what is the modern synthesis? how do we define evolution? · • evolutionary species concept:...

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• What is the modern synthesis?

• How do we define evolution?

Modern Synthesis

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Tuesday, March 1, 2011

Questions

• What are the four forces of evolution?

• How does each change gene frequencies within and between populations?

• What is a population?

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The four forces• What is genetic drift?

• When is genetic drift most effective?

• Why is mutation so important?

• What is the role of gene flow in maintaining species?

• What are the different ways in which Natural selection works?

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Genetic Drift

• The random factor

• Greatest effect in small populations

• Founder effect

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Genetic Drift - Bottleneck

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Genetic bottleneck

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Natural selection

• differential reproductive success over multiple generations

• some variations are more successful than others, leading to a change in the entire population over time

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Natural Selection

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Pepper Moths

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Four Forces and Populations

Within Between

Mutation

Genetic Drift

Natural Selection

Gene Flow

=

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MICROEVOLUTION = the small changes in gene frequencies in a population from generation to

generation

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MACROEVOLUTION = the cumulative effect of these small changes over a long period of time - may lead to

SPECIATION

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What is a species?What is speciation?

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What is a species?

• an interbreeding group of animals or plants that are reproductively isolated though anatomy, ecology, behavior, or geographic distribution from all other such groups

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Reproductive Isolating Mechanisms

• Premating RIMsHabitat isolationTemporal isolationBehavioral isolationMechanical incompatibility

• Postmating RIMsSperm-egg incompatibilityZygote inviabilityEmbryonic or fetal inviability

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Theridon grallator

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Pan

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Common Chimpanzee Bonobo

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Gorilla

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Mountain Eastern lowland Western lowland

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X

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X =

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Species Concepts• Biological species concept: Defines species as

interbreeding populations reproductively isolated from other such populations.

• Evolutionary species concept: Defines species as evolutionary lineages with their own unique identity.

• Ecological species concept: Defines species based on the uniqueness of their ecological niche.

• Recognition species concept: Defines species based on unique traits or behaviors that allow members of one species to identify each other for mating.

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Modes of Evolutionary Change

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Cladogenesis

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Sympatric speciation

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Speciation creates clades

• clades are evolutionarily related groups

• Classification is the naming of these groups

• Started with Linnaeus

• tried to group organisms together based on relationship

• based on similarity

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Tempo of Speciation?• Gradualism?

• Punctuated Equilibrium?

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