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EVALUATING THE TB HAZARD POSED TO CATTLE FROM WILDLIFE ACROSS EUROPE
Joanne L. Hardstaff, Glenn Marion, Michael R. Hutchings and Piran C.L. White
APPENDIX A
Distribution of wild species
1
Figure A1. The distribution of badgers (top left), fallow deer (top right), red deer (middle left), roe deer
(middle right), wild boar (bottom left). Black shows species presence, pale green species absence and blue
inadequate information.
2
APPENDIX B
The results of the sensitivity analyses using Latin hypercube sampling.
Table B. The sensitivity results from the LHS.Input into LHS
Lower bounds Upper bounds R2 R2 R2
Prev Density Overlap PrevDensity Overlap Country Species R2 GCV Overlap Density Prev
0 0 0 0.153 49.5 1 UK Badger 73.6 0.3325 23.5 24.6 25.7
0 0 0 0.185 6.42 1 Fallow deer 72.6 0.0082 24.3 23.8 24.1
0 0 0 0.015 30.5 1 Red deer 72.6 0.0012 24.7 24 24.5
0 0 0 0.015 28.5 1 Roe deer 73.5 0.0011 24.3 24.8 24.5
0 0 0 0.0001 0.312 1 Czech Badger 72.9 0.3303 24.4 24.1 23.9
0 0 0 0.0001 0.25 1 Fallow deer 100 0.0000 0.00929 100 100
0 0 0 0.0001 19.5 1 Red deer 72.8 0.0000 24.4 24.1 25.2
0 0 0 0.0001 25.5 1 Roe deer 73.1 0.0000 25.3 24.7 23.6
0 0 0 0.0001 0.56 1 Wild boar 75.3 0.0000 22.8 31.6 22.5
0 0 0 0.02 1.6 1 France Badger 72.8 0.3317 24 24.2 24.4
0 0 0 0.24 8 1 Red deer 73.6 0.0212 24.2 24.7 24.3
0 0 0 0.019 25 1 Roe deer 73.7 0.0013 23.8 23.1 27.2
0 0 0 0.42 10 1 Wild boar 73.1 0.1036 24.8 25 23.8
0 0 0 0.238 1 1 Spain Badger 73.1 0.3319 24.1 24.8 23.5
0 0 0 0.287 20 1 Fallow deer 72.8 0.1896 24.6 23.7 23.4
0 0 0 0.4 14 1 Red deer 72.5 0.1865 24.4 24.4 24.3
0 0 0 0.06 7.5 1 Roe deer 75.4 0.0012 22.6 23.6 32
0 0 0 1 10 1 Wild boar 73.2 0.5723 25.1 24.6 23.8
0 0 0 0.0001 5.9 1 Germany Badger 73 0.3299 24.6 24.4 24.5
0 0 0 0.0001 0.263 1 Fallow deer 100 0.0000 0.0026 100 100
0 0 0 0.0001 0.304 1 Red deer 73 0.0000 24.9 24.2 24.2
0 0 0 0.0001 9.5 1 Roe deer 73.3 0.0000 25.1 24.1 24
0 0 0 0.0001 5.13 1 Wild boar 73 0.0000 24 23.4 24.5
0 0 0 0.0001 0.015 1 Sweden Badger 73.5 0.0000 24.8 23.9 24.2
0 0 0 0.0001 0.1 1 Fallow deer 73.3 0.0000 25.1 24 24.1
0 0 0 0.0001 0.1 1 Red deer 0.016 0.0000 0.00851 0.0022 0.005
0 0 0 0.0001 14 1 Roe deer 72.8 0.0000 24.6 24.4 24.4
0 0 0 0.0001 1.33 1 Wild boar 75 0.0000 22.4 30.1 22.5
3
APPENDIX C
Information on literature used for species densities, distribution and TB prevalence. Question marks in
the tables indicate that a species is known to be present but we were unable to locate any relevant data.
Gaps in the tables indicate that there are no records relating to whether a species is present.
4
Table C1. Data used for badger densities and distribution across Europe
Badger densities Badger distribution
CountryDensities (km2) References
Max Density used References
Austria ? ? ? ?Belgium ? ? ? ?Bulgaria 2.67 Grigorov (1986) via wwf 2.67Croatia ? ? ? ?Czech Republic
0.118-0.312 Matyastik and Bicik (1999) 0.312
Denmark 1.5 Aaris-Sorensen (1995) 1.5Aaris-Sorensen et al. (1995)
Estonia ? ? ? ?Finland 0.26 Kauhala et al. (2006) 0.26 Kauhala et al. (2005)
France 0.36-0.48, 1.5-20
Rosalino et al. (2004), Ecologie Nature (2010) 1.6 ONCFS (2010)
Germany 5.9 Keuling et al. (2010) 5.9 ?Greece ? ? ? ?
Hungary 0.095-0.324 Kozak et al. (2006) 0.325 ?
Ireland1.9-35
Muldowney et al. (2003); Olea-Popelka et al. (2003)
Italy ~1 Remonti et al. (2006) 1.2Latvia ? ? ? ?Lithuania ? ? ? ?Macedonia (FYR) 0.028 Melovski (2008) 0.029Netherlands ? ? ? ?Norway 0.12-0.588 Broseth et al. (1997) 0.59Poland 0.15-2.11 Kowalczyk et al. (2003) 2.11Portugal 0.36-0.48 Rosalino et al. (2004) 0.48Romania 4; 10 Murariu (2003) and (2008) 10Slovakia ? ? ? ?Slovenia ? ? ? ?
Spain 0.2-1 Revilla and Palomares, (2002) 1
Sweden 0.015 Cederlund et al. (2004); Annual survey,2003/4;Tannerfeldt et al. (2004)
0.015
U.K. 4.9-49.5 Delahay et al. (2006); Palphramand et al.(2007) 25.3
5
Table C2. Data used for badger TB prevalence across EuropeBadger TB prevalenceCountry Prevalences
(%)References Max prevalence
used (proportion)Mode prevalence used (proportion)
Austria ? ? ? ?Belgium ? ? ? ?Bulgaria 0.0001 0.00005Croatia ? ? ? ?Czech Republic 0.0001 0.00005Denmark ? ? 0.0001 0.00005Estonia ? ? ?Finland 0.0001 0.00005France 1.8 Zanella et al. (2008) 0.018 0.005Germany 0.0001 0.00005Greece ? ? ? ?Hungary 0.0001 0.00005
Ireland 11-28.5Olea-Popelka et al. (2003) and (2006) 0.285 0.11
Italy 0.0001 0.00005Latvia ? ? ? ?Lithuania ? ? ? ?Macedonia (FYR) 0.0001 0.00005Netherlands ? ? ? ?Norway 0.0001 0.00005Poland 0.0001 0.00005Portugal 0.0001 0.00005Romania 0.0001 0.00005Slovakia ? ? ? ?Slovenia ? ? ? ?
Spain 7 or 16.7-23.8Martin-Atance, (2006) 0.238 0.07
Sweden 0.0001 0.00005
U.K.
6-15.3
Clifton-Hadley et al. (1993); Courteney et al. (2006) 0.153 0.06
6
Table C3. Data used for fallow deer densities and distributions across Europe
Fallow deer densities Fallow deer distribution
CountryDensities (km2) References
Max Density used References
Austria 1Reimoser and Reimoser (2010) 1
Reimoser and Reimoser (2010)
Belgium 0.0118Casear and Licoppe (2010) 0.12 Casear and Licoppe (2010)
Bulgaria 0.52Genov pers. comm (2010). 1
Croatia 0.039D. Konjevic pers. comm. (2010) 0.045 Kusak and Krapinec (2010)
Czech Republic 0.24
Bartos, Kottba and Pintir (2010) 0.25
Denmark 0.14Andersen and Hothe (2010) 0.14 Andersen and Hothe (2010)
Estonia ? ? ?
Finland 0.0018Ruusila and Kojola (2010) 0.0018 Ruusila and Kojola (2010)
France ? ? ? ?Germany 0.263 Schettler et al. (2006) 0.263
Greece 0.008 Papaioannou et al. (2010) 0.008
European ungulate management (2010); Tracje pers com (2010) about fallow deer being on the boarder between Albania and Greece.;Papaioannou (2010)
Hungary
5 in game forest(inc. arable)
Palotas et al. (1996) 5 Csanyi and Lehoczki (2010)
Ireland 78.23 McElligott et al. (2002) 14 www.irishdeersociety.ie
Italy9.9-14.2; 2.9->53.8;
Rossi et al. (1994); Focardi et al. (2002); Focardi et al. 2006
14.59 Apollonio et al. (2010)
Latvia ? ? ?Lithuania ? ? ?Macedonia (FYR) ? ? ? ?
Netherlands 0.0270Van Wierene and Groot Bruinderink (2010) 0.028
Van Wierene and Groot Bruinderink (2010)
Norway ? ? ?
