beach seine catch data from secheltinlet and agamemnon … · except for a few beaches and...
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Beach Seine Catch Data from Sechelt Inletand Agamemnon Channel, British Columbia
1+1
C.D. Levings, I.K. Birtwell, and G.E. Piercey
Fisheries and Oceans CanadaScience Branch - Pacific RegionWest Vancouver Laboratory4160 Marine DriveWest Vancouver, British Columbia V7V lN6
2003
Canadian Data Report ofFisheries and Aquatic Sciences 1110
Fisheries and Oceans Peches et OceansCanada , Canada Canada
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Canadian Data Report of
Fisheries and Aquatic Sciences 1110
2003
BEACH SEINE CATCH DATA FROMSECHELT INLET AND AGAMEMNON CHANNEL, BRITISH COLUMBIA
by
C.D. Levings, LK. Birtwell, and G.E. Piercey
Fisheries and Oceans CanadaScience Branch, Pacific Region
West Vancouver Laboratory4160 Marine Drive
West Vancouver, B.C. V7V IN6
© Her Majesty the Queen in Right of Canada, 2003,as represented by the Minister of Fisheries & Oceans
Cat. No. Fs 97-13/11lOE ISSN 0706-6465
Correct citation for this publication:
Levings, C.D., I.K. Birtwell, and G.B. Piercey. 2003. Beach seine catch data fromSechelt Inlet and Agamemnon Channel, Blitish Columbia. Can. Data Rep. Fish.Aquat. Sci. 1110: iii + 23 p.
11
ABSTRACT
Levings, C.D., I.K. BiltweIl, and G.E. Piercey. 2003. Beach seine catch data fromSechelt Inlet and Agamemnon Channel, Blitish Columbia. Can. Data Rep. Fish.Aquat. Sci. 1110: iii + 23 p.
Data are presented from beach seine surveys in May, June, and July 1987 at
Agamemnon Channel and Sechelt Inlet on the east side of the Strait of Georgia. Chum
salmon fry dominated catches of salmonids except for one location, where large catches
of coho smolts that had escaped from a salmon net pen operation were obtained. Chum
fry in Agamemnon Channel were larger relative to those from Sechelt Inlet and were
likely fish migrating to sea after a period of growth in the southern Strait. Thirty-seven
taxa of fish were caught in the surveys. Habitat observations at each of the sample sites
are also given.
RESUME
Levings, C.D., I.K. BirtweIl, and G.E. Piercey. 2003. Beach seine catch data fromSechelt Inlet and Agamemnon Channel, British Columbia. Can. Data Rep. Fish.Aquat. Sci. 1110: iii + 23 p.
Les donnees presentees rendent compte de releves effectues ala senne de plage en
mai, juin et juillet 1987 dans Ie chenal Agamemnon et dans Ie bras Sechelt, sur la rive
orientale du detroit de Georgia. Les alevins de ketas dominaient les prises de salmonides,
sauf aun endroit, ou on a capture une grande quantite de smolts de cohos echappes d'une
salmoniculture. Les alevins de ketas du chenal Agamemnon etaient plus gros que ceux du
bras Sechelt, et il s'agissait vraisemblablement de poissons prets amigrer vers Ie large
apres une periode de croissance dans Ie sud du detroit. Nous avons capture 37 taxons de
poissons pendant les releves. Nous presentons aussi des observations sur l'habitat achacun des lieux d'echantillonnage.
111
INTRODUCTION
The migratory routes and rearing habitats ofjuvenile salmon from the Strait ofGeorgia and adjacent inlets are poorly known but are important data for habitat managersand coastal planners. In this report we present data from nearshore surveys conducted intwo contrasting coastal seaways on the east side of the Strait, an area that has not beeninvestigated by researchers. The work was conducted in May, June, and July 1987.
Agamemnon Channel is an open, northeast tending channel connecting MalaspinaStrait with the mouth of Jervis Inlet. This channel affords northward migrating juvenilesalmon from the Fraser River (e.g. chum salmon fry, Oncorhynchus keta) a route from theestuary to the northern end of the Strait and hence to sea (Levings and Tompkins 1985).Sechlet Inlet, in contrast, is an enclosed fjord with a narrow entrance (SkookchumkukNarrows, 2 km wide) - only local populations of salmonids are the likely users of theinlet.
Our studies were also stimulated by a concern, at the time, that during theirmigration and rearing phases juvenile salmon would be preyed upon by farmed fish (cohoand chinook, 0. kisutch and 0. tshawytscha respectively). In 1986 there were 18approved salmon farm locations on Sechelt Inlet and seven approved locations onAgamemnon Channel, with numerous other applications pending (Keller and Leslie 1996,p.29). Due to logistics and time constraints, we only made a few observations on thistopic; in particular we documented catches of escaped coho salmon from a net penoperation.
METHODS
SAMPLING STATIONS
Stations were selected on the basis of substrate, slope, ease of access andproximity to operating and planned fish farms. Except for a few beaches and mini-deltasat the mouths of creeks, most of the shorelines of the Agamemnon Channel and SecheltInlet are steep and rocky, so that beach seining was possible at only a few locations.Table 1 and Figure 1 give descriptions and locations of the sample stations.
SAMPLING PROCEDURES
A 14.7 beach seine, with wings 4.9 m (l em stretch mesh), bunt 4.9 m (3 mmmesh), and depth 1.5 m, was used in the sampling, deployed by boat or by wading,depending on water depth at the particular locations. Stations on Sechelt Inlet weresampled in all three months. Agamemnon Channel was only sampled in May and Julybecause weather conditions prevented sampling during the June trip. All fish specieswere identified, enumerated, and released, except for the June trip to Sechelt Inlet. On
that trip only salmonids were enumerated. During the May trip to Sechelt Inlet, surfacetemperature and salinity were measured with a Beckman RS-5 inductive salinometer.
Subsamples of chum salmon fry at a few locations were collected and preserved in10% formalin for length measurement and stomach analyses. A total of 91 chum salmonfry were retained. On May 25, the stomach contents of two coho (est.l.5-0.8 kg) salmonand three chinook salmon (est. 0.5 kg) from an aquaculture operation near Station 3 wereanalysed. The fish were sampled from the rearing pen with a dipnet and contents wereexamined in the field.
LABORATORY METHODS
Chum fry were measured to the nearest millimetre. Stomachs were removed, gutcontents weighed, and organisms identified using a Wild M5 Stereomicroscope.
DATA TABULATIONS
Table 2 provides variable descriptions and codes used in the report. A descriptionof species names and codes for fish caught are given in Table 3.
Stomach analysis data are given in Table 6 with an associated list of organismcodes in Table 5.
COMMENTS ON RESULTS
1. FISH CATCHES
Fish catch data are tabulated in Table 4. The majority of the salmonids caughtwere chum salmon fry except at Station 3 (Agamemnon Channel) on May 20 where about1400 coho salmon (0 kistuch) were caught. These were escapees from an adjacent netpen rearing operation and ranged up about 4 kg in weight. A few chinook salmon fry andsmolts (0 tshawytscha) and sockeye smolts (0 nerka) were also taken, with the latterfrom Agamemnon Channel only. Cutthroat trout (0 clarkii clarkii) were only caught inSechelt Inlet (Stations 9 and 12).
2. CHUM SALMON LENGTH FREQUENCY
Length frequency data for chum salmon fry are shown in Figure 2. More largechum salmon fry (> 90 mm) were caught in Agamemnon Channel relative to SecheltInlet. This suggests the former fish originated from rivers in the southern Strait that hadgrown since leaving their natal streams.
2
3. STOMACH CONTENTS
None of the coho salmon or chinook salmon examined from the net pen containedmacroscopic organisms. Commercial fish food pellets was the only matelial observed inthe stomachs.
Stomach content data for chum salmon fry showed they had been feeding on a widevariety of invertebrates. Twenty-five taxa were observed in the stomachs (Table 6,7).
ACKNOWLEDGEMENTS
Thanks are owing to Bruce Hillaby, Mel Kotyk, Rebecca Stewart, and BarryLawley for their help during the fieldwork. Rebecca Stewart conducted the laboratorystudies and Gaye Sihin assisted with data compilation and report preparation.
REFERENCES
Keller, B.c. and R.M. Leslie. 1996. Sea Silver. Inside British Columbia's SalmonFarming Industry. Horsdal and Schubart Publishers Ltd, VictOIia, BC 138 p.
Levings, C.D. and A. Tompkins. 1985. Habitats of Fraser River chum - an overview. p.162-167. In B.G. Shepherd (Rapporteur). Proc. 1985 Northeast Pacific Pink andChum Salmon Workshop. Dept. of Fish. and Oceans, 1090 West Pender Street,Vancouver, B.C. 212 p.
3
Table 1. Sampling station coordinates and descriptions.
Stn. Lat. Long. Geographic Description Habitat Comments
4945.901 N Nile Pt. Northwest side Steep shore, boulders
12400.428 W Oysters present
2 4945.927 N Nile Pt, southeast side Sand/gravel beach
12400.228 W
3 4945.157 N Agamemnon Bay Gravel beach, creek mouth,
12359.352 W oysters present. Fish farm
and hatchery.
4 4945.967 N West side of Agamemnon Channel Gravel beach, boulders in low
12400.874 W intertidal, small creek mouth.
5 4945.074 N West of Nile Pt on Agamemnon Channel Rocky beach, boulders in low
12402.252 W intertidal, small creek mouth.
6 4945.200 N Immediately west of ferry slip at Earls Cove Rocky beach
12400.735 W
7 4945.724 N Southwest side of Ag.amemnon Channel, Very steep shore
12401.309 W Immediately upstream of power lines
8 4938.122 N Salmon Inlet, Kunachin Bay Gravel beach, small creek mouth
12347.442 W
9 4937.969 N Salmon Inlet, Gravel beach, small creek mouth
12347.831 W Kunachin Bay at Marine Park sign
10 4936.374 N Salmon Inlet, Sand/gravel beach, gentle slope
12346.775 W Bible Camp immediately east of Nine Mile Pt
11 4936.114 N Sechelt Inlet, west side at Halfway Islets Eelgrass bed
12349.522 W
12 4932.664 N Sechelt Inlet, mouth of Carlson Creek Gravel beach, rockweed (Fucus
12347.790 W sp.); some cobble with barnacles;
sedges at high intertidal
13 4933.038 N Sechelt Inlet, Behind Tuwanek Islets, Gravel beach, boulders in subtidal
12345.925 W mouth of Irvine Creek
14 4930.569 N Sechelt Inlet, mouth of Angus Creek Sand beach
12345.289 W
15 4928.894 N End of Sechelt Inlet, Porpoise Bay Mud/sand beach, sedge
12345.156 W
4
Table 2. Descriptions of the variables and codes in Tables 3 and 4.Variable Name Variable Description Code
DATE
STATION
SET
TEMP
SAL
TIME
SPECIES
STAGE
TOTAL
Sampling date(yyyymmdd)
Sampling Sites
Sequential set per site
Surface Temperature(OC)Data not available
Surface Salinity (psu)Data not available
Sampling Time (PDT)Data not available notused
Species (defined inTables)No Catch (NC)Data not available
Life history stageSmoltFryJuvenileStage not noted
Catch by species andstageIndicates numberestimated
5
o
SMFRJU
SNN
*
Table 3. Scientific and common names and species codes of fishes.
Family/Genus Scientific Name Common Name Code
CLUPEIDAE Clupea harengus pallasi Pacific Herring PAHE
SALMONIDAE Salmonidae Salmon Unidentified to species SALMOncorhynchus keta Chum Salmon CHUMO. nerka Sockeye Salmon SOCKO. tshawytscha Chinook Salmon CHINO. kisutch Coho Salmon COHOO. clarki clarki Cutthroat Trout CUTT
AMMODYTIDAE Ammodytes hexapterus Pacific Sand Lance PASL
GASTEROSTEIDAE Gasterosteus aculeatus Threespine Stickleback THSK
SYNGNATHIDAE Syngnathus griseolineatus Bay Pipefish BAