonda Hyde, Ph.D.B.S., Yale University

Entered 2000 * Defended September 8, 2008Advisor: Anne Hart

Gene c Analysis of Protein Kinase C and MAPK in C. elegansmechanosensa on_______________Biology Instructor

Biology Instructor, Upli Educa on

Alex Jackson, Ph.D.B.Sc., McGill University

Entered 2000 * Defended September 11, 2006Advisor: Bruce Bean

Subthreshold Currents Underlying the Spontaneous Firing ofHypothalamic Pacemaker Neurons

_______________Assistant Professor of Physiology

University of Connec cut

Vuk Koprivica, Ph.D.B.A., Earlham University

Entered 2000 * Defended November 22, 2005Advisor: Zhigang He

Mechanisms of Myelin Regenera on Inhibi on_______________

Director of Business DevelopmentVanda Pharmaceu cals

Page 64: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Sebas an Kraves, Ph.D.University of Buenos Aires

Entered 2000 * Defended May 5, 2005Advisor: Chuck Weitz

Molecular Analyses of the Circadian Control of Locomotor Ac vityin Mammals

_______________Co Founder

miniPCR

Claire McKellar, Ph.D.B.S., Yale University

Entered 2000 * Defended November 17, 2005Advisor: Carla Shatz

A Program of New Gene Expression Induced by Synaptogenesis_______________Research Specialist

Howard Hughes Medical Ins tute

Brooke Pa en, Ph.D.B.S., Princeton University

Entered 2000 * Defended December 8, 2005Advisor: Gabriel Corfas

Molecular Mechanisms of Radial Glia Forma on in the DevelopingCerebellum

_______________Assistant Vice President

MedStar Washington Hospital Center

Ben Samuels, Ph.D.B.S., Johns Hopkins University

Entered 2000 * Defended May 2, 2006Advisor: Li Huei Tsai

Physiological Func ons of Cdk5 in the Nervous System_______________

Assistant Professor of PsychiatryColumbia University

Page 65: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Sujith Vijayan, PH.D.A.B., Amherst College

Entered 2000 * Defended September 17, 2007Advisor: Emery Brown

Characteriza on of Ac vity in the Primary Somatosensory Cortex DuringAc ve Behavior and Sleep

_______________Postdoctoral Fellow

Massachuse s General Hospital—Cash Lab

Kevin Wang, Ph.D.B.S., M.S., Stanford UniversityM.Phil., Cambridge University

M.D., University of California at San FranciscoEntered 2000 * Defended March 19, 2003

Advisor: Zhigang HeSignaling Mechanisms Underlying Inhibi on

of Axonal Regenera on by CNS Myelin_______________

Assistant Professor of DermatologyStanford University School of Medicine

Lisa Zakhary, M.D./Ph.D.B.S., Columbia University

Entered 2000 * Defended April 25, 2005Advisor: Catherine Dulac

Anatomical and Func onal Analysis of MicroRNAs in MammalianOlfactory Neurogenesis

_______________Psychiatrist

Massachuse s General Hospital

Page 66: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Perry Bri s, Ph.D. B.A., Colgate College

M.D., Harvard Medical SchoolEntered 2001 * Defended January 29, 2004

Advisor: John FlanaganRNA Based Mechanisms for Locally Regula ng Axon Guidance Decisions

_______________Founder and Chief Execu ve O cer

SliceLabs, LLC

Brenda Bloodgood, Ph.D. University of California, San Diego

Entered 2001 * Defended July 14, 2006Advisor: Bernardo Saba ni

Biochemical Compartmentaliza on and Synap c Signaling in Dendri c Spines of Hippocampal Pyramidal Neurons

_______________Assistant Professor of Neurobiology University of California, San Diego

James Berg, Ph.D. B.S., Rutgers University

Entered 2001 * Defended May 5, 2008Advisor: Gary Yellen

Imaging Neuronal Metabolism_______________

Manager, Electrophysiology The Allen Ins tute for Brain Science

Dinu Florin Albeanu, Ph.D. B.S., Massachuse s Ins tute of TechnologyEntered 2001 * Defended September 8, 2008

Advisor: Venkatesh MurthyMul modal Imaging of Sensory Inputs and Neuronal Coding in the

Rodent Olfactory Bulb _______________

Assistant Professor of Neuroscience Cold Spring Harbor Laboratory

Page 67: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Eunju Chung, Ph.D. B.S., Boston College

Entered 2001 * Defended May 9, 2007Advisor: Tom Schwarz

Axonal Transport of Synap c Components and Synaptogenesis in Drosophila

_______________Senior Manager, Strategic Planning

Genexine, Inc.

Yeang Ch’ng, M.D./Ph.D. B.S., M.S., Yale University

Entered 2001 * Defended September 17, 2007Advisor: Clay Reid

Func onal Imaging of Cellular and Vascular Ac vity in Visual Cortex _______________

Radiologist Fine Art Photographer

Jay Chyung, M.D./Ph.D. A.B., Harvard College

Entered 2001 * Defended September 1, 2004Advisor: Dennis Selkoe

Cell Surface Events in the Amyloidogenic Processing of the Amyloid Precursor Protein

_______________Advisor

Recon Strategy

Sophie Currier, M.D./Ph.D. B.S., Massachuse s Ins tute of TechnologyEntered 2001 * Defended June 7, 2004

Advisor: Christopher A. WalshGene cs of Walker Warburg Syndrome: A Disorder of Brain, Muscle,

and Eye Development _______________

Physician London, England

Page 68: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

John Dekker, M.D./Ph.D. B.A., Wesleyan University

Entered 2001 * Defended May 2, 2006Advisor: Gary Yellen

Molecular Analysis of Ga ng in Voltage Dependent Ion Channels _______________

Sta physician NIH Clinical Center

Elizabeth Glater, Ph.D. B.A., Swarthmore College

Entered 2001 * Defended May 24, 2006Advisor: Thomas Schwarz

The mechanism for Transport of Mitochondria in Drosophila _______________

Assistant Professor of Neuroscience Pomona College

Robert Gri n, M.D./Ph.D. B.A., Swarthmore College

Entered 2001 * Defended September 20, 2005Advisor: Cli ord Woolf

Di eren al Gene Expression Associated with Neuropathic Pain _______________Anesthesiologist

New York City

Dana Harrar, Ph.D. B.A., Johns Hopkins UniversityM.D., Vanderbilt University

Entered 2001 * Defended February 28, 2007Advisor: Michael Greenberg

An RNA Interference Screen Iden es New Molecules Required for Mammalian Synapse Development

_______________Resident in Child Neurology Boston Children’s Hospital

Page 69: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Mac Hooks, Ph.D. A.B., Harvard College

Entered 2001 * Defended March 26, 2007Advisor: Gina Turrigiano

Func onal Development of the Re nogeniculate Synapse _______________

Assistant Professor of Neurobiology University of Pi sburgh

Vikram Khurana, Ph.D. B.A., B.S., M.D., University of Sydney

Entered 2001 * Defended May 11, 2006Advisor: Mel Feany

Mitogenic signaling and cell cycle ac va on mediate Neurodegenera on in Drosophila

_______________Neurologist

Massachuse s General Hospital

Misao Higashi Kovtun, Ph.D. Sc..B., Brown University

Entered 2001 * Defended April 13, 2009Advisor: Thomas Schwarz

Characteriza on of an Impor n beta11 Mutant at the Drosophila Neuromuscular Junc on

_______________Homemaker and Philanthropist

Brad Molyneaux, M.D./Ph.D. B.A., Dartmouth University

Entered 2001 * Defended September 23, 2005Advisor: Je rey Macklis

Molecular Development of Cor cospinal Motor Neurons _______________

Assistant Professor of Neurobiology University of Pi sburgh Medical Center

Page 70: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Patrick Safo, M.D./Ph.D. B.S., State University of New York at Stony BrookEntered 2001 * Defended September 23, 2005

Advisor: Wade RegehrMolecular Development of Cor cospinal Motor Neurons

_______________Dermatologist

Wisconsin

Gilberto Solver Llavina, Ph.D. B.S., University of Puerto Rico

Entered 2001 * Defended August 8, 2007Advisor: Bernardo Saba ni

Compartmentaliza on and Modula on of Synap c Signaling in Dendrites and Spines

_______________Inves gator

Novar s, Inc.