Poland 3 Borkowski and Pudelko (2007) 4
Portugal 0.0053-0.0325 Vingada et al. (2010) 0.32 Vingada et al. (2010)
Romania 0.025 Micu et al. (2010) 0.025 Micu et al. (2010)
Slovakia 0.15Find’o and Skuban (2010) 0.15
Slovenia 2.2 Stergar M., pers. comm (2011) 2.2 Stergar M., pers. comm
(2011)
Spain 3-22; 28; 12.08;
Alvarez et al. (1988); Malo et al. (2000), Marco et al. (2002); Martin-Atance et al. (2006);
3 Carranza et al. (2010)
Sweden 27 on an Cederlund et al. (2004) ; 0.1 Libery et al. (2010)
7
island, mean in oskarshamn 0.1
Tannerfeldt et al. (2004)
U.K. 5.86 Ward (2005) 6.24 British Deer Society
8
Table C4. Data used for TB prevalence in fallow deer across EuropeFallow deer TB prevalence
CountryTB prevalence (%)
References Max Prevalence used
Mode Prevalence used
Austria 0.0001 0.00005Belgium ? ? ?Bulgaria 0.0001 0.00005Croatia 0.0001 0.00005Czech Republic 0.0001 0.00005Denmark 0.0001 0.00005Estonia 0.0001 0.00005Finland 0.0001 0.00005France 0.7 Zanella et al. (2008) n/a n/aGermany 0.0001 0.00005Greece 0.0001 0.00005Hungary 0.0001 0.00005Ireland 0.16 EFSA (2008) 0.16 0.005Italy 0.0001 0.387Latvia 0.0001 0.00005Lithuania 0.0001 0.00005Macedonia (FYR) ? ? ? ?
Netherlands 0.0001 0.00005Norway 0.0001 0.00005Poland 0.0001 0.00005Portugal 0.185 EFSA (2008) 0.0001 0.185Romania 0.0001 0.00005Slovakia 0.0001 0.00005Slovenia 0.0001 0.00005
Spain 18.5, 38.7 Gortazar et al. (2008), EFSA (2008) 0.0001 0.287
Sweden 0.0001 0.00005U.K. 1.02 Delahay et al. (2007) 0.0001 0.185
9
Table C5. Data used for red deer densities and distributions across EuropeRed deer densities Red deer distribution
Country Densities (km2) References Max density
used References
Austria 0.48;1Schdmitt (2002); Reimoser and Reimoser (2010)
1 Reimoser and Reimoser (2010)
Belgium 3; 0.593 (Wallonia)
Licoppe, (2003);Casear and Licoppe (2010)
3 Casear and Licoppe (2010)
Bulgaria 1.74 Genov pers. comm, (2010) 3
Croatia 0.21 D. Konjevic, pers. comm., (2010) 0.21 Kusak and Krapinec (2010)
Czech Republic
0.9(0.1-1.7)-2.1; 19.5 av, 1st winter 30, 2nd winter 9; 0.3
Homolka, (2003), Prokesova et al. 2006; European ungulate management (2010)
19.5 & 9 & 30
Denmark 0.35 Andersen and Hothe (2010) 0.35 Andersen and Hothe (2010)
Estonia 0.022-0.044 Balciauskas et al. (2010) 0.044 Balciauskas et al. (2010)
Finland ? ? ? ?
France
0.4-0.9 (vosges), 0.34-2.67, 7-10
ONCFS (2010) doc, Pichon et al. (1999); Richard, 2010,
8ONCFS (2010); Richomme et al. 2010, stated that red deer are on Corsica.
Germany 0.304 Schettler 0.304 Worschikowsky (2010)
Greece 0.008 Papaioannou (2010) 0.008IUCN red list, via largeherbivore.org.;Papaioannou (2010)
Hungary 0.125; 6 Palotas et al. , 1996; Biro, 2006 6 Csanyi and Lehoczki (2010)
Ireland 14 www.irishdeersociety.ie
Italy 2, 0.55-5.434, 2.7
Mattioli (2001); Apollonio, (2004); Ramazin et al. (2007)
5.434 Mattioli (2001)
Latvia 0.45 European ungulate management (2010) 0.45 Balciauskas et al. (2010)
Lithuania 0.208-0.258; 0.184
Balciauskas (2009); European ungulate management (2010)
0.26 Balciauskas et al. (2010)
Macedonia (FYR) ? ? ? European ungulate
management (2010)
Netherlands 2-3; 0.065 Kuiters (2005), 3 Van Wierene and Groot Bruinderink (2010)
Norway 0.4 European ungulate management (2010) 0.4 European ungulate
management (2010)
Poland
1.8-14.1; 2.86-8.58; 0.7-2.0-carpathians; up to 12.7;
Bobek (1984); Jedrzejewska et al. (1994); Jedrezjewski, (2002); Gula et al. (2006), European ungulate management (2010)
8.58
Portugal 0.7; 0.17-0.22
Bugalho et al. (2003); European ungulate management (2010)
2 Vingada et al. (2010)
10
Romania5-8; 5-38; 1.514; 5.838-0.058;
Murariu (2003), Cristescu et al. (2007) ; European ungulate management (2010)
8
Slovakia 0.775 Find’o and Skuban (2010) 0.78 European ungulate
management (2010)
Slovenia0.5-0.7/59.9, cull density up to 2.7;
European ungulate management (2010), Trdan et al. et al. (2008); Stergar M., pers. comm (2011)
8 Stergar M., pers. comm,2011
Spain 10-98; 14;
Malo et al. (2000); Gortazar et al. (2005); Santin-Duran et al. (2005);
14 Carranza J. (2010)
Sweden
0 to 0.01, mean in oskarshamm 0.1;0.02
Cederlund et al. (2004); Annual survey,2003/4; European ungulate density; Tannerfeldt et al. (2004)
0.1 Libery et al. (2010)
U.K. 7.57-30.5 Clutton-Brock, 1982; Ward (2005) 30.5 British Deer Society
11
Table C6. Data used for TB prevalence in red deer across Europe
Red deer TB prevalence
CountryTB prevalence (%)
References Max Prevalence used
Mode Prevalence used
Austria 0.0001 0.00005Belgium 0.0001 0.00005Bulgaria 0.0001 0.00005Croatia 0.0001 0.00005Czech Republic 0.0001 0.00005
Denmark 0.0001 0.00005Estonia 0.0001 0.00005Finland ? ? ? ?
France 13-24, 21.8 Zanella et al. (2008); EFSA 2008 0.24 0.13
Germany 0.0001 0.00005Greece 0.0001 0.00005Hungary 5.88 Csivincsik(2010) 0.0588 0.0001Ireland 2.9 Quigley et al. (1997) 0.0001 2.9Italy 0.0001 0.00005
Latvia 0.0001 0.00005Lithuania 0.0001 0.00005Macedonia (FYR) ? ? ? ?
Netherlands 0.0001 0.00005Norway 0.0001 0.00005Poland 0.0001 0.00005
Portugal 14.1&19.6Direccao De Services Veterinaros De Regiao Alentejo (2010)
0.015 0.196
Romania 0.0001 0.00005Slovakia 0.0001 0.00005Slovenia 0.0001 0.00005
Spain 0-44; 0.1-2.91
Hermoso de Mendoza et al. (2006); Vicente et al. (2007); 0.0001 0.4
Sweden 0.0001 0.00005
U.K. 7.2-18.5, 4.3 Delahay et al. (2001) & (2007) 0.0001 0.015
12
Table C7. Data used for roe deer densities and distributions across Europe.Roe deer densities Roe deer distribution
Country Densities (km2) References Max Density used References
Austria 4 Reimoser and Reimoser (2010) 4 Reimoser and Reimoser
(2010)
Belgium 1.48 Flanders and 2.37 Wallonia
Casear and Licoppe (2010) 2.4 Casear and Licoppe
(2010)
Bulgaria 121,160 in 1992; 7;
wwf, Genov pers. comm. (2010) 8.5
Croatia 1.04 D. Konjevic, pers comm (2010) 1.05 Kusak and Krapinec
(2010)
Czech Republic
0.22(0.07-0.37)-0.5 (mountain) 18.4 av, 1st winter 25.5, 2nd winter 11.3; 3.7
Homolka, (2003); Prokesova et al. (2006); European ungulate management (2010)
25.5
Denmark 2.57 Andersen and Hothe (2010) 2.56 Andersen and Hothe
(2010)
Estonia 1.2;1.1Sumilo et al. (2008); European ungulate management (2010)
1.2 Balciauskas et al. (2010)
Finland 0.045-0.06 European ungulate management (2010) 0.06 Ruusila and Kojola
(2010)
France 3; 2-25; 0.9-4.8, 5-25; 8-10;
Vincente et al. (1995), ONC;Pichon et al. (1999); Pays et al.(2007)
5
Germany 7.6-9.5;5.8 Radeloff, 2000; Schettler, 2006 9.5
Greece
13.9;continental thrace,macedonia, epirus,thessaly=0.14-0.39 to 0.58-1.82; Epirius 1.81-2.22;
Tsaparis et al. (2008); Giannakopoulos -roe deer meeting;European ungulate management (2010)
13.9Giannakopoulos (2009);Papaioannou (2010)
Hungary
2-6 nationally, 6 in game forest(with arable included); 2.5; 2.4 to 4.6
Palotas et al. (1996), Biro, (2006); Kiraly (2007)
6 Csanyi and Lehoczki (2010)
Ireland N/A N/A N/A N/A
Italy 5-7; 8.5, 6.11, 12.2, 1.8 and 4.34
Focardi et al. (2002); Apollino (2004); Borger (2006); Ramazin et al. (2007); Rizzoli (2009)
12.4 Apollonio et al. (2010)
Latvia 2;2Sumilo et al. (2008); European ungulate management (2010)
2 Balciauskas et al. (2010)
Lithuania 1.3;0.686-1.06; 1.23
Sumilo et al. (2008); Balciauskas (2009); European ungulate management (2010)
1.3 Balciauskas et al. (2010)
Macedonia (FYR) 0.083333333 Melovski, 2008 0.083
Netherlands 1.43 Van Wierene and Groot Bruinderink (2010) 1.43 Van Wierene and Groot
Bruinderink (2010)
Norway 0.124-0.154 European ungulate management (2010) 0.154 Andersen
Poland 1.15-8.2; up to 20Jedrzejewski (2002); European mammal management
8.2
13
Portugal 0.033-0.054 Vingada et al. (2010) 0.7 Vingada et al. (2010)
Romania 10-15; 16; 0-5; 0.7; 1.09-0.34;
Murariu (2003) & (2008); Cristescu et al. (2007); European ungulate management (2010)
15.1
Slovakia 1.63-1.73 Find’o and Skuban (2010) 1.73
Slovenia 1.73-2.22; 3-6/52.247
Pokorny, RoeDeerConference; European ungulate management (2010),Stergar M., pers com,2011
6.1 Stergar M., pers com,2011
Spain 3; 7.5; 4.29-23.6; 2-18;
Herrero,99, Saez-Royez and tellaria1991 via Acevedo,2005, Mateos-Quesada, 2005;Aragon 1995.
7.5
Sweden
14; 4.8-9.36; mean in oskarshamn 5;0.27,0.01-3.5;
Cederlund et al. (2004); Kjellander et al. (2004); Annual survey,2003/4; European ungulate management (2010); Tannerfeldt et al. (2004) Wiken-Ros (2009);
14 Libery et al. (2010)
U.K. 7.2; 16.55; 28.2;Chapman et al. (1993); Hemami (2005); Ward et al. (2005)
28.2 British Deer Society
14
Table C8. Data used for TB prevalence in roe deer across Europe
Roe deer TB prevalence
CountryTB prevalence (%)
References Max Prevalence used
Mode Prevalence used
Austria 0.0001 0.00005Belgium 0.0001 0.00005Bulgaria 0.0001 0.00005Croatia 0.0001 0.00005Czech Republic 0.0001 0.00005
Denmark 0.0001 0.00005Estonia 0.0001 0.00005Finland 0.0001 0.00005France 1.89 Zanella et al. (2008) 0.019 0.005Germany 0.0001 0.00005Greece 0.0001 0.00005Hungary 0.0001 0.00005Ireland N/A N/A N/A N/AItaly 2.27 Balseiro et al. (2009) 2.27 0.005Latvia 0.0001 0.00005Lithuania 0.0001 0.00005Macedonia (FYR) 0.0001 0.00005Netherlands 0.0001 0.00005Norway 0.0001 0.00005Poland 0.0001 0.00005Portugal 0.0001 0.00005Romania 0.0001 0.00005Slovakia 0.0001 0.00005Slovenia 0.0001 0.00005Spain 0.60 Balseiro et al. (2009) 0.06 0.006Sweden 0.0001 0.00005U.K. 1.02 Delahay et al. (2007) 0.015 0.006
15
Table C9. Data used for wild boar densities and distributions across Europe
Wild boar densities Wild boar distribution
Country Densities (km2) References Max Density
used References
Austria 4 Reimoser and Reimoser (2010) 4 Reimoser and
Reimoser (2010)
Belgium 1.246 Casear and Licoppe (2010) 1.25 Casear and Licoppe (2010)
Bulgaria 6.35 Genov pers. comm. (2010) 7.5
Croatia 0.34 Konjevic D., pers. comm. 2010. 0.342 Kusak and Krapinec
(2010)
Czech Republic 0.56 Bartos et al. (2010) 0.56
Denmark ? ? ?
Estonia 0.45;0.154Sumilo et al. (2008); European ungulate management (2010)
0.45 Balciauskas et al. (2010)
Finland 1.33 (Sweden) 1.33 Ruusila and Kojola (2010)
France 0.6-2.9, 0.03 - 4.77
Dardaillon et al. (1986);ONCFS (2010) 10
Germany 5.13 minimum; Keuling et al. (2008) 5.13
Greece
mean0.98+-0.21, (range 3.14-0.13), Epirius-2.44-3.21
Tscachalidis et al. (2009); European ungulate management (2010)
3.21 Tsachalidis et al. and Hadjisterkotis (2009)
Hungary
2.91 in game forest(with arable included)
Palotas et al. (1996) 2.91 Csanyi and Lehoczki (2010)
Ireland N/A N/A N/A N/A
Italy 3-18; 3.9-23.6; 3.6;
Mattioli (1995); Massei et al. (1998); Focardi (2002); 23.91
Kept in Italy due to Torina et al. (2008) referring to Boar on Sicily. Apollonio et al. (2010)
Latvia 0.75;0.263Sumilo et al. (2008); European ungulate management (2010)
0.75 Balciauskas et al. (2010)
Lithuania 0.45, 0.3-0.38;0.199
Sumilo et al. (2008); Balciauskas (2009); European ungulate management (2010)
0.45 Balciauskas et al. (2010)
Macedonia (FYR) 0.108796296 Melovski (2008) 1.09
Netherlands 2; 0.055 Kuiters (2005); European ungulate management (2010) 2
Van Wierene and Groot Bruinderink (2010)
Norway ? ? ? ?
Poland0.18-5.9;, 2.09-6.78; up to 17.9
Jedrzejewska et al. (1994), Jedredjewski et al. (2002); European ungulate management (2010); Gulaet al. (2006)
6.78
Portugal at least 7.6; 0.0868
Braga (2010); European ungulate management (2010) 7.6 Vingada et al. (2010)
Romania 10-20; 9-74; 0.237, 0.5-0.07
Cristescu et al. (2007); European ungulate management (2010) Murariu
20.1
16
(2008);Slovakia 0.587 Find’o and Skuban (2010) 0.59
Slovenia min=0.36/35.67
European ungulate management (2010)/Stergar M., pers com,2011
2.9 Stergar M., pers. comm. (2011)
Spain 0.5-48 Martin-Atance et al. (2006); Mendoza et al. (2006) 10 Carranza J. (2010)
Sweden
60000=1.33; forsmark 0.01;0.26 simpevarp and 0.12 Blankaholm;0.09
Annual survey,2003/4; European ungulate management (2010); Cederlund et al. (2004); Tannerfeldt et al. (2004); Rosvold and Andersen (2008);
1.33 Libery et al. (2010)
U.K. ? ? ? ?
17
Table C10. Data used for TB prevalence in wild boar across EuropeWild boar TB prevalence
Country TB prevalence (%) References Max Prevalence
usedMode Prevalence used
Austria 0.0001 0.00005Belgium 0.0001 0.00005Bulgaria 1.75 Bachvarova et al. (1995) 0.0175 0.0001Croatia 0.0001 0.00005Czech Republic 0.0001 0.00005Denmark 0.0001 0.00005Estonia 0.0001 0.00005Finland 0.0001 0.00005
France 29-42, 31.7 Zanella et al. (2008), EFSA (2008) 0.42 0.29
Germany 0.0001 0.00005Greece 0.0001 0.00005Hungary 0.0001 0.00005Ireland N/A N/A 0.0001 0.00005
Italy3.3 (1991-1992), 4.25-27, 23.8
Serraino (1999); EFSA (2008); 0.27 0.033
Latvia 0.0001 0.00005Lithuania 0.0001 0.00005Macedonia (FYR) 0.0001 0.00005
Netherlands 0.0001 0.00005Norway ? ? 0.0001 0.00005Poland 0.0001 0.00005
Portugal 10.8(6.9-16.4);34& 39
Direccao De Services Veterinaros De Regiao Alentejo (2010); Santos et al. (2010)
0.39 0.32
Romania 0.0001 0.00005Slovakia 0.0001 0.00005Slovenia 0.0001 0.00005
Spain 0-100, 23.95, 1.1-7.7
Hermoso de Mendoza et al. (2006); Vicente et al. (2007); EFSA (2008);
1 0.24
Sweden 0.0001 0.00005U.K. ? ? 0.0001 0.00005
18
Table C11. Data used for TB prevalence and density of cattle across Europe
Countries Regions of countries Cattle TB prevalence (%) Cattle Density
Belgium Antwerp 0.0224 110.2797Limburg 0.0224 55.9238East Flanders 0.0224 108.7525Flemish Brabant 0.0224 51.4720West Flanders 0.0224 130.9788Walloon Brabant 0.0224 51.7841Hainaut 0.0224 88.7105Liège 0.0224 72.2425Luxembourg 0.0224 83.3671Namur 0.0224 71.5066
Bulgaria Vidin 0.0000 4.6138Vratsa 0.0000 4.6138Montana 0.0000 4.6138Lovech 0.0000 4.6138Pleven 0.0000 4.6138Ruse 0.0000 6.0373Veliko Tarnov 0.0000 6.0373Gabrovo 0.0000 6.0373Razgrad 0.0000 6.0373Targovishte 0.0000 6.0373Silistra 0.0000 6.3215Dobrich 0.0000 6.3215Varna 0.0000 6.3215Shumen 0.0000 6.3215Burgas 0.0000 4.8486Yambol 0.0000 4.8486Sliven 0.0000 4.8486Stara Zagora 0.0000 4.8486Sofia 0.0000 3.6101Grad Sofiya 0.0000 3.6101Kyustendil 0.0000 3.6101Pernik 0.0000 3.6101Blagoevgrad 0.0000 3.6101Plovdiv 0.0000 7.6954Kardzhali 0.0000 7.6954Haskovo 0.0000 7.6954Pazardzhik 0.0000 7.6954Smolyan 0.0000 7.6954
Czech Republic Praha 0.0000 2.0161
