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    Phylum: Chordates

    Subphylum: Urochordata andCephalochordata

    AA & SSPeriod 1

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    http://www.bio.miami.edu/dana/160/chordatephylogeny.gif

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    Chordates

    Chordates are organismsthat possess: A structure called a

    notochord at somepart of theirdevelopment.

    A dorsal, hollow nervecord

    Pharyngeal slits Muscular post-analtail.

    http://instruct1.cit.cornell.edu/courses/biog105/labs/deuts/chordates.html

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    Chordates

    Notochord a rod that extends most of thelength of the body that provides support duringlocomotion.

    Nerve cord- dorsal hollow nerve cord that isunique to chordates animals from other phylahave solid nerve chords. The nerve cord of a chordate is the central

    nervous system: brain and spinal cord.

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    Chordates

    Pharyngeal slits allow water to exit from themouth without going through the entiredigestive tract. Slits have become modified for

    gas exchange, jaw support, hearing, and otherfunctions during vertebrate evolution. Chordates have a tail. Usually, non-chordates

    have a digestive tract that extends the wholelength of the body. The chordate tail hasmuscles, and skeletal elements and providespropulsion in water.

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    Invertebrates

    Invertebrates are animals that lack a backbone. Invertebrates include 98% of the animals on

    Earth.

    Invertebrates usually have a shell or a hardexoskeleton for protection

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    Invertebrate Chordates

    The adults of Urochordates have neither anotochord nor a dorsal tubular nervous systemSo how can they be chordates?

    These animals disperse themselves with free-swimming larvae that have a dorsal tubular nervous system

    notochord gill slits

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    Invertebrate Chordates

    Chordates include two subphyla of invertebrates:-UROCHORDATES -CEPHALACHORDATES

    https://reader009.{domain}/reader009/html5/0426/5ae100297aced/5ae1002ea3073.jpg http://www-biol.paisley.ac.uk/courses/Tatner/biomedia/pictures/amphi8.htm

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    Invertebrate Chordates Tunicate larva - free-

    swimming and exhibits allchordate characteristics: notochord, a dorsal

    nerve cord, pharyngealslits, and a post-analtail.

    The larvae swims until iteventually attaches to ahard substrate it loses its tail and

    ability to move, and itsnervous system

    disintegrates.

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    Urochordates

    Commonly called tunicates Most are marine animals Some adhere to rocks while others are

    planktonic. Seawater enters through an incurrent passes

    through the pharyngeal slits into atrium and exits

    through an excurrent siphon (Atriopore). Thefood that is filtered from this water is passed by cilia into the intestine. The anus empties into theexcurrent siphon.

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    Urochordates

    Have a notochord thatextends from just behind the head tothe tail

    Urochordata means"tail- cord

    There are three

    classes within thissubphylum: Ascidiacea, Thaliacea,and Larvacea.

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    Class Ascidiacea

    Adult ascidians arecommonly called "seasquirts

    Ascidiacea areinhabitants of theintertidal zone they may be either

    solitary or colonial. All are filter feeders.

    http://upload.wikimedia.org/wikipedia/commons/1/16/Ascidias.jpg

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    Class Thaliacea

    barrel-shaped Known as "salps".

    They are surrounded by circular muscle bands and both endsof their cylindrical body are open. Contractions of the circular muscles make the body pulsate;

    This creates a form of jet propulsion and a mechanism forfilter-feeding (suspended food is removed from the water by the pharyngeal gill slits).

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    Class Larvacea

    Larvacians resemble ascidian larvae Most specialized urochordates. Build a home that is mobile within which they

    travel through the course of their lives. The walls of their mobile homes give little

    protection.

    Their pharyngeal gill slits are used for feeding.