Rebecca Swimmer, A.M. University of Illinois, Urbana

Entered September 2001 * Withdrew June 1, 2005_______________

Withdrew with Master’s degree Veterinarian in Venice, California

Cindy Chen, A.M. B.S., Massachuse s Ins tute of TechnologyEntered 2002 * Withdrew June 1, 2008

_______________Withdrew with Master’s Degree

Whereabouts unknown

Page 71: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kenneth Colodner, Ph.D. B.A., Northwestern University

Entered 2002 * Defended June 2, 2008Advisor: Mel Feany

Using Drosophila to Inves gate the Glial Contribu on to Neurodegenera ve Disorders

_______________Visi ng Assistant Professor

Mt. Holyoke College

Steven Flavell, Ph.D. B.A., Oberlin College

Entered 2002 * Defended October 17, 2008Advisor: Michael Greenberg

Regula on of synapse development by the ac vity regulated transcrip on factor MEF2

_______________Assistant Professor

Massachuse s Ins tute of Technology

Raj Gopal, M.D./Ph.D. B.S., University of California, Irvine

Entered 2002 * Defended June 26, 2007Advisor: Bruce Yankner

Regula on of DNA Double Strand Break Repair by Parkin _______________

Clinical Fellow in Medicine Brigham and Women’s Hospital

Brian Ha er, M.D./Ph.D. B.S., Princeton University

Entered 2002 * Defended November 7, 2006Advisor: David Rowitch

Transcrip onal Regula on of Olig2 in Motor Neuron Development _______________Ophthalmologist

Boston, Massachuse s

Page 72: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Todd Herrington, M.D./Ph.D. B.S., Stanford University

Entered 2002 * Defended June 15, 2007Advisor: John Assad

A en onal Dynamics in Parietal Cortex _______________

Instructor in Neurology Harvard Medical School

Elizabeth Hong, Ph.D. B.S., California Ins tute of Technology

Entered 2002 * Defended December 1, 2008Advisor: Michael Greenberg

Func onal Signi cance of Neuronal Ac vity Dependent Transcrip onal Regula on in the Nervous System

_______________Assistant Professor of Neuroscience

California Ins tute of Technology

Michael Hughes, Ph.D. B.S., Stanford University

Entered 2002 * Defended December 19, 2006Advisor: Dietmar Schmucker

A Gene c and Biochemical Analysis of Dscam Signaling in Dendrite Morphogenesis

_______________Assistant Professor of Biology

University of Missouri at St. Louis

John Nicholas Hunter, Ph.D. B.S., University of Washington

Entered 2002 * Defended June 16, 2009Advisor: Richard T. Born

Spa al Integra on of Mo on Signals in Macaque MT _______________

Whereabouts unknown

Page 73: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Eirene Kontopoulos, Ph.D. B.A., Haverford College

Entered 2002 * Defended May 4, 2007Advisor: Mel Feany

Mechanisms of alpha synuclein Neurotoxicity in Parkinson’s Disease _______________

Biotech Analyst Fidelity Investments

Edmund Koundakjian, Ph.D. B.S., University of California, Riverside

M.S., California State University, San BernardinoEntered 2002 * Defended May 15, 2008

Advisor: Lisa GoodrichAn Analysis of the Development of the Cochlear Ves bular Ganglion

_______________Patent Agent

Cooley LLP

Alice Lu, Ph.D. B.A., Columbia University

Entered 2002 * Defended August 11, 2008Advisor: Rudolph Tanzi

Poten al Roles for Ubiquilin 1 in Alzheimer’s Disease Pathogenesis _______________

Healthcare Strategy Consultant Har ord, CT

Ju Lu, Ph.D. B.Engr., Tsinghua University

Entered 2002 * Defended April 21, 2008Advisor: Je Lichtman

Connectomic Studies of the Mammalian Neuromuscular System _______________

Assistant Project Scien st University of California, Santa Cruz (Beijing, China)

Page 74: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Laura Magno , Ph.D. B.A., University of Pennsylvania

Entered 2002 * Defended May 27, 2010Advisor: David Paul

Gap Junc ons Between Astrocytes and Oligodendrocytes Are Cri cal for Normal Glial Func on

_______________ Preceptor in Molecular and Cellular Biology, Harvard University

Senior Editor, American Journal Experts

David Simon, Ph.D. B.S., M.S., Brandeis University

Entered 2002 * Defended April 30, 2008Advisor: Josh Kaplan

microRNA Regula on of Synapse Func on in C. elegans _______________

Postdoctoral Fellow The Rockefeller University—Tessier Lavigne Lab

Glenn Yiu, M.D./Ph.D. B.S., Columbia University

Entered 2002 * Defended May 26, 2006Advisor: Zhigang He

Signaling Mechanisms Limi ng Axon Regenera on in the Adult Central Nervous System

_______________Assistant Professor of Ophthalmology

University of California, Davis

Victoria Abraira, Ph.D. B.S., University of Southern California

Entered 2003 * Defended March 30, 2009Advisor: Lisa Goodrich

Molecular Mechanisms of Inner Ear Morphogenesis _______________

Postdoctoral Fellow Harvard Medical School —Ginty Lab

Page 75: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Joshua Jacob Buchman, Ph.D. B.A., Rutgers University

Entered 2003 * Defended October 1, 2009Advisor: Li Huei Tsai

Roles of centrosome proteins associated with primary microcephaly in mammalian cor cogenesis

_______________Science Advisor

Goodwin Procter LLP

Adam Christensen, Ph.D. B.S., University of Iowa

Entered 2003 * Defended March 30, 2009Advisor: David Corey

The Role of TRP Channels in Mechanosensa on _______________

Special Agent Federal Bureau of Inves ga on

Stephanie Courchesne Schlink, Ph.D. B.S., Davidson College

Entered 2003 * Defended November 3, 2009Advisor: Rosalind Segal

Spa al Aspects of Neuronal Survival Signaling in Sensory Neurons: A Role for Mef2D and Bcl w

_______________Health Sciences Policy Analyst Na onal Ins tutes of Health

Luis de la Torre Ubieta, Ph.D. B.S., University of Puerto Rico

Entered 2003 * Defended May 4, 2010Advisor: Azad Bonni

Transcrip onal Regula on of Neuronal Polarity _______________

Postdoctoral Fellow University of California at Los Angeles—Geschwind Lab

Page 76: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Xuecai Ge, Ph.D. B.S., Beijing University

M.S., Tsinghua UniversityEntered 2003 * Defended April 8, 2009

Advisor: Li Huei TsaiFunc ons of the Centrosome in Embryonic Neurogenesis

_______________Postdoctoral Fellow

Stanford University Medical Center—Sco Lab

Johannes Haushofer, Ph.D. B.A., Oxford University

Entered 2003 * Defended March 26, 2008Advisor: Margaret Livingstone

Cor cal Object Representa ons _______________

Assistant Professor of Psychology Princeton University

Ma hew Hemming, Ph.D. B.S., University of California, San Diego

M.D., Harvard Medical SchoolEntered 2003 * Defended September 10, 2007

Advisor: Dennis SelkoeRegula on of Beta Amyloid Protein Levels by Proteoly c Degrada on

and Its Implica ons for Alzheimer’s Disease _______________

Clinical Fellow in Oncology Dana Farber Cancer Ins tute—Bradner Lab

Youn Young Kate Hong, Ph.D. B.S., Brown University

Entered 2003 * Defended April 21, 2010Advisor: Joshua Sanes

Structure and Development of Re nal Ganglion Cells _______________

Postdoctoral Fellow Columbia University—Bruno Lab

Page 77: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Aaron Kerlin, Ph.D. B.A., Oberlin College

Enterd 2003 * Defended January 18, 2011Advisor: R. Clay Reid

Response proper es of inhibitory neuron subtypes in mouse visual cortex _______________

Postdoctoral Fellow Janelia Farm—Svoboda Lab

Allison Terra Knoll, Ph.D. B.S., University of Wisconsin at Madison

Entered 2003 * Defended November 9, 2009Advisor: Bill Carlezon

Role of Kappa Opioid Receptors in Fear and Anxiety like Behavior in Rats

_______________Postdoctoral Fellow

University of Southern California—Zilkha Lab

Camilo Libedinsky, Ph.D. B.S., University of Chile

Entered 2003 * Defended May 6, 2009Advisor: Marge Livingstone

Neuronal Mechanisms of Visual Percep on: Role of Early Visual and Prefrontal Cor cal Areas in Conscious Awareness

_______________Assistant Professor of Psychology Na onal University of Singapore

Timothy Mosca, Ph.D. B.S., M.S., Yale University

Entered 2003 * Defended April 23, 2009Advisor: Thomas Schwarz

Nuclear Import and Synapse to Nucleus Signalling at the Drosophila Neuromuscular Junc on

_______________Postdoctoral Fellow

Stanford University—Luo Lab

Page 78: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Carlos Ponce, M.D./Ph.D. B.S., University of Utah

Entered 2003 * Defended May 22, 2008Advisor: Rick Born

Integra ng Visual Informa on through Parallel Pathways _______________

Postdoctoral Fellow Harvard Medical School — Livingstone Lab

Pavan Ramdya, Ph.D. B.A., Drew University

Entered 2003 * Defended May 1, 2009Advisor: Florian Engert

Direc on selec vity in the larval zebra sh op c tectum _______________

SNSF Advanced Postdoctoral Fellow California Ins tute of Technology—Dickinson Lab

James Fitzhugh Sturgill, Ph.D. B.S., Davidson College

Entered 2003 * Defended September 30, 2009Advisor: Bernardo Saba ni

Ac vity dependent Regula on of Synapse Structure and Func on: Roles of PSD 95 and the Metabolic Sensor, AMPK

_______________Postdoctoral Fellow

Cold Spring Harbor Laboratory—Kepecs Lab

Geo rey Tanner, Ph.D. B.S., M.A.L.S., Wesleyan University

Entered 2002 * Defended July 29, 2009Advisor: Gary Yellen

K(ATP) Channels, Metabolism, and Neuronal Excitability _______________Visi ng Lecturer

Mt. Holyoke College

Page 79: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Max Tisch eld, Ph.D. B.A., Rutgers University

Entered 2003 * Defended April 23, 2009Advisor: Elizabeth Engle

Human Eye Movement Disorders Help Iden fy Molecular and Gene c E ologies

of Novel Neurological Syndromes _______________

Postdoctoral Fellow Boston Children’s Hospital—Engle Lab

Rochelle Wi , M.D./Ph.D. B.S., University of Illinois, Urbana

Entered 2003 * Defended June 22, 2010Advisor: Rosalind Segal

Of Structure and Func on: The Nature of the Sonic Hedgehog Heparan Sulfate Proteoglycan Interac on

_______________Child Neurologist

New York City

Eiman Abdel Azim, Ph.D. B.S., B.A., Stanford University

Entered 2004 * Defended November 20, 2009Advisor: Je rey Macklis

Molecular Controls over Neocor cal Neuronal Diversity and Oligodendrocyte Development

_______________Postdoctoral Fellow

Columbia University Medical Center—Jessel Lab

Jessica Appler, Ph.D. B.S., University of Southern CaliforniaEntered 2004 * Defended April 22, 2010

Advisor: Lisa GoodrichThe Molecular and Morphological Development

of Spiral Ganglion Neurons _______________

Public Health Analyst BARDA/U.S. Department of Health and Human Services

Page 80: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Parizad Bilimoria, Ph.D. B.S., M.S., Brandeis UniversityEntered 2004 * Defended May

Advisor: Azad BonniSignaling Mechanisms Regula ng Neuronal Morphogenesis

in the Cerebellar Cortex _______________

Director of Programming and Outreach Harvard University—Conte CBS Colloquium on Mental Health

Dravida Das Bock, Ph.D. B.S., Brown University

Entered 2004 * Defended January 10, 2011Advisor: R. Clay Reid

Network Anatomy and In Vivo Physiology from a Group of Visual Cor cal Neurons

_______________Group Leader Janelia Farm

Kelly Dakin, Ph.D. B.A., Bowdoin College

Entered 2004 * Defended September 6, 2011Advisor: Bruce Yankner

Novel Biological Func ons of Hemoglobin in the Brain

_______________Database Curator

Alzforum

Nathan Gouwens, Ph.D. B.A., Northwestern University

Entered 2004 * Defended July 15, 2009Advisor: Rachel Wilson

Processing of Neural Signals in the Drosophila Olfactory System _______________

Scien st I The Allen Ins tute for Brain Science

Page 81: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Peri Kurshan, Ph.D. B.Sc., Brown University

Entered 2004 * Defended March 4, 2010Advisor: Thomas Schwarz

The Role of Alpha2delta 3 in Calcium Channel Localiza on, Synap c Func on and Bouton Forma on

_______________Postdoctoral Fellow

Stanford University—Shen Lab

Jason Neal, Ph.D. B.S., Hampton University

M.D., Columbia University Medical Center (2014)Entered 2004 * Defended August 3, 2010

Advisor: Christopher A. WalshInsights into the role of Filamin A and Arfgef2 genes

in periventricular heteropia forma on _______________

Resident in Neurology Columbia University Medical Center

Shawn R. Olsen, Ph.D. B.S., University of Utah

Entered 2004 * Defended August 11, 2008Advisor: Rachel Wilson

Synap c and Circuit Mechanisms of Odor Processing in Drosophila _______________

Assistant Inves gator The Allen Ins tute for Brain Science

Irit Rappley, Ph.D. B.S., Massachuse s Ins tute of TechnologyEntered 2004 * Defended May 1, 2009

Advisor: Dennis SelkoeSelec ve E ects of Alpha Synuclein

on Membrane Phospholipids and Mitochondrial Func on _______________

Scien st II Celgene

Page 82: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Leigh Amanda Sadacca, Ph.D. B.A., University of Pennsylvania

Entered 2004 * Defended July 15, 2010Advisor: Charles Weitz

Circadian Clocks in Behavior and Physiology _______________

Laboratory Manager for Director of Division of Biosciences University College London

Ganesh Shankar, M.D./Ph.D. B.A., Stanford University

Entered 2004 * Defended December 17, 2007Advisor: Dennis Selkoe, Bernardo Saba ni

Soluble Abeta Oligomers and Synap c Dysfunc on in Alzheimer’s Disease

_______________Medical Resident in Neurosurgery Massachuse s General Hospital

Yue Yang, Ph.D. B.S., Stanford University

Entered 2004 * Defended May 3, 2010Advisor: Azad Bonni

A Cdc20 APC Ubiqui n Signaling Pathway Regulates Presynap c Di eren a on

_______________Postdoctoral Fellow

Washington University in St. Louis —Bonni Lab

Kevin Michael Cury, Ph.D. B.S., University of California at BerkeleyEntered 2005 * Defended April 29, 2011

Advisor: Naoshige UchidaThe Role of Spike Timing in Odor Coding During Sni ng

_______________Postdoctoral Fellow

Columbia University—Axel Lab

Page 83: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Danos Christodoulou B.S., University of Wisconsin at MadisonEntered 2005 * Withdrew May 11,2006

_______________Withdrew without taking degree

Transferred to BBS, defended 2013 Postdoctoral Fellow

Harvard Medical School—Seidman Lab

Jamie Fitzgerald, Ph.D. B.S., Stanford University

Entered 2005 * Defended August 23, 2011

Advisor: John Assad

Generalized associa ve representa ons in parietal cortex

_______________

Postdoctoral Fellow

The Rockefeller University—the Maimon Lab

Masha Gelfand, Ph.D. B.S., Washington University

Entered 2005 * Defended April 13, 2012Advisor: Chenghua Gu

Characteriza on of the in vivo role of Neuropilin1 during development _______________

Scien c Editor Cell Press

Andrew Jacob Giessel B.S., University of Kansas

Entered 2005 * Defended August 19, 2010Advisor: Bernardo Saba ni

Local, Non linear Regula on of Synap c Signals at Dendri c Spines of CA1 Pyramidal Cells

_______________

Data Scien st SenseAI

Page 84: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

J. Stefan Kaczmarek, Ph.D. B.A., Williams College

Entered 2005 * Defended August 2, 2011Advisor: David ClaphamEntered September 2005

Novel ac va on mechanisms of TRPC5 reveal its par cipa on in neuronal growth cone collapse

_______________Sta Scien st II

Genzyme

Nadia Li erman, Ph.D. A.B., Princeton University

Entered 2005 * Defended April 12, 2011Advisor: Azad Bonni

Regula on of Golgi Morphogenesis and Dendrite Pa erning by Ubiqui n Signaling _______________

Scien st and Project Manager 23andMe

Luis Mejia, Ph.D. B.S., University of Illinois – Urbana

Entered 2005 * Defended June 6, 2013Advisor: Azad Bonni

Interac on proteomics iden es a novel Hap1 Tsc1 connec on thatcontrols Neuronal mTORC1 signaling and pyramidal neuron

morphogenesis_______________

Postsdoctoral Fellow Cold Spring Harbor Laboratory — Bo Lab

Ellio Merriam B.S., University of Wisconsin at MadisonEntered 2005 * Withdrew May 11,2006

_______________Withdrew without taking degree

Enrolled in the University of Wisconsin Ph.D. program in neuroscience Patent Agent, Sea le, Washington

Page 85: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Michael Hideki Myoga, Ph.D. B.S., University of Washington

Entered 2005 * Defended February 25, 2011Advisor: Wade Regehr

Calcium Dependent Synap c Plas city in the Cerebellum _______________

Group Leader Ludwig Maximilians Universtat—Munich

Allison Paradise, A.M. B.Sc., Brown University

Entered 2005 * Le November 1, 2009_______________

Withdrew with Master’s degree Execu ve Director, My Green Lab

Simina Popa B.S., University of Washington

Entered 2005 * Le July 19, 2007_______________

Withdrew without taking degree Earned Ph.D. from University of Washington

Lecturer, University of Washington

Zeev Waks B.S., Technion Israel Ins tute of TechnologyEntered 2005 * Withdrew November 27, 2006

_______________Transferred to Systems Biology

Research Scien st, IBM

Page 86: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Yi (Joey) Zhou, Ph.D. B.S., Duke University

Entered 2005 * Defended December 16, 2011Advisor: Rachel Wilson

Transduc on in Drosophila olfactory receptor neurons is invariant to air speed

_______________

Quan ta ve Researcher TrexQuant Management

Page 87: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Rachel Bortnick, M.D./Ph.D.B.A., University of Chicago

M.Phil, Cambridge UniversityEntered 2006 * Defended October 21, 2009

Advisor: Dietmar SchmuckerGene c Dissec on of the Dscam Pathway

_______________Pediatrician

University Hospital—Freiberg

Daniel Brady, Ph.D.B.S., University of California at Berkeley

Entered 2006 * Defended October 28, 2011Advisor: Takao Hensch

Mechanisms of cross modal re nement by visual experience_______________Lead Data Scien st

Hello Alfred

Bre Carter, Ph.D.B.S., George Ins tute of Technology

Entered 2005 * Defended July 27, 2011Advisor: Bruce Bean

Sodium current during and between ac on poten als of GABAergicand glutamatergic neurons

_______________Postdoctoral Fellow

Oregon Health & Science University—Jahr Lab

Eun Young Choi, Ph.D.B.A., Columbia University

Entered 2006 * Defended July 29, 2013Advisor: Randy Buckner

The Organiza on of Cor costriatal Connec vity in the Human Brain_______________

Postdoctoral FellowUniversity of Rochester—Haber Lab

Page 88: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Irina De la Huerta, Ph.D.A.B., Harvard College

M.D., University of TorontoEntered 2006 * Defended April 19, 2012

Advisor: Joshua SanesExpression and role of cadherins in the mammalian visual system

_______________Medical Resident in Ophthalmology

University of California at San Francisco Medical Center

Adriana Eisner, Ph.D.B.A., University of California at BerkeleyEntered 2006 * Defended June 26, 2013

Advisor: Rosalind SegalPhosphatome RNAi Screen Iden es Eya1 as a Posi ve Regulator

Of Hedgehog Signal Transduc on_______________

AssociateMcKinsey & Company

Soyon Hong, Ph.D.B.S., University of Washington

Entered 2006 * Defended April 18, 2012Advisor: Dennis Selkoe

Probing the in vivo economy of amyloid beta proteinduring the development of Alzheimer’s disease type pathology

_______________Postdoctoral Fellow

Harvard Medical School—Stevens Lab

Kuo Hua Huang, Ph.D.B.S., Na onal Tsinghua University

Entered 2006 * Defended April 18, 2012Advisor: Florian Engert

Control of Turning Behaviors by Spinal Projec on Neurons_______________

Postdoctoral FellowFriedrich Meisser Ins tute, Switzerland

Page 89: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Yessenia Ibarra, Ph.D.B.S., San Diego State University

Entered 2006 * Defended February 14, 2014Advisor: David Clapham

Characteriza on of human TRPA1 and TRPV1 channelsin response to naturally occurring defensive compounds

_______________Public Health Associate

Center for Disease Control (Boston)

Mar n Kurtev, A.M.B.S., Massachuse s Ins tute of Technology

Entered 2006 * Withdrew with Master’s degree on 3/31/2014Advisor: David Clapham

_______________Whereabouts unknown

Brendan Lehnert, Ph.D.B.S., Stanford University

Entered 2006 * Defended April 10, 2012Advisor: Rachel Wilson

Dis nct roles of TRP channels in auditory transduc onand ampli ca on in Drosophila

_______________Postdoctoral Fellow

Harvard Medical School—Ginty Lab

Alan Mardinly, Ph.D.B.A., University of Chicago

Entered 2006 * Defended January 16, 2013Advisor: Michael E. Greenberg

Regula on of Synapse Development by Ac vity DependentTranscrip on in Inhibitory Neurons

_______________Postdoctoral Fellow

University of California, Berkeley—Adesnik Lab

Page 90: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Amy Meesun Ni, Ph.D.B.A., Dartmouth University

Entered 2006 * Defended December 19, 2011Advisor: John Maunsell

Visual A en on and the Role of Normaliza on

_______________Analyst

The New Teachers ProjectBrooklyn, New York

Alexandra Smolyanskaya, Ph.D.B.S., State University of New York at Stony Brook

Entered 2006 * Defended August 24, 2012Advisor: Richard T. Born

Dynamics of cor cal decision circuitsduring changes in the delity of sensory representa ons

_______________Data Scien st

S tch Fix

Michael Soskis, Ph.D.B.A., Yale University

Entered 2006 * Defended June 20, 2012Advisor: Michael E. Greenberg

A chemical gene c study of EphB receptor tyrosine kinasesignaling in the developing nervous system

_______________Food Opera ons ManagerSpring: Dine on Demand

David Stoppel, Ph.D.B.A., New York University

Entered 2006 * Defended January 29, 2014Advisor: Ma hew Anderson

Social Behavior and Gene Expression Disturbances in Mouse Modelsof Angelman Syndrome and Idic15 Au sm

_______________Whereabouts Unknown

Page 91: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ashlee Van’t Veer, Ph.D.B.A., Rutgers University

Entered 2006 * Defended January 24, 2013Advisor: Bill Carlezon

Role of Kappa Opioid Receptors in Stress Induced Behaviors_______________

Health Program SpecialistNa onal Ins tute of Neurological Disorders

Nina Vujovi , Ph.D.B.A., University of Virginia

Entered 2006 * Defended October 8, 2013Advisor: Cli ord Saper

Hypothalamic neural circuits underlying circadian organiza on ofbehavior and physiology

_______________Postdoctoral Fellow

Harvard Medical School — Medical Chronobiology Program

Oulu Wang, Ph.D.A.B., Harvard College

Entered 2006 * Defended April 13, 2012Advisor: Joseph Majzoub

Glucocor coids Promote Obesity and Suppressionof Reproduc ve Func on During Stress

_______________Enrolled in Harvard Law School in August 2015

Zheng (Herbert) Wu, Ph.D.B.S., Fudan University

Entered 2006 * Defended July 25, 2013Advisor: Catherine Dulac

Molecular Dissec on of Neural Circuitsunderlying Parental Behavior in Mice

_______________Postdoctoral Fellow

Columbia University—Shadlen Lab

Page 92: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Allison Rosen Bialas, Ph.D.B.A., Cornell University

Entered 2007 * Defended April 29, 2013Advisor: Beth Stevens

Ini a ng Complement dependent Synap c Re nement: Mechanisms ofNeuronal C1q Regula on

_______________Postdoctoral Fellow

Harvard Medical School — Carroll Lab

Mónica Feliú Mójer, Ph.D.B.S., University of Puerto Rico

Entered 2007 * Defended January 15, 2013Advisor: Joshua Kaplan

Coupling the small GTPase Rab3 to the Synap c Vesicle Cycle_______________Program Manager

Ciencia Puerto Rico

David Ferrero, Ph.D.B.S., M.S., Swiss Federal Ins tute of Technology

Diploma, Imperial College LondonEntered 2007 * Defended December 11, 2012

Advisor: Steve LiberlesOlfactory Control of Ins nc ve Behavior

_______________Consultant

The Boston Consul ng Group

Vijay Ganesh, M.D./Ph.D.A.B., Harvard College

Entered 2007 * Defended June 19, 2012Advisor: Christopher A. Walsh

Iden ca on and Characteriza on of Genes Essen al for Human BrainDevelopment

_______________Returned to Harvard Medical School

Residency at Massachuse s General Hospital

Page 93: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jonathan Garst Orozco, Ph.D.A.B., Washington University

Entered 2007 * Defended May 9, 2014Advisor: Bence Ölveczky

Learning Related Changes in the Func onal Connec vityWithin the Zebra Finch Song Control Circuit

_______________

Senior Scien stP zer Pharmaceu cals

Julie e Han, Ph.D.B.S., University of California at Los AngelesEntered 2007 * Defended April 18, 2012

Advisor: Mustafa SahinThe Role of TSC in Oligodendrocyte Di eren a on and Myelina on

_______________Engagement ManagerMcKinsey & Company

Jocelyn LeBlanc, Ph.D.B.A., Colby College

Entered 2007 * Defended February 25, 2013Advisor: Michela Fagiolini

Cri cal period plas city and sensory func onin a neuroligin 3 model of au sm

_______________Postdoctoral fellow

Harvard Medical School—Nelson Lab

Emily Lehrman, Ph.D.B.A., Stanford University

Entered 2007 * Defended April 30, 2014Advisor: Beth Stevens

Innate Immune Molecules Direct Microglia Mediated DevelopmentalSynap c Re nement_______________

Postdoctoral FellowBoston Children’s Hospital—Stevens Lab

Page 94: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Shan Lou, Ph.D.B.S., Fudan University

Entered 2007 * Defended April 1, 2013Advisor: Qiufu Ma

Development and Func ons of C Low Threshold Mechanoreceptors_______________

Postdoctoral FellowHarvard University—Adam Cohen Lab

Eric Luth, Ph.D.B.A., Colby College

Entered 2007 * Defended April 1, 2014Advisor: Dennis Selkoe

Physiological and Pathological Characteriza onof Alpha Synuclein Oligomers

_______________TEACRS Postdoctoral Fellow

Tu s University School of Medicine

Athar Malik, M.D./Ph.D.B.S., Johns Hopkins University

Entered 2007 * Defended May 9, 2013Advisor: Michael Greenberg

Genome wide iden ca on and characteriza on ofs mulus responsive enhancers in the nervous system

_______________Resident in Neurosurgery

Massachuse s General Hospital

Lia Yoon Hong Min, Ph.D.B.S., B.F.A., University of Michigan at Ann Arbor

Entered 2007 * Defended July 26, 2012Advisor: Michela Fagiolini

Experience Dependent Loss of Cross Modal Plas cityin Mouse Visual Cortex

_______________Art Science Research Fellow

University of Michigan School of Art & Design

Page 95: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kathleen Quast, Ph.D.B.F.A., University of Wisconsin at Madison

Entered 2007 * Defended September 17, 2012Advisor: Takao Hensch

Func onal Development and Plas city of Parvalbumin Cells in VisualCortex: Role of Thalamocor cal Input

_______________Postdoctoral Fellow

Baylor College of Medicine—Arenkiel Lab

Heather Rice, Ph.D.B.A., University of Oklahoma at NormanEntered 2007 * Defended May 3, 2013

Advisor: Dennis SelkoeRegula on of the proteoly c processing and func on of Amyloid

Precursor Protein by candidate ligands_______________

Postdoctoral FellowKU Leuven—De Strooper Lab

Douglas Ru , Ph.D.B.S., College of William and Mary

Entered 2007 * Defended January 12, 2012Advisor: Richard T. Born

Neuronal Tuning and Its Role in A en on_______________

Postdoctoral FellowUniversity of Pi sburgh—Cohen Lab

John Salogiannis, Ph.D.B.A., B.S., University of Maryland at College Park

Entered 2007 * Defended June 27, 2013Advisor: Michael Greenberg

Regula on of excitatory synapse development by the RhoGEF Ephexin 5_______________

Postdoctoral FellowHarvard Medical School —Reck Peterson Lab

Page 96: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Daniel E. Smith, A.M.B.S., Carnegie Mellon UniversityEntered 2007 * Le July 1, 2010

Advisor: Clay Reid_______________

Withdrew with Master’s degreeWhereabouts Unknown

Alice Yiquing Wang, Ph.D.B.A., Columbia University

Entered 2007 * Defended July 26, 2012Advisor: Naoshige Uchida

Neural substrates of choosing ac ons and mo va onal drive,a role for the striatum_______________

Postdoctoral FellowHarvard Medical School—Harvey Lab

Sara J. Fenstermacher, Ph.D.B.A., Bucknell University

Entered 2009 * Defended February 27, 2015Advisor: Rosalind Segal

From nucleus to axon: Spa al regula on of bclw mRNApromotes neurotrophin dependent axon survival

_______________Postdoctoral Fellow

Columbia University — Jessell Lab

Mehmet Fisek, Ph.D.A.B., Brandeis University

Entered 2008 * Defended April 17, 2014Advisor: Rachel Wilson

Connec vity and Computa onsin Higher Order Olfactory Neurons in Drosophila

_______________Postdoctoral Fellow

University College London—Hausser Lab

Page 97: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Allison Provost, Ph.D.B.A., University of California, Berkeley

Entered 2008 * Defended March 10, 2015Advisor: Venkatesh Murthy

Serotoninergic Modula on of the Olfactory Bulb_______________

Scien c Program ManagerOrion Bionetworks, Inc.

Ting Ren, A.M.B.S., Beijing Normal University

Entered 2008 * Le September 1, 2010Advisor: Catherine Dulac

_______________Withdrew with Master’s degree

Vice President of a company in Beijing, China(no English transla on given)

Cameron Sadegh, M.D./Ph.D.S.B., Massachuse s Ins tute of TechnologyEntered 2008 * Defended June 14, 2013

Advisor: Je rey MacklisDirected di eren a on of mouse embryonic stem cells

into neocor cal output neurons_______________

Intern in NeurosurgeryMassachuse s General Hospital

Arpiar Saunders, Ph.D.B.A., Swarthmore College

Entered 2008 * Defended July 28, 2014Advisor: Bernardo Saba ni

A direct GABAergic output from the basal ganglia to frontal cortex_______________

Postdoctoral FellowHarvard Medical School — McCarroll Lab

Page 98: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Alexandra Krol, Ph.D.B.A., M.Sci, Cambridge University

Entered 2008 * Defended March 20, 2015Advisor: Lisa Goodrich

Regula on of neuronal dendrite development and migra onby the atypical cadherin Fat3

_______________Postdoctoral Fellow

Massachuse s Ins tute of Technology—Feng Lab

Allison Nishitani, Ph.D.B.A., University of Southern CaliforniaEntered 2008 * Defended July 25, 2014

Advisor: Lisa GoodrichThe role of Netrin 1 in semicircular canal morphogenesis

_______________Teacher

Choate Rosemary Hall

Ian Oldenburg, Ph.D.B.S., Carnegie Mellon University

Entered 2008 * Defended July 11, 2014Advisor: Bernardo Saba ni

Basal Ganglia Modula on of Cor cal Firing Ratesin a Behaving Animal_______________

Postdoctoral FellowUniversity of California, Berkeley — Adesnik Lab

Rajesh Poddar Ph.D.A.B., Princeton University

Entered 2008 * Defended April 9, 2015Advisor: Bence Ölveczky

A high throughput system for automated trainingcombined with con nuous long term neural recordings in rodents

_______________Quan ta ve AnalystJump Trading, L.L.C.

Page 99: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

David Strochlic, Ph.D.B.A., Tu s University

Entered 2008 * Defended May 5, 2015Advisor: Steve Liberles

Molecular and Gene c Analysis of the Vagus Nerve_______________

Postdoctoral FellowHarvard Medical School—Liberles Lab

Monica Thanawala, Ph.D.A.B.., Harvard University

Entered 2008 * Defended May 1, 2014Advisor: Wade Regehr

Control of Neurotransmi er Release Proper esby Presynap c Calcium

_______________Postdoctoral Fellow

Harvard University—Zhuang Lab

Mary Wertz, Ph.D.B.A., Boston College

Entered 2008 * Defended April 13, 2015Advisor: Mustafa Sahin

Aberrant microRNA Expression in SMA Motor Neurons_______________

Postdoctoral FellowHarvard Medical School—Sahin Lab

Jessica Yano, A.M.B.S., California Ins tute of TechnologyEntered 2008 * Withdrew June 18, 2012

_______________

Withdrew with a Master’s degreeResearch Scien st

California Ins tute of Technology (Hsiao Lab)

Page 100: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Wenjia You, Ph.D.B.A., Tsinghua University

Entered 2008 * Defended May 7, 2015Advisor: Connie Cepko

Lineage and Func onal Analyses of Speci c Subsetsof Re nal Progenitor Cells

_______________Consultant

McKinsey & Company

Allison Baker Chang, Ph.D.B.A., Dartmouth University

Entered 2009 * Defended April 28, 2015Advisor: Rachel Wilson

Encoding and decoding mechanosensory informa onon mul ple mescales_______________

Enrolled in Harvard Medical School August 2015

Joseph Bell, M.D./Ph.D.A.B., Harvard University

Entered 2009 * Defended April 17, 2015Advisor: Rachel Wilson

Non linear s mulus integra on in olfactory behavior_______________

Returned to Harvard Medical School

Luke Bogart, Ph.D.B.A., Rochester University

Entered 2009 * Defended August 14, 2015Advisor: Takao Hensch

Synap c speci city and plas city in parvalbumin basket cell circuits_______________

Plans uncertain as of August 2015

Page 101: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Anna Chambers, Ph.D.B.A., Johns Hopkins University

Entered 2009 * Defended December 9, 2014Advisor: Daniel Polley

Progressive recovery of cor cal and midbrain sound feature encodingFollowing profound cochlear neuropathy

_______________Postdoctoral Fellow

University of Mainz—Rumpel Lab

Timothy Dunn, Ph.D.B.A., University of California, BerkeleyEntered 2009 * Defended April 20, 2015

Advisor: Florian EngertBrain wide neural dynamics underlying looming evoked escapes

and spontaneous explora on_______________

Postdoctoral FellowHarvard University Engert Lab

Jessica Hauser, Ph.D.B.A., Oberlin College

Entered 2009 * Defended July 17, 2014Advisor: Chinfei Chen

Mechanisms Shaping Excitatory Transmissionat the Developing Re nogeniculate Synapse

_______________Returned to Baylor College of Medicine

to complete MD/Ph.D. program

David Hildebrand, Ph.D.B.A., Northeastern University

Entered 2009 * Defended March 23, 2015Advisor: Florian Engert

Whole brain func onal and structural examina onin larval zzebra sh_______________

Postdoctoral FellowThe Rockefeller University—Freiwald Lab

Page 102: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Tilly Kimm, Ph.D.B.A., Massachuse s Ins tute of TechnologyEntered 2009 * Defended December 3, 2014

Advisor: Bruce BeanPotassium and Sodium Currents Regula ng

Pacemaking and Burst Firing in Substan a Nigra Dopamine Neurons_______________

Professional Tango Dancer

Dimitar Kostadinov, Ph.D.B.A., McGill University

Entered 2009 * Defended May 13, 2015Advisor: Joshua Sanes

Mechanism and func on of dendri c self avoidance and self/non selfdiscrimina on in the mammalian nervous system

_______________Postdoctoral Fellow

University College London—Hausser Lab

Andrew Lutas, Ph.D.B.A., Pennsylvania State University

Entered 2009 * Defended April 7, 2015Advisor: Gary Yellen

Cellular Metabolism Modulates Ion Channelsthat Regulate Neuronal Excitability

_______________Postdoctoral Fellow

Harvard Medical School—Andermann Lab

Caleigh Mandel BrehmB.A., University of Wisconsin—Madison

Entered 2009 * Defended October 28, 2015Advisor: Michael E. Greenberg

A Behavioral and Molecular Approachfor Understanding Angelman Syndrome

_______________Postdoctoral Fellow

Harvard Medical School—Greenberg Lab

Page 103: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Alicia NugentB.A., University of California, Berkeley

Entered 2009Advisor: Elizabeth Engle

_______________S ll enrolled

Stanislav PashkovskiB.S., Brandeis University

Entered 2009Advisor: Sandeep Robert Da a

_______________S ll enrolled

Rebecca Reh, Ph.D.B.A., University of Washington

Entered 2009 * Defended July 16, 2015Advisor: Takao Hensch

A rapid peak in gamma power signals cri cal period plas cityin the developing mouse visual cortex

_______________

Plans uncertain as of August 2015

Jason Yamada Han , Ph.D.B.A., Swarthmore College

Entered 2009 * Defended December 16, 2014Advisor: Bruce Bean

Regula on of hippocampal pyramidal neuron excitabilityBy subthreshold voltage dependent conductances

_______________Star ng up an educa onal outreach company

(as of August 2015)

Page 104: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Daniel BearB.A., Harvard College

Entered 2010Advisor: Sandeep Robert Da a

_______________S ll enrolled

Rachel DonahueB.A., Bowdoin College

Entered 2010Advisor: William Carlezon

_______________S ll enrolled

Alan EmanuelB.S., Washington State University

Entered 2010Advisor: Michael Do

_______________S ll enrolled

Catherine HartmanB.A., Duke University

Entered 2010Advisor: Michael Wolfe

_______________Withdrew without taking degree

Associate, The Boston Consultant Group

Page 105: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Xun (Helen) HouB.S., Massachuse s Ins tute of Technology

Entered 2010Advisor: Bernardo Saba ni

_______________S ll enrolled

Wen Fan Hu, M.D./Ph.D.B.A., Yale University

Entered 2010 * Defended June 10, 2014Advisor: Christopher A. Walsh

Building a Bigger Brain: Centriole Controleof Cerebral Cor cal Development

_______________Returned to Harvard Medical School

Divya Jayaraman, M.D./Ph.D.B.A., Harvard College

Entered 2010 * Defended June 15, 2015Advisor: Christopher A. Walsh

The role of centriole biogenesis in control of brain size_______________

Returned to Harvard Medical School

Wendy Liu, M.D./Ph.D.B.A., Princeton University

Entered 2010 * Defended April 14, 2014Advisor: Rachel Wilson

When Hot Is Not the Opposite of Cold:Thermosensory Processing in the Drosophila Brain