Stredoceský 0.0000 13.3908Jihozápad 0.0000 20.2759Plzenský 0.0000 20.2759Karlovarský 0.0000 8.6600Ústecký 0.0000 8.6600Liberecký 0.0000 21.2701Pardubický 0.0000 21.2701Královéhradecký 0.0000 21.2701Jihoceský 0.0000 19.5854Vysocina 0.0000 19.5854Olomoucký 0.0000 16.6721
19
Zlínský 0.0000 16.6721Moravskoslezský 0.0000 14.8871
Denmark Frederiksborg 0.0000 12.1046Bornholm 0.0000 12.1046Copenhagen 0.0000 12.1046Storstrøm 0.0000 11.9621Vestsjælland 0.0000 11.9621Roskilde 0.0000 11.9621Vejle 0.0000 50.3650Ribe 0.0000 50.3650Fyn 0.0000 50.3650South Jutland 0.0000 50.3650Viborg 0.0000 38.7308Ringkøbing 0.0000 38.7308Århus 0.0000 38.7308North Jutland 0.0000 47.9374
Germany Baden-Württemberg 0.2687 29.3186Bayern 0.2687 48.5982Berlin 0.2687 0.6726Brandenburg 0.2687 19.9803Bremen 0.2687 26.7327Hamburg 0.2687 8.6093Hessen 0.2687 23.3436Mecklenburg-Vorpommern 0.2687 24.3874Niedersachsen 0.2687 54.0358Nordrhein-Westfalen 0.2687 41.6987Rheinland-Pfalz 0.2687 19.7250Saarland 0.2687 21.0977Sachsen 0.2687 27.3784Sachsen-Anhalt 0.2687 17.1304Schleswig-Holstein 0.2687 74.7895Thüringen 0.2687 21.7104Estonia 0.0000 5.1959
Ireland Donegal 0.4939 70.2899Mayo 0.4939 70.2899Leitrim 0.4939 70.2899Sligo 0.4939 70.2899Monaghan 0.4939 70.2899Cavan 0.4939 70.2899Roscommon 0.4939 70.2899Galway 0.4939 70.2899Louth 0.4939 70.2899Longford 0.4939 70.2899Westmeath 0.4939 70.2899Offaly 0.4939 70.2899Laoighis 0.4939 70.2899Meath 0.4939 98.4408Wicklow 0.4939 98.4408Kerry 0.4939 98.4408Limerick 0.4939 98.4408Tipperary 0.4939 98.4408Kilkenny 0.4939 98.4408Carlow 0.4939 98.4408Cork 0.4939 98.4408
20
Wexford 0.4939 98.4408Waterford 0.4939 98.4408Kildare 0.4939 98.4408Clare 0.4939 98.4408Dublin 0.4939 98.4408
Greece Anatoliki Makedonia kai T 0.7139 7.5087Kentriki Makedonia 0.7139 9.6008Dytiki Makedonia 0.7139 4.7826Thessalia 0.7139 7.3235Stereá Elláda 0.7139 10.4894Ipeiros 0.7139 6.0741Ionioi Nisoi 0.7139 2.7308Dytiki Ellada 0.7139 5.3100Peloponnisos 0.7139 1.0717Attiki 0.7139 1.1029Voreio Aigaio 0.7139 1.8918Notio Aigaio 0.7139 5.3180Kriti 0.7139 0.2422
Spain Galicia 0.0257 32.7923Principado de Asturias 0.0435 37.2690Cantabria 0.4511 53.1103País Vasco 0.0342 20.9842Comunidad Foral de Navarr 0.3015 10.3455La Rioja 0.1171 6.9574Aragón 0.1852 5.7336Comunidad de Madrid 0.5908 13.9761Castilla y León 0.4380 12.7357Castilla-la Mancha 1.9560 5.8122Extremadura 0.4670 16.9501Cataluña 0.0754 17.1794Comunidad Valenciana 0.2484 2.3694Islas Baleares 0.0000 5.2484Andalucía 1.2496 6.8547Región de Murcia 0.1945 5.0473Islas Canarias 0.4977 2.3768
France Île-de-France 0.0195 2.4975Champagne-Ardenne 0.0195 23.7444Picardie 0.0195 27.3210Haute-Normandie 0.0195 50.7429Basse-Normandie 0.0195 93.2969Bourgogne 0.0195 41.2577Nord-Pas-de-Calais 0.0195 54.3829Lorraine 0.0195 40.4298Alsace 0.0195 19.8068Franche-Comté 0.0195 37.2176Pays de la Loire 0.0195 81.0112Bretagne 0.0195 74.4607Poitou-Charentes 0.0195 30.0283Aquitaine 0.0195 18.4948Midi-Pyrénées 0.0195 27.8292Limousin 0.0195 65.6357Rhône-Alpes 0.0195 22.9301Auvergne 0.0195 60.4698Languedoc-Roussillon 0.0195 8.8980
21
Provence-Alpes-Côte-d'Azu 0.0195 2.1338Corse 0.0195 7.8341
Italy Piemonte 0.0948 31.2965Valle d'Aosta 0.2056 12.5345Liguria 0.2985 3.2835Lombardia 0.0000 69.6366Provincia Autonoma Bolzano-Bozen 0.0000 20.3649
Trentino-Alto Adige 0.0561 3.7701Veneto 0.0000 50.5573Friuli-Venezia Giulia 0.0557 13.2018Emilia-Romagna 0.0008 28.0645Toscana 0.0038 4.5876Umbria 0.0000 7.3084Marche 0.1668 7.4995Lazio 0.1432 14.3314Abruzzo 0.0022 7.6617Molise 0.0023 12.7084Campania 0.1103 15.4321Apulia 0.1968 8.3153Basilicata 0.1116 8.0464Calabria 0.4300 7.1021Sicily 1.1038 11.5178Sardegna 0.0640 10.4234Cyprus 0.0000 6.0121Latvia 0.0000 5.8555Lithuania 0.0000 759.4000Luxembourg (Grand-Duché) 0.0000 75.8970
Hungary Budapest 0.0000 8.2382Pest 0.0155 8.2382Komárom-Esztergom 0.0251 7.5643Fejér 0.0219 7.5643Veszprém 0.0000 7.5643Gyor-Moson-Sopron 0.0000 9.2783Vas 0.0000 9.2783Zala 0.0000 9.2783Tolna 0.0000 5.2932Baranya 0.0041 5.2932Somogy 0.0084 5.2932Borsod-Abaúj-Zemplén 0.0000 4.9896Heves 0.0000 4.9896Nógrád 0.0000 4.9896Szabolcs-Szatmár-Bereg 0.0000 9.5780Hajdú-Bihar 0.0000 9.5780Jász-Nagykun-Szolnok 0.0000 9.5780Bács-Kiskun 0.0019 7.7431Csongrád 0.0000 7.7431Békés 0.0000 7.7431Malta 0.0000 51.5823
Netherlands Groningen 0.0004 84.1563Friesland 0.0004 163.0337Drenthe 0.0004 84.4058Overijssel 0.0004 192.0649Gelderland 0.0004 183.3166
22
Flevoland 0.0004 44.3975Utrecht 0.0004 147.9076Noord-Holland 0.0004 59.5506Zuid-Holland 0.0004 68.8432Zeeland 0.0004 29.0828Noord-Brabant 0.0004 134.9868Limburg 0.0004 67.3479
Austria Burgenland 0.1038 5.5472Niederösterreich 0.1038 23.8344Wien 0.1038 0.2528Kärnten 0.1038 20.8684Steiermark 0.1038 20.9492Oberösterreich 0.1038 48.9816Salzburg 0.1038 23.2457Tirol 0.1038 14.6743Vorarlberg 0.1038 25.3749
Poland Lódz 0.0042 26.2309Masovian 0.0360 27.6422Lesser Poland 0.0175 12.4106Silesian 0.0158 9.8015Lublin 0.0008 14.7007Subcarpathian 0.0000 6.3259Swietokrzyskie 0.0457 14.1278Podlachian 0.0000 45.8050Greater Poland 0.0050 27.3188West Pomeranian 0.0062 3.9823Lubusz 0.0000 4.2047Lower Silesian 0.0104 4.9581Opole 0.0000 12.9091Kuyavian-Pomeranian 0.0000 26.6170Warmian-Masurian 0.0026 17.0606Pomeranian 0.0033 9.4135
Portugal Viana do Castelo 0.0340 15.1753Braga 0.0340 15.1753Vila Real 0.0340 15.1753Bragança 0.0340 15.1753Porto 0.0340 15.1753Faro 0.0340 1.6445Aveiro 0.0340 7.8941Viseu 0.0340 7.8941Guarda 0.0340 7.8941Coimbra 0.0340 7.8941Castelo Branco 0.0340 7.8941Leiria 0.0340 7.8941Lisboa 0.0340 16.9358Beja 0.0340 18.0470Santarém 0.0340 18.0470Portalegre 0.0340 18.0470Évora 0.0340 18.0470Setúbal 0.0340 18.0470Região Autónoma dos Açores (PT) 0.0000 104.4757Região Autónoma da Madeira (PT) 0.0000 6.3670
Romania Maramures 0.0034 11.8563Satu Mare 0.0034 11.8563
23
Bistrita-Nasaud 0.0034 11.8563Cluj 0.0034 11.8563Bihor 0.0034 11.8563Salaj 0.0034 11.8563Alba 0.0034 10.5718Harghita 0.0034 10.5718Mures 0.0034 10.5718Sibiu 0.0034 10.5718Brasov 0.0034 10.5718Covasna 0.0034 10.5718Bacau 0.0034 17.6662Botosani 0.0034 17.6662Suceava 0.0034 17.6662Iasi 0.0034 17.6662Neamt 0.0034 17.6662Vaslui 0.0034 17.6662Constanta 0.0034 8.1679Galati 0.0034 8.1679Vrancea 0.0034 8.1679Buzau 0.0034 8.1679Braila 0.0034 8.1679Tulcea 0.0034 8.1679Ialomita 0.0034 9.9191Calarasi 0.0034 9.9191Dâmbovita 0.0034 9.9191Arges 0.0034 9.9191Prahova 0.0034 9.9191Giurgiu 0.0034 9.9191Teleorman 0.0034 9.9191Bucuresti - Ilfov 0.0034 9.5003Vâlcea 0.0034 8.8799Gorj 0.0034 8.8799Mehedinti 0.0034 8.8799Olt 0.0034 8.8799Dolj 0.0034 8.8799Timis 0.0034 5.7731Arad 0.0034 5.7731Hunedoara 0.0034 5.7731Caras-Severin 0.0034 5.7731
Slovenija Koroška 0.0000 26.0563Pomurska 0.0000 26.0563Podravska 0.0000 26.0563Savinjska 0.0000 26.0563Zasavska 0.0000 26.0563Spodnjeposavska 0.0000 26.0563Jugovzhodna Slovenija 0.0000 26.0563Notranjsko-kraška 0.0000 26.0563Obalno-kraška 0.0000 18.8190Goriška 0.0000 18.8190Osrednjeslovenska 0.0000 18.8190Gorenjska 0.0000 18.8190
Slovakia Bratislavský 0.0000 36.4130Trenciansky 0.0000 34.3521Trnavský 0.0000 34.3521
24
Žilinský 0.0000 34.3521Nitriansky 0.0000 7.8634Banskobystrický 0.0000 7.8634Košický 0.0000 19.9936Prešov 0.0000 19.9936
Finland Eastern Finland 0.0000 4.0366Southern Finland 0.0000 2.6219Western Finland 0.0000 3.5567Lapland 0.0000 1.3997Oulu 0.0000 1.3997Åland 0.0000 4.7036
Sweden Stockholm 0.0000 3.0826Uppsala 0.0000 6.6479Orebro 0.0000 6.6479Södermanland 0.0000 6.6479Västra Götaland 0.0000 6.6479Östergötland 0.0000 6.6479Kronoberg 0.0000 11.8096Jönköping 0.0000 11.8096Gotland 0.0000 11.8096Kalmar 0.0000 11.8096Blekinge 0.0000 16.3660Skåne 0.0000 16.3660Västra Götaland 0.0000 11.5820Halland 0.0000 11.5820Gävleborg 0.0000 1.9010Dalarna 0.0000 1.9010Värmland 0.0000 1.9010Jämtland 0.0000 0.8521Västernorrland 0.0000 0.8521Norrbotten 0.0000 0.3979Västerbotten 0.0000 0.3979
UK: England Darlington 0.0051 31.5410Stockton-on-Tees 0.0051 31.5410Redcar and Cleveland 0.0051 31.5410Durham 0.0074 31.5410Tyne and Wear 0.0000 31.5410Northumberland 0.0000 31.5410Cheshire 0.0990 67.2785Merseyside 0.0000 67.2785Halton 0.0990 67.2785Warrington 0.0000 67.2785Manchester 0.0000 67.2785Lancashire 0.0033 67.2785Cumbria 0.0095 67.2785North East Lincolnshire 0.0080 36.5759South Yorkshire 0.0278 36.5759North Lincolnshire 0.0080 36.5759North Yorkshire 0.0051 36.5759West Yorkshire 0.0085 36.5759East Riding of Yorkshire 0.0327 36.5759Northamptonshire 0.0714 38.2959Rutland 0.0714 38.2959Derbyshire 0.1078 38.2959
25
Leicestershire 0.0299 38.2959Nottinghamshire 0.1037 38.2959Lincolnshire 0.0080 38.2959Staffordshire 0.2153 39.4351West Midlands 0.0163 39.4351Warwickshire 0.2082 39.4351Herefordshire 0.4634 39.4351Worcestershire 0.4634 39.4351Shropshire 0.2035 39.4351Hertfordshire 0.0000 10.7741Essex 0.0000 10.7741Milton Keynes 0.0000 10.7741Norfolk 0.0866 10.7741Bedfordshire 0.0000 10.7741Suffolk 0.0000 10.7741Cambridgeshire 0.0000 10.7741Harrow 0.0000 0.0000Berkshire 0.0233 23.1986Westminster 0.0000 0.0000Enfield 0.0000 23.1986Merton 0.0000 0.0000Hackney 0.0000 0.0000Havering 0.0000 23.1986Kent 0.0000 23.1986Hampshire 0.0304 23.1986Richmond upon Thames 0.0000 0.0000Bromley 0.0000 0.0000Surrey 0.0000 23.1986Sutton 0.0000 0.0000Croydon 0.0000 0.0000West Sussex 0.1346 23.1986Southampton 0.0304 23.1986East Sussex 0.0386 23.1986Poole 0.0926 23.1986Portsmouth 0.0000 23.1986Isle of Wight 0.0000 23.1986Oxfordshire 0.2096 23.1986Buckinghamshire 0.0414 23.1986Gloucestershire 0.4674 74.5310Bristol 0.0000 0.0000South Gloucestershire 0.4674 74.5310Wiltshire 0.2519 74.5310North Somerset 0.1605 74.5310Bath and North East Somer 0.1605 74.5310Devon 0.2501 74.5310Somerset 0.1605 74.5310Cornwall 0.2461 74.5310Dorset 0.0926 74.5310
UK: Wales Anglesey 0.0046 53.7562Conwy 0.0046 53.7562Denbighshire 0.0545 53.7562Flintshire 0.0545 53.7562Gwynedd 0.0046 53.7562Wrexham 0.0545 53.7562
26
Powys 0.1958 53.7562Ceredigion 0.2951 53.7562Pembrokeshire 0.2951 53.7562Carmarthenshire 0.2951 53.7562Monmouthshire 0.2188 53.7562Swansea 0.2514 53.7562Neath Port Talbot 0.2514 53.7562Rhondda, Cynon, Taff 0.0291 53.7562Merthyr Tydfil 0.0291 53.7562Blaenau Gwent 0.2188 53.7562Torfaen 0.2188 53.7562Caerphilly 0.2188 53.7562Newport 0.2188 53.7562Bridgend 0.0291 53.7562Vale of Glamorgan 0.0000 53.7562Cardiff 0.0000 53.7562
UK: Scotland Shetland Islands 0.0000 22.4064Orkney Islands 0.0000 22.4064Eilean Siar 0.0000 22.4064Highland 0.0172 22.4064Moray 0.0000 22.4064Aberdeenshire 0.0116 22.4064Argyll and Bute 0.0250 22.4064Perthshire and Kinross 0.0000 22.4064Angus 0.0616 22.4064Stirling 0.0000 22.4064Fife 0.0000 22.4064West Dunbartonshire 0.0000 22.4064Inverclyde 0.0250 22.4064North Ayshire 0.0101 22.4064East Dunbartonshire 0.0000 22.4064Falkirk 0.0000 22.4064East Lothian 0.0000 22.4064Renfrewshire 0.0310 22.4064Glasgow 0.0000 22.4064North Lanarkshire 0.0000 22.4064West Lothian 0.0000 22.4064Edinburgh 0.0000 22.4064East Ayrshire 0.0101 22.4064East Renfrewshire 0.0310 22.4064South Lanarkshire 0.0000 22.4064Midlothian 0.0000 22.4064Scottish Borders 0.0081 22.4064South Ayrshire 0.0101 22.4064Dumfries and Galloway 0.0046 22.4064
UK: Northern Ireland 0.0619 112.2589
27
References for supplementary tables
Aaris-Sorensen, J., 1995. Road-kills of badgers (Meles meles) in Denmark. Annales. Zoologici.
Fennici 32, 31-36.
Andersen, R., Holthe, V., 2010. Ungulates and their management in Denmark. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 71-85.
Andersen, R., Lund, E., Solberg, E.J. Saether, B.E., 2010. Ungulates and their management in
Norway. In M. Appollonio, R. Andersen and R. Putman Eds. European Ungulates and their
management in the 21st Century, Cambridge: Cambridge University Press, pp 14-36.
Apollonio, M., Di Vittorio I., 2004. Feeding and reproductive behaviour in fallow bucks (Dama
dama). Naturwissenschaften 91, 597-584.
Apollonio, M., Ciuti, S., Pedrotti, L., Banti, P., 2010. Ungulates and their management in Italy. In M.
Appollonio, R. Andersen and R. Putman Eds. European Ungulates and their management in the 21st
Century, Cambridge: Cambridge University Press, pp 507-526.
Aragon, S., Braza, F., San Jose, C., 1995., Socioeconomic, physiognomic, and climatic factors
determining the distribution pattern of roe deer Capreolus capreolus in Spain. Acta Theriologica 40,
37-43.
Bachvarova, Y., Libkov, B., Karov, R., 1996. Tuberculosis in wild boar. In Proceedings of the 22nd
congress of the international union of game biologists: the game and man, N. Botev Ed. Pensoft
Publishers, Sofia, pp 492-494.
Balciauskas, L., Kawata, Y., 2009. Estimation of carrying capacity and growth rate of wolf in
Lithuania. Acta Zoologica Litnanica 19, 79-83.
28
Balciauskas, L., Ozolms, J., Randveer, T., Tonisson, J., 2010. Ungulates and their management in the
Baltics (Estonia, Latvia and Lithuania). In M. Appollonio, R. Andersen and R. Putman Eds. European
Ungulates and their management in the 21st Century, Cambridge: Cambridge University Press, pp
103-128.
Balseiro, A., Oleaga, A., Orusa, R., Robetto, S., Zoppi, S., Dondo, A., Goriaa, M., Gortazar, C., Garcia
Marin, J.F., Domenis, L., 2009. Tuberculosis in Roe deer from Spain and Italy. Veterinary Record
164, 468-470.
Biro, Z., Szemthy, L., Katona, K., Heltai, M., Peto, Z., 2006. Seasonal distribution of red deer (Cervus
elaphus) in forest-agricultural habitat in Hungary. Mammalia 70, 70-75.
Bobek, B., Boyce, M.S., Kosobucka, M., 1984. Factors affecting red deer (Cervus elaphus) population
density in Southeastern Poland. Journal of Applied Ecology 21, 881-890.
Borger, L., Franconi, N., Ferretti, F., Meschi, F., De Michele, G., Gantz, A., Coulson, T., 2006. An
integrated approach to identify spatiotemporal and individual-level determinants of animal home range
size. American Naturalist 168, 417-485.
Borkowski, J., Pudelko, M., 2007. Forest habitat use and home range size in radio-collared fallow
deer. Annales of Zoologici Fennicci 44, 107-114.
Braga, C., Alexandre, N., Fernandez-Llario, P., Santos P., 2010. Wild boar (Sus scrofa) harvesting
using the espera hunting method: side effects and management implications. European Journal of
Wildlife Research 56, 465-469.
British deer society. www.bds.org.uk, 05/10/2010.
29
Broseth, H., Bevanger, K., Knutsen, B, 1997. Function of multiple badger Meles meles setts:
distribution and utilisation. Wildlife Biology 3, 89-96.
Bugalho, M.N., Milne, J.A., 2003. The composition of the diet of red deer (Cervus elaphus) in a
Mediterranean environment: a case of summer nutritional constraint? Forest Ecology and Management
181, 23-29.
Carranza, J., 2010. Ungulates and their management in Spain. In M. Appollonio, R. Andersen and R.
Putman Eds. European Ungulates and their management in the 21st Century, Cambridge: Cambridge
University Press, pp 419-440.
Casear, J., Licoppe, A., 2010. Ungulates and their management in Belgium. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 184-200.
Cederlund, G., Hammarstrom, A., Wallin, K., 2004 Surveys of mammal populations in the areas
adjacent to Forsmark and Okarshamn: results from 2003. In Svensk Karnbranslehantering AB (SKB),
Stockholm, pp 1-43.
Csivincsik, A., Janosi, S.Z., Szabo, J., Nagy, G., Nemes, C.S., Nagy, J., Sugar, L., Bogdan, T., Tuboly,
S., Lovey, L., 2010. The possible role of red deer (Cervus elaphus) in the maintenance of bovine
tuberculosis in Zselic region of Hungary. Enclosures: a dead-end? InInfluence on game biology,
conservation and hunting symposium proceedings, Hungary.
Chapman, N., Claydon, K., Claydon, M., Forde, P.G., Harris, S., 1993. Sympatric populations of
muntjac (Muntiacus reevesi) and Roe deer (Capreolus capreolus): a comparative analysis of their
ranging behaviour, social aggregation and activity. Journal of Zoology 229, 623-640.
30
Clifton-Hadley, R.S., Wilesmith, J.W., Stuart F.A., 1993. Mycobacterium bovis in the European
badger (Meles meles): epidemiological findings in the tuberculous badgers from a naturally infected
population. Epidemiology and Infection 111, 9-19.
Clutton-Brock, T.H., Guiness, F.E., Albon, S.D., Red deer: wildlife behaviour and ecology series.
University of Chicago press, Chicago, 1st edit 1982.
Courteney, O., Reilly, L.A., Sweeney, F.P., Hibberd, V., Bryan, S., U-Hassan, A., Newman, C.,
MacDonald, D.W., Delahay, R.J., Wilson, G.J., Wellington, E.M.H., 2006. Is Mycobacterium bovis in
the environment important for the persistence of bovine tuberculosis? Biology Letters 2, 460-462.
Cristescu, B., Ion, I., 2007. Density of ungulates at preferential feeding sites in two hunting grounds of
Vrancea county (Romania), with comparison between two study methods. Analele Stiintifice ale
Universitatii Iasi: Biologie Animala 53, 245-250.
Csanyi, S., Lehoczki, R., 2010. Ungulates and their management in Hungary. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 291-318.
Dardaillon, M., 1986. Seasonal variations in habitat selection and spatial distribution of wild boar (Sus
scrofa) in the Camargue, Southern France. Behavioural Processes 13, 251-268.
Delahay, R.J., Cheeseman, C.L., Clifton-Hadley, R.S., 2001. Wildlife disease reservoirs: the
epidemiology of Mycobacterium bovis infection in the European badger (Meles meles) and other
British mammals. Tuberculosis 81, 43-49.
Delahay, R.J., Walker, N.J., Forrester, G.J., Harmsen, B., Riordan, P., MacDonald, D.W., Newman,
C., Cheeseman, C.L., 2006. Demographic correlates of bite wounding in Eurasian badgers, Meles
meles L., in stable and perturbed populations. Animal Behaviour 71, 1047-1055.
31
Delahay, R.J., Smith, G.C., Barlow, A.M., Walker, N.J., Harris, A., Clifton-Hadley, R.S., Cheeseman,
C.L., 2007. Bovine tuberculosis infection in wild mammals in the Southwest region of England: A
survey of prevalence and a semi-quantitative assessment of the relative risks to cattle. Veterinary
Journal 173, 287-301.
Direccao General De Veterinaria 2010. Programma de erradicacao da tuberculose bovina, 1-32.
Direccao De Services Veterinaros De Regiao Alentejo (2010) Controlo tuberculose bovina caso regiao
Moura-Barrancos, 1-17.
Ecologie Nature, www.ecologie.nature.free.fr/pages/mammiferes, 27/10/2010.
EFSA (European Food Safety Authority Reports),www.EFSA.europa.eu.en,
2004;2005;2006;2007;2008.
Eurostat. www.epp.eurostat.ec.europa.eu/portal/food/data/database, 02/07/2010.
Find’o S., Skuban, M., 2010. Ungulates and their management in Slovakia. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 262-290.
Focardi, S., Isotti, R., Tinelli, A., 2002. Line transect estimates of ungulate populations in a
Mediterranean forest. Journal of Wildlife Management 66, 48-58.
Focardi, S., Aragno, P., Montanaro, P., Riga, F., 2006. Inter-specific competition from fallow deer
(Dama dama) reduces habitat quality for the Italian roe deer Capreolus capreolus italicus. Ecography
29, 407-417.
32
Genov P., pers com, 2010.
Giannakopoulos, A., Tsaparis et al. , D., Birtsas, P., Poirazidis, Iliopoulos, Y., Manios, N., 2009.
Current distribution and population status of roe deer in Greece. In 9 th European roe deer meeting,
Edinburgh, 2009.
Gortazar, C., Jose Torres, M., Vicente, J., Acevedo, P., Reglero, M., De La Fuente, J., Jose Negro, J.,
Aznar-Martin, J., 2008. Bovine Tuberculosis in Donana Biosphere reserve: The role of wild ungulates
as disease reservoirs in the last Iberian Lynx strongholds. PLoS ONE 3, e2776.
Gula, R., 2006. Wolf predation on domestic animals in the Polish Carpathian mountains. Journal of
Wildlife Management 72, 283-289.
Hemami, M-R, Watkinson, A.R., Dolman, P.M., 2005. Population densities and habitat associations of
introduced muntjac Munitacus reevesi and native roe deer Capreolus capreolus in a lowland pine
forest. Forest Ecology and Management 215, 234-238.
Hermoso de Mendoza, J., Parra, A., Tato, A., Alonso, J.M., Rey, J.M., Pena, J., Garcia-Sanchez, A.,
Larrasa, J., Teixido, J., Manzano, G., Cerrato, R., Pereira, G., Fernandez-Llario, P., Hermoso de
Mendoza, M., 2006. Bovine tuberculosis in wild boar (Sus scrofa), red deer (Cervus elaphus) and
cattle (Bos taurus) in a Mediterranean ecosystem (1992-2004). Preventative Veterinary Medicine, 74,
239-247.
Homolka, M., Heroldova, M., 2003. Impact of large herbivores on mountain forest stands in the
Beskydy Mountains. Forest Ecology and Management 181,119-129.
Irish deer society, www.irishdeersociety.ie, 03/11/2011.
33
Jedrzejewska, B., Okarma, H., Jedrzejewski, W., Milkowski, L., 1994. Effects of exploitation and
protection on forest structure, ungulate density and wolf predation in Bialowieza Primeval Forest,
Poland. Journal of Applied Ecology 31, 664-676.
Jedrzejewski, W., Sauf, J., Schmidt, K., Theurkauf, J., Jedrzejewska, B., Selva, N., Zub, K., Szymura,
L., 2002. Kill rates and Predation by wolves on ungulate populations in Bialowieza Primeaval Forest
(Poland). Ecology 83, 1341-1356.
Kauhala, K., 1995 Changes in distribution of the European badger Meles meles in Finland during the
rapid colonization of the racoon dog. Annales Zoologici Fenninci 32, 183-191.
Kauhala, K., Holmala, K., 2006 Contact rate and risk of rabies spread between medium-sized
carnivores in Southeast Finland. Annales Zoologici Fennici 43, 348-357.
Keuling, O., Stier ,N., Roth, M., 2008. Annual and seasonal space use of different age classes of
female wild boar Sus scofa L. European Journal of Wildlife Research 54, 403-412.
Keuling, O., Greiser, G., Grauer, A., Strauss, E., Bartel-Steinbach, M., Klein, R., Wenzelides, L.,
Winter, A., 2010. The German wildlife information system (WILD): population densities and den use
of red foxes (Vulpes vulpes) and badgers (Meles meles) during 2003-2007 in Germany. European
Journal of Wildlife Research 57, 95-105.
Kiraly, I., Egri, B., 2007. Epidemiological characteristics of Cephenemyia stimulator (Clark, 1815)
larval infestation in European roe deer (Capreolus capreolus) in Hungary. Acta Zoologica Academiae
Scientiarum Hungaricae 53, 271-279.
Kjellender, P., Hewison, A.J.M., Liberg, O., Angibault, J-M., Bideau, E., Cargnelutti, B., 2004.
Experimental evidence for density-dependence of home-range size in roe deer (Capreolus capreolus
L.): a comparison of two long-term studies. Oecologia 139, 478-485.
34
Kowalczyk, R., Zalewski, A., Jedrzejewska, B., Jedrzejewski, W., 2003. Spatial organisation and
demography of badgers (Meles meles) in Bialowieza primeval Forest, Poland and the influence of
earthworms on badger densities in Europe. Canadian Journal of Zoology 81, 74-87.
Kozak, L., Heltai, M., 2006. Distribution, habitat selection and facial morphology of setts of badgers
(Meles meles Linnaeus, 1758) in the region of Erdospusztak (Eastern Hungary). Analele Universitatii
din Oradea, Fascicula Biologie 13, 48-51.
Kuiters, A.T., Groot Bruinderink, G.W.T.A., Lammertsma, D.R., 2005. Facilitative and competitive
interactions between sympatric cattle, red deer and wild boar in Dutch woodland pastures. Acta
Theriologica 50, 241-252.
Kusak, J., Krapinec, K., 2010. Ungulates and their management in Croatia. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 527-539.
Libery, O., Bergstrom, R., Kindway, J., Von Essen, H., 2010 Ungulates and their management in
Sweden. In M. Appollonio, R. Andersen and R. Putman Eds. European Ungulates and their
management in the 21st Century, Cambridge: Cambridge University Press, pp 37-70.
Licoppe, A.M., De Crombrugghe, S.A., 2003 Assessment of spring habitat selection of red deer
(Cervus elaphus L.) based on census data. Z. Jagdwiss 49, 1-13.
Malo, J.E., Jimenez, B., Suarez, F., 2000. Herbivore dunging and endozoochorous seed deposition in a
Mediterranean dehesa. Journal of Range Management 53, 322-328.
Marco, I., Ruiz, M., Juste, R., Garrido, J.M., Lavin, S., 2002. Paratuberculosis in free-ranging Fallow
deer in Spain. Journal of Wildlife Disease 38, 629-632.
35
Martin-Atance, P., Leon-Vizcaino, L., Palomares, F., Revilla, E., Gonzalez-Candela, M., Calzada, J.,
Cuero-Pablo, M.J., Delibes, M., 2006. Antibodies to Mycobacterium bovis in wild carnivores from
Donana national park (Spain). Journal of Wildlife Disease 42, 704-708.
Massei, G., Bacon, P., Genov P.V., 1998. Fallow deer and Wild boar pellet group disappearance in a
Mediterranean area. Journal of Wildlife Management 62, 1086-1094.
Mateos-Quesada, P., 2005. Densidad poblacional y uso del espacio del corzo en el centro de la
peninsula iberica. Galemys 17, 3-12.
Mattioloi, S., Meneguz, P.G., Brugnoli, A., Nicoloso S., 2001, Red deer in Italy: recent changes in
range and numbers. Hystrix Italian Journal of Mammology 12, 27-35.
Matyastik, T., Bicik, V., 1999. Distribution and habitat selection of badger (Meles meles) in Northern
Moravia. Acta Universitatis Palackianae olomucensis facultas rerum naturalium: Biologica 37, 77-88.
McElligott, A.G., Altwegg, R., Hayden, T.J., 2002 Age-specific survival and reproductive
probabilities: evidence for senescence in male fallow deer (Dama dama). Proceedings of the Royal
Soceity of London B 269, 1129-1137.
Melovski, D., Ivanov, G., Stojanov, A., Trajce, A., Zimmermann, F., Von Arx, M., 2008. First
camera-trap survey in the national park of Mavrovo, Macedonia. International conference on
Biological and environmental sciences. University of Tirana: Faculty of natural sciences, 312-315.
Micu, I., Nahlik, A., Negus, S., Mihalache, I., Szabo, I., 2010. Ungulates and their management in
Romania. In M. Appollonio, R. Andersen and R. Putman Eds. European Ungulates and their
management in the 21st Century, Cambridge: Cambridge University Press, pp 319-337.
36
Muldowney, J., Curry, J.P., O’Keefe, J., Schmidt, O., 2003. Relationships between earthworm
populations,grassland management and badger densities in County Killkenny, Ireland. Pedobiologia
47, 913-919.
Murariu, D., 2003. Mammals (Mammalia) from the southern area of Piatra Craiuli national park
(Romania). Travaux du museum national d’Histoire Naturelle 45, 381-393.
Murariu, D., 2008. Faunology, biology, ecology and protection statutge of the mammals (Mammalia)
of the Macin mountains national park (Romania). Travaux du museum national d’Histoire Naturelle
51, 273-301.
Office National de la Chase et de la Faune Sauvage 2010. Tableaux de chasse ongules sauvages saison
2008-2009. ONCFS 287.
Olea-Popelka, F.J., Griffin, J.M., Collins, J.D., McGrath, G., Martin, S.W., 2003. Bovine tuberculosis
in badgers in four areas in Ireland: does tuberculosis cluster? Preventive Veterinary Medicine 59, 103-
111.
Olea-Popelka, F.J., Phelan, J., White, P.W., McGrath, G., Collins, J.D., O’Keefe, J., Duggan, M.,
Collins, D.M., Kelton, D.F., Berke, O., More, S.J., Martin, S.W., 2006. Quantifying badger exposure
and the risk of bovine tuberculosis for cattle herds in county Kilkenny, Ireland. Preventive Veterinary
Medicine 75, 34-46.
Palotas et al. , G., Bartucz, P., Kovacs, G., 1996. The world famous fallow deer (Dama dama)
population and its habitat. In Proceedings of the 22nd congress of the international union of game
biologists: the game and man, N. Botev Ed. Pensoft Publishers, Sofia, pp 155.
37
Palotas, G., Vadasz, I., Gal, B., 1996. Population structure and dynamics of roe deer (Capreolus
capreolus) in the Great Hungarian plains. In Proceedings of the 22nd congress of the international
union of game biologists: the game and man, N. Botev Ed. Pensoft Publishers, Sofia, pp 156.
Palphramand, K.L., Newton-Cross, G., White, P.C.L., 2007. Spatial organisation and behaviour of
badgers (Meles meles) in a moderate-density population. Behavioural Ecological Sociobiology 61,
401-413.
Papageorgiou, N.K. 1978. Food preferences, feed intake and protein requirements of red deer in
Central Greece. Journal of Wildlife Management 42, 940-943.
Papaioannou, H., 2010. Ungulates and their management in Greece. In M. Appollonio, R. Andersen
and R. Putman Eds. European Ungulates and their management in the 21st Century, Cambridge:
Cambridge University Press, pp 540-562.
Pays, O., Benhamou, S., Helder, R., Gerard, J-F., 2007. The dynamics of group formation in large
mammalian herbivores: an analysis in the European roe deer. Animal Behaviour 74, 1429-1441.
Pichon, B., Mousson, L., Figureau, C., Rodhain, F., Perez-Eid 1999. Density of deer in relation to the
prevalence of Borrelia burgdorferi L. in Ixodes ricinus nymphs in Rambouillet forest, France.
Experimental and Applied Acarology 23, 267-275.
Prokesova, J., Barancekova, M., Homolka, M., 2006 Density of red and roe deer and their distribution
in relation to different habitat characteristics in a floodplain forest. Folia Zoologica 55, 1-14.
Quigley, F.C., Costello, E., Flynn, O., Gogarty, A., McGuirk, J., Murphy, A., Egan, J., 1997. Isolation
of mycobacteria from the lymph node lesions in deer. Veterinary Record 141, 516-518.
38
Radeloff, V.C., Pidgeon, A., Hostert, P., 1999. Habitat and population modelling of roe deer using an
interactive geographic information system. Ecological Modelling 144, 287-304.