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    Urochordata

    There are roughly 1,600 species of urochordates; most

    are small solitary animals but some arecolonial, organisms. Tunicates Sea Squirts

    http://www.dscc.edu/bwilliams/Biology2/bio2animcont.htm

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    Physiology

    No coelom - has been replaced by proliferationof connective tissue

    Complex nervous system

    Locomotion propulsion Reproduction:

    Most are hermaphrodites, producing both

    eggs and sperm hat are either free-spawned or brooded. Most species are self-sterile, butsome are self- fertile

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    Physiology

    Urochordates have: dual nervous system - different nerves

    segmented (somatic nerves) unsegmented

    (visceral nerves) regions; segmented muscles. locomotion by contraction of alternate sides

    as in vertebrates

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    Physiology Brain is functions similarly to the vertebrate brain segmented muscle (myomeres) pull alternate sides

    (stiffened by notochord.) Pharynx & gills used only for filter feeding. Circulation limited, but has a ventral pump & major

    dorsal artery. No hemoglobin; oxygen carried in solution; no kidney, but similar excretory cells. No connection between segmented & unsegmented

    nerves. Semi-sedentary filter feeder Both Cephelochordata and

    Urochordata trap detritus in mucus; water moved by cilia mouth and gut have little or no musculature.

    No paired fins therefore less coordination.

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    Anatomy Urochordata

    http://bio.classes.ucsc.edu/bio136/urochordata/urochordata.html

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    Cephalochordata Derived from Greek

    meaning both endspointed

    The Cephalochordatasubphylum commonly includes vertebrates,tetrapods, and amniotes

    Lampreys and hagfish arethe only agnathans(jawless fish) that are notextinct today

    They are commonly known as lancelets

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    Lampreys (Cephalaspidomorphi )

    Feed upon fish bloodfrom the flank

    Born in fresh water

    streams and remainthere until they reachmaturity in lakes oropen sea

    Larva (ammocoete)are scavengers sincetheir mouths are notdeveloped for rasping

    http://upload.wikimedia.org/wikipedia/en/thumb/e/e5/Lamprey_illustration_side.png/800px-Lamprey_illustration_side.png

    http://io.uwinnipeg.ca/~simmons/16cm05/1116/34-09a-Lamprey.jpg

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    Hagfish (Myxini) Hagfish secrete slime,

    then tie themselves in aknot to scrape it off, toclean themselves

    Pair of horizontalstructures that haveprojections that pick upfood

    Feed on alive/dead fishsinsides

    Tie themselves in a knotand slide towards themouth, the knot pullingthe mouth off http://newport.pmel.noaa.gov/heceta/source_files/source_jpegs/logbook_images/r609hagfi

    sh_link.jpg

    http://lemonodor.com/scruz-2003-01-24/med/130-lo-hagfish.jpg

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    Anatomy of Cephalochordata

    1= brain-like blister 2= notochord

    3= dorsal nerve cord 4= post anal tail 6= food canal 10= gill slit

    http://upload.wikimedia.org/wikipedia/commons/thumb/b/b1/Lancetnikinside.png/800px-Lancetnikinside.png

    http://upload.wikimedia.org/wikipedia/commons/b/b1/Lancetnikinside.png
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    Physiology I

    Chordates generally have bilateralsymmetry that is differentiated into head,trunk, and tail

    All have notochord, nerve cord, visceralclefts and arches (gills)

    Notochord: rod-like structure found incephalochordata embryos; gives axialsupport. Made from mesoderm cells.

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    Physiology II

    Nerve cord: develops above the notochordas a hollow tube. Differentiates brainanteriority and spinal cord from the trunk to the tail. Makes up central nerve system.

    Visceral clefts & arches (gills): appears aspouches that push out to make clefts,connecting the pharynx to the exterior.Site of major gas exchange.

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    Quiz

    Of which of the following features does acephalochordata not possess?

    I notochord

    II jawsIII nerve cord A) I only

    B) II only C) I and IID) I and IIIE) I, II, and III

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    Quiz

    Which classes are not extinct in jawless vertebrates today?

    A) Myxini and CephalaspidomorphiB) LanceletsC) AgnathaD) Cephalaspidomorphi and MammaliaE) Amphibia and Reptilia

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    Quiz

    How do the adults of Urochordates andCephelochordates belong to the groupchordates?

    What are the three classes within the subphylum

    Urochordata?

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    QUIZ

    These animals disperse themselves with free-swimming larvae that have a dorsal tubular nervous system

    notochord gill slits

    There are three classes within this subphylum: Ascidiacea, Thaliacea, and Larvacea.