_______________Returned to Harvard Medical School

Page 106: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ian McLachlanB.S., University of Michigan (Ann Arbor)

Entered 2010Advisor: Maxwell Heiman

_______________S ll enrolled

Daniel MillmanB.A., M.S., Johns Hopkins University

Entered 2010Advisor: Venkatesh Murthy

_______________S ll enrolled

Sarah PeaseB.S., Brandeis University

Entered 2010Advisor: Rosalind Segal

_______________S ll enrolled

Alexander Murphy, Ph.D.B.A., Massachuse s Ins tute of TechnologyEntered 2010 * Defended April 30, 2015

Advisor: Je rey MacklisRNA and protein networks that locally control brain wiring

during development_______________

ConsultantMcKinsey & Company

Page 107: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Michael Susman, M.D./Ph.D.B.A., Stanford University

Entered 2010 * Defended June 24, 2015Advisor: Michael E. Greenberg

The Wnt5a Ror Signaling Pathway in the Morphogenesisof the Embryo

_______________Returned to Harvard Medical School

Taralyn TanB.S., Oregon State University

Entered 2010Advisor: Sandeep Robert Da a

_______________S ll enrolled

Ju TianB.E., Tsinghua University

Entered 2010Advisor: Naoshige Uchida

_______________S ll enrolled

David Roberson, Ph.D.B.A., M.B.A., Harding University

Entered 2010 * Defended September 18, 2015Advisor: Cli ord Woolf

Analysis of voluntary behavior to interrogate neural func on_______________

Unknkown

Page 108: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Alexander TroB.S., Brandeis University

Entered 2010Advisor: Rick Born

_______________S ll Enrolled

Caroline WeeB.A., Cornell University

Entered 2010Advisor: Florian Engert

_______________S ll enrolled

Alexander WiltschkoB.E., Tsinghua University

Entered 2010Advisor: Naoshige Uchida

_______________S ll enrolled

Willie TobinB.S., University of California, Santa Cruz

Entered 2010 *Advisor: Rachel Wilson

_______________S ll enrolled

Page 109: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ryan Dosumu JohnsonB.S., University of California, Los Angeles

Entered 2011Advisor: Susan Dymecki

_______________S ll Enrolled

Laura DriscollB.S., University of California, Berkeley

Entered 2011Advisor: Christopher Harvey

_______________S ll enrolled

Masha BloomB.A., Barnard College

Entered 2011Advisor: Sandeep Robert Da a

_______________S ll enrolled

Lauren CarvalhoA.B., Harvard College

Entered 2011 * Withdrew July 26, 2013Advisor: Venkatesh Murthy

_______________

Withdrew without degreeEnrolled at Tu s Veterinary School

Page 110: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Nick JikomesB.S., University of Wisconsin, Madison

Entered 2011Advisor: Mark Andermann

_______________S ll enrolled

Mounir Koussa, Ph.D.B.A., University of Arizona

Entered 2011 * Defended April 15, 2015Advisor: David Corey

The Biophysics of Vertebrate Hearing: A Single Molecule Approach_______________

Founder of Canary, Inc.

GaYoung LeeB.S., Korea Advanced Ins tute of Science and Technology

Entered 2011Advisor: Tom Schwarz

_______________S ll enrolled

Neir Eshel, M.D./Ph.D.B.A., Princeton University

M.S.C., University College, LondonEntered 2011 * Defended May 9, 2014

Advisor: Naoshige UchidaArithme c and Local Circuitry Underlying Dopamine Predic on Errors

_______________Returned to Harvard Medical School

Page 111: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Jinyue LiuB.A., Cambridge University

Entered 2011Advisor: Joshua Sanes

_______________S ll enrolled

Ellio MilnerB.A., Oberlin College

Entered 2011Advisor: Michael Do

_______________S ll enrolled

Zhihao (Thomas) LuoA.B., Harvard College

Entered 2011Advisor: John Maunsell

_______________S ll enrolled

Elizabeth LitvinaB.A., M.A., Wesleyan University

Entered 2011Advisor: Chinfei Chen

_______________S ll enrolled

Page 112: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Nicole NeubarthB.A., University of Chicago

Entered 2011Advisor: David Ginty

_______________S ll enrolled

Tim OtchyB.A., Georgia State University

M.A., Tu s UniversityB.S., Georgia Ins tute of Technology

Entered 2011Advisor: Bence Ölveczky

_______________S ll enrolled

Shay NeufeldB.S., McGill University

Entered 2011Advisor: Bernardo Saba ni

_______________S ll enrolled

Ari MorcosB.S., University of California, San Diego

Entered 2011Advisor: Christopher Harvey

_______________S ll enrolled

Page 113: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Alex SimonB.A., Lewis and Clark College

Entered 2011Advisor: Michela Fagiolini

_______________S ll enrolled

Gregory SousaB.S., Bates College

Entered 2011 * Withdrew September 1, 2011

_______________Withdrew without degree

Enrolled in University of Pennsylvania Veterinary School

Priya Srikanth, M.D./Ph.D.B.A., Washington University

Entered 2011 * Defended May 11, 2015Advisor: Dennis Selkoe

Schizophrenia relevant DISC1 interrup on reveals subtle altera onsin Wnt signaling and cell fate in human iPSC derived neurons

_______________Returned to Harvard Medical School

Hye Yeon Park, A.M.B.S., M.S., Korea Advanced Ins tute of Science and TechnologyEntered 2011 * Withdrew with Master’s degree 11/14/2014

Advisor: Maria Leh nen

_______________Scien st

Novar s, Inc.

Page 114: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Zhaoying XuB.A., Grinnell College

Entered 2011Advisor: Je rey Macklis

_______________S ll enrolled

Zhanlei YeB.Sc., Beijing University

Entered 2011Advisor: Takao Hensch

_______________S ll enrolled

Zhiqi (Candice) YipB.S., University of Southern California

Entered 2011Advisor: Maxwell Heiman

_______________S ll enrolled

Erika WilliamsB.A., Williams College

Entered 2011Advisor: Steve Liberles

_______________S ll enrolled

Page 115: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Gregory BrymanB.S., University of Rochester

Entered 2012Advisor: Michael Do

_______________S ll enrolled

Selmaan Che hB.A., Swarthmore College

Entered 2012Advisor: Christopher Harvey

_______________S ll enrolled

Benjamin AndreoneB.S., University of Maryland—College Park

Entered 2012Advisor: Chenghua Gu

_______________S ll enrolled

YunXiang Chu, M.D./Ph.D.B.A., Stanford University

Entered 2012 * Defended April 15, 2014Advisor: Wade Regehr

The Role of Protein Kinase C in Short Term Plas city_______________

Returned to Harvard Medical School

Page 116: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ershela DurresiB.S., M.S., Yale University

Entered 2012Advisor: Michael E. Greenberg

_______________S ll enrolled

Richard HeldB.S., Tulane University

Entered 2012Advisor: Pascal Kaeser

_______________S ll enrolled

Michael CoulterA.B., Harvard College

Entered 2012Advisor: Christopher Walsh

_______________S ll enrolled

Benyam KindeB.S., University of Maryland—Bal more County

Entered 2012Advisor: Michael E. Greenberg

_______________S ll enrolled

Page 117: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ma hias MindererB.S., University of California, San Diego

Entered 2012Advisor: Christopher Harvey

_______________S ll enrolled

Tara RaamB.S., University of California, San Diego

Entered 2012Advisor: Amar Sahay

_______________S ll enrolled

Gil MandelbaumB.S., Tel Aviv University

Entered 2012Advisor: Bernardo Saba ni

_______________S ll enrolled

Rohan RameshB.S., Brown University

Entered 2012Advisor: Mark Andermann

_______________S ll enrolled

Page 118: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Christopher TzengB.A., Rice University

Entered 2012Advisor: Michael E. Greenberg

_______________S ll enrolled

Hugo Vega RamirezB.S., University of California, Davis

Entered 2012Advisor: Amar Sahay

_______________S ll enrolled

Hannah SomhegyiB.A., Lewis and Clark College

Entered 2012Advisor: Dragana Rogulja

_______________S ll enrolled

Chris na WelshB.S., Massachuse s Ins tute of Technology

Entered 2012Advisor: Beth Stevens

_______________S ll enrolled

Page 119: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Andrea YungB.S., Stanford University

Entered 2012Advisor: Lisa Goodrich

_______________S ll enrolled

Alex BatchelorB.A., Cambridge University

Entered 2013Advisor: Rachel Wilson

_______________S ll enrolled

Yuwen WuB.A., University of Chicago

Entered 2012Advisor: Beth Stevens

_______________S ll enrolled

Michael BukwichB.S., New York University

Entered 2013Advisor: Naoshige Uchida

_______________S ll enrolled

Page 120: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Brian ChowB.S., University of California, San Diego

Entered 2013Advisor: Chenghua Gu

_______________S ll enrolled

Nicholas DeSte noB.S., B.A., University of Pi sburgh

Entered 2013Advisor: Jesse Gray

_______________S ll enrolled

Chester ChiaB.A., Rutgers University

Entered 2013Advisor: Lisa Goodrich

_______________S ll enrolled

Michelle FrankB.A., Saint Olaf College

Entered 2013Advisor: Dragana Rogulja

_______________S ll enrolled

Page 121: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Grigori GuitchountsB.A., Boston University

Entered 2013Advisor: David Cox

_______________S ll enrolled

Chong GuoB.S., University of Michigan, Ann Arbor

B.S., Cornell UniversityEntered 2013

Advisor: Wade Regehr

_______________S ll enrolled

Morgan FreretB.S., Stanford University

Entered 2013Advisor: Susan Dymecki

_______________S ll enrolled

James HarrisB.A., Bowdoin College

Entered 2013Advisor: Paola Arlo a

_______________S ll enrolled

Page 122: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kee Wui HuangB.S., M.S., Stanford University

Entered 2013Advisor: Bernardo Saba ni

_______________S ll enrolled

Minsuk HyunB.S., Stanford University

Entered 2013Advisor: Bernardo Saba ni

_______________S ll enrolled

Stephen HoltzB.S., University of Virginia

Entered 2013Advisor: Rachel Wilson

_______________S ll enrolled

Robert JohnsonB.S., Washington University

Entered 2013Advisor: Florian Engert

_______________S ll enrolled

Page 123: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Angela KimB.S., Yonsei University

M.S., Korea Ins tute of Science and TechnologyEntered 2013

Advisor: Bradford Lowell

_______________S ll enrolled

Elizabeth LamkinB.S., Haverford College

Entered 2013Advisor: Maxwell Heiman

_______________S ll enrolled

Jessleen KanwalB.S., University of Virginia

Entered 2013Advisor: Aravinthan Samuel

_______________S ll enrolled

Sung Ling (Dean) LeeB.S., University of California, Los Angeles

Entered 2013Advisor: Paola Arlo a

_______________S ll enrolled

Page 124: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Ivan San agoB.S., M.A., City College of New York

Entered 2013Advisor: Ma hew Pecot

_______________S ll enrolled

Bryan SongB.S., University of Texas, Aus n

Entered 2013Advisor: Dragana Rogulja

_______________S ll enrolled

Aleksandr (Sasha) RayshubskiyB.S., M.Eng., Cornell University

Entered 2013Advisor: Rachel Wilson

_______________S ll enrolled

Stephen ThornquistB.S., Washington University

Entered 2013Advisor: Michael Crickmore

_______________S ll enrolled

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Huaiying (Shan Shan) WangB.S., Massachuse s Ins tute of Technology

Entered 2013Advisor: Pascal Kaeser

_______________S ll enrolled

Xingjie (Stephen) ZhangB.A., Grinnell College

Entered 2013Advisor: Dragana Rogulja

_______________S ll enrolled

Josef TurecekB.S., University of Washington

Entered 2013Advisor: Wade Regehr

_______________S ll enrolled

Asa Barth MaronB.S., Lehigh University

Entered 2014Advisor: Rachel Wilson

_______________S ll enrolled

Page 126: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Cesar EchavarriaB.S., Massachuse s Ins tute of Technology

Entered 2014Advisor: David Cox

_______________S ll enrolled

Kelly GirskisB.A., University of Southern California

Entered 2014Advisor: Christopher Walsh

_______________S ll enrolled

Ma hew BaumB.S., M.S., Yale UniversityM.Sc., University of Dublin

Entered 2014Advisor: Beth Stevens

_______________S ll enrolled

Vivian HemmelderB.Sc., Universiteit van Amsterdam

Entered 2014Advisor: Naoshige Uchida

_______________S ll enrolled

Page 127: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Kris ne LyonB.A., Lewis and Clark College

Entered 2014Advisor: Susan Dymecki

_______________S ll enrolled

Michael MarquisB.S., Arizona State University

Entered 2014Advisor: Rachel Wilson

_______________S ll enrolled

Aziz KarakhanyanB.A., Rutgers University

Entered 2014Advisor: Ma hew Pecot

_______________S ll enrolled

Eric MulhallB.S., University of California, Santa Cruz

Entered 2014Advisor: David Corey

_______________S ll enrolled

Page 128: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Aurel NagyB.S., Brown University

Entered 2014Advisor: Michael E. Greenberg

_______________S ll enrolled

Paola PatellaB.S., University of GenoaM.Sc., University of Trento

Entered 2014Advisor: Rachel Wilson

_______________S ll enrolled

Sivapratha Nagappan Che arB.A., University of California, Berkeley

Entered 2014Advisor: Hisashi Umemori

_______________S ll enrolled

Noah Pe tB.Sc., University College London

Entered 2014Advisor: Christopher Harvey

_______________S ll enrolled

Page 129: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Nesibe TemizB.S., M.Sc., University of Bogazici

Entered 2014Advisor: Cli ord Woolf

_______________S ll enrolled

He YangB.S., Tsinghua University

Entered 2014Advisor: Sam Kunes

_______________S ll enrolled

Rachel Rei RodinB.S., Emory University

Entered 2014Advisor: Christopher Walsh

_______________S ll enrolled

Ee Lynn YapB.S., Yale University

Entered 2014Advisor: Michael E. Greenberg

_______________S ll enrolled

Page 130: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

James BohnslavA.B., Harvard College

Entered 2015

_______________S ll enrolled

Senan EbrahimA.B., Harvard College

Entered 2015Advisor: Syd Cash

_______________S ll enrolled

Qiyu ZhangB.S., Brandeis University

Entered 2014Advisor: David Ginty

_______________S ll enrolled

Alexander GrebenB.Sc., McGill University

Entered 2015

_______________S ll enrolled

Page 131: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Xiaomeng (Mona) HanB.S., Tsinghua University

Entered 2015

_______________S ll enrolled

Lingfeng HouB.S., Tsinghua University

Entered 2015

_______________S ll enrolled

Allison HamilosSc.B., Massachuse s Ins tute of Technology

Entered 2015

_______________S ll enrolled

Narendra JoshiB.A., Amherst College

Entered 2015

_______________S ll enrolled

Page 132: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Emma KrauseB.S., Davidson College

Entered 2015

_______________S ll enrolled

Lauren KershbergB.A., Pomona College

Entered 2015

_______________S ll enrolled

Jae Eon LeeB.S., Korea Advanced Ins tute of Science and Technology

Entered 2015

_______________S ll enrolled

Kelly McGuireA.B., Sc.B., Brown University

Entered 2015

_______________S ll enrolled

Page 133: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Alexandra MooreB.S., University of Oregon

Entered 2015

_______________S ll enrolled

Rebecca SenB.A., Swarthmore College

Entered 2015

_______________S ll enrolled

Clara StarkweatherB.S., Duke University

Entered 2015Advisor: Naoshige Uchida

_______________S ll enrolled

Aniqa TasnimB.A., Johns Hopkins University

Entered 2015

_______________S ll enrolled

Page 134: Alumni Director cover page - HMS · PDF fileNicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Po ©er A Physiological and Histological

Nivanthika WimalasenaA.B., Harvard College

Entered 2015

_______________S ll enrolled