Ramanzin, M., Sturaro, E., Zanon, D., 2007. Seasonal migration and home range of roe deer
(Capreolus capreolus) in the Italian eastern Alps. Canadian Journal of Zoology 85, 280-289.
Reimoser, F., Reimoser, S., 2010. Ungulates and their management in Austria. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 103-128.
Remonti, L., Balestrieri, A., Prigioni, C., 2006. Range of the Eurasian badger (Meles meles) in an
agricultural area of northern Italy. Ethology, Ecology and Evolution 18, 61-7.
Revilla, E., Palomares, F., 2002. Spatial organisation, group living and ecological correlates in low-
density populations of Eurasian badgers, Meles meles. Journal of Animal Ecology 71, 497-512.
Richard, E., Gaillard, J-M, Said, S., Hamann, J-L, Klein, F., 2010. High red deer density depresses
body mass of roe deer fawns. Oceologica 163, 91-97.
Richomme, C., Boschiroli, M.L., Hars, J., Casablanca, F., Ducrot, C., 2010. Bovine tuberculosis in
Livestock and wild boar on the Mediterranean island, Corsica. Journal of Wildlife Diseases 46, 627-
631.
Rosalino, L.M., MacDonald, D.W., Santos-Reis, M., 2004. Spatial structure and land-cover use in a
low density Mediterranean population of Eurasian badgers. Canadian Journal of Zoology 82, 1493-
1502.
Rosvold, J., Andersen, R., 2008. Wild boar in Norway – is climate a limiting factor? NTU
Vitenkapsmuseet. Rapp. Zool. Ser. 1, 1-23
39
Ruusila, V., Kojola, I., 2010. Ungulates and their management in Finland. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 86-102.
Saez-Royez and Tellaria 1991. In Acevedo, P. Delibes-Mateos, M., Escudero, M.A., Vicente, J.,
Marco, J., Gortazar C., 2005. Environmental constraints in the colonization sequence of the roe deer
(Capreolus capreolus Linnaeus, 1758) across the Iberian mountains, Spain. Journal of Biogeography
32, 1671-1680.
Santin-Duran, M., Alunda, J.M., Hoberg, E.P., De La Fuente, C., 2004. Abomasal parasites in wild
sympatric cervids, red deer, Cervus elaphus and fallow deer, Dama dama, from three localities across
central and western Spain: relationship to host density and park management. Journal of Parasitology
90, 1378-1386.
Santos, N., Geraldes, M., Afonso, A., Almeida, V.. Correia-Neves 2010. Diagnosis of tuberculosis in
the wild boar (Sus scrofa): A comparison of methods applicable to hunter-harvested animals. PLoS
ONE 5, e12663.
Schettler, E., Steinbach, F., Eschenbacher-Kaps, I., Gert, K., Meussdoerffer, F., Risch, K., Jurgen
Streich, W., Frolich, K., 2006. Surveillance for prion disease in cervids, Germany. Emerging
Infectious Diseases 12, 319-322.
Schmidt, K.T., Hoi, H., 2002. Supplemental feeding reduced natural selection in juvenile red deer.
Ecography 25, 265-272.
Serraino A., Marchetti G., Sanguinetti V., Rossi M. C., Zanoni R. G., Catozzi L., Bandera A., Dini W.,
Mignone W., Franzetti F. and Gori A. (1999) Monitoring the transmission of tuberculosis between
40
wild boar and cattle: Genotypical analysis of strines my molecular epidemiology techniques. Journal
of Clinical Microbiology, 37 (9), 2766-2771.
Spiridonov G. and Spassov N. (www.worldwildlife.org/bsp/publications/europe/bulgaria/bulgaria17,
13/10/2010, sub ref Grigorov, 1986)
Sumilo, D., Bormane, A., Asokliene, L., Vasilenko, V., Golovljova, I., Avsic-Zupanc, T., Hubalek, Z.,
Randolph, S.E. 2008. Socio-economic factors in the differential upsurge of tick-borne encephalitis in
Central and Eastern Europe. Reviews in Medical Virology 18, 81-95.
Tannerfeldt, M., Thiel, L., Miljobyra, L., 2004. Okarshamn site investigation: Survey of mammal
populations at Simpevarp, spotlight survey 2004. In Svensk Karnbranslehantering AB (SKB),
Stockholm, pp 1-21.
Torina, A., Alongi, A., Naranjo, V., Scimeca, S., Nicosia, S., Di Marco, V., Caracappa, S., Kocan,
K.M., De La Fuente, J., 2008. Characterisation of Anaplasma infections in Sicily, Italy. Animal
Biodiversity and Emerging Diseases 1149, 90-93.
Trdan, S., Vidrih, M., 2008. Quantifying the damage of red deer (Cervus elaphus) grazing on
grassland production in Southeastern Slovenia. European Journal of Wildlife Management 54, 138-
141.
Tsachalidis, E.P., Hadjisterkotis, E., 2009. Current distribution and population status of wild boar (Sus
scrofa L.) in Greece. Acta Silvatica Lignaria. Hungaria 5, 153-157.
Tsaparis, D., Katsanevakis, S., Stamouli, C., Legakis, A., 2008. Estimation of roe deer Capreolus
capreolus and mouflon Ovis aries densities, abundance and habitat use in a mountainous
Mediterranean area. Acta Theriologica 53, 87
41
Van Wierene, S.E., Groot Bruinderink, G.W.T.A., 2010. Ungulates and their management in the
Netherlands. In European Ungulates and their management in the 21st Century, Eds. M. Appollonio,
R. Andersen and R. Putman, Cambridge: Cambridge University Press, pp 165-183.
Vicente, J.P., Bideau, E., Hewison, A.J.M., Angibault, J.M., 1995. The influence of increasing density
on body weight, kidd production, home range and winter grouping in roe deer (Capreolus capreolus).
Journal of Zoology 236, 371-382.
Vicente, J., Hofle, U., Garrido, J.M., Fernandez de Mera, I.G., Acevedo, P., Juste, R., Barral, M.,
Gortazar, C., 2007. Risk factors associated with the prevalence of tuberculosis-like lesions in fenced
wild boar and red deer in south central Spain. Veterinary Research 38, 451-464.
Vingada, J., Fonseca, C., Cancela, J, Ferreira, J., Eira, C., 2010. Ungulates and their management in
Portugal. In M. Appollonio, R. Andersen and R. Putman Eds. European Ungulates and their
management in the 21st Century, Cambridge: Cambridge University Press, pp 392-418.
Ward, A. I., 2005. Expanding ranges of wild and feral deer in Great Britain. Mammal Review, 35,
165-173.
Wikenros, C.S. H., Wabakken, P., Liberg, O., Pedersen, H.C., 2009. Wolf predation on moose and roe
deer: chase distances and outcome of encounters. Acta Theriologica 54, 207-218.
Worschikowsky, U., 2010 Ungulates and their management in Germany. In M. Appollonio, R.
Andersen and R. Putman Eds. European Ungulates and their management in the 21st Century,
Cambridge: Cambridge University Press, pp 201-222.
Zanella, G., Durand, B., Hars, J., Moutou, F., Garin-Bastuji, B., Duvauchelle, A., Ferme, M., Karoui,
C., Boschiroli, M. L., 2008. Mycobacterium bovis in wildlife in France. Journal of Wildlife Diseases
44, 99-108
42
Zanella, G., Durand, B., Hars, J., Moutou, F., Garin-Bastuji, B., Duvauchelle, A., Ferme, M., Karoui,
C., Boschiroli, M. L., 2008. Premier foyer de tuberculose a M. bovis dans une population de cerfs et de
sangliers sauvages en France. Bulletin Epidemiologique 29, 1-5
43
APPENDIX D
Table of land class categories for species within the study
Table D. Categorisation of relative species densities in different habitats, relative to the maximum density
value from the literature (see Appendix Tables A3, A5, A7, A9, A11 and A12 for details for each species).
Category 5, ≥ 90% of maximum density value; category 4, 70-89%; category 3, 50-69%; category 2, 30-
49%; category 1, 10-29%; category 0, <10%.
Habitats\Species Badgers (M. meles)
Fallow Deer (D. dama)
Red Deer (C. elaphus)
Roe Deer (C. capreolus)
Wild Boar (S. scrofa)
Cow (B. taurus)
Agro-forestry areas 5 5 5 5 5 0Annual crops associated with permanent crops
1 3 2 3 3 0
Broad-leaved forest 5 5 5 5 5 0Complex cultivation patterns
3 3 2 3 3 0
Coniferous forest 4 2 4 4 4 0Fruit trees and berry plantations
3 2 1 2 3 0
Green urban areas 3 0 0 1 0 0Inland marshes 1 1 1 1 1 0Land principally occupied by agriculture, with significant areas of natural vegetation
5 5 5 5 5 0
Mixed forest 5 5 5 5 5 0Moors and heathland 1 1 3 1 1 0Natural grasslands 4 4 4 4 4 5Non-irrigated arable land 3 3 2 3 3 0Olive groves 3 2 1 2 2 0Pasture 5 5 3 2 3 5Peat bogs 1 1 3 1 1 0Permanently irrigated land
3 3 2 3 3 0
Sclerophyllous vegetation
3 2 3 2 3 5
Transitional woodland-shrub
5 5 5 4 4 0
Vineyards 2 2 2 2 2 0
44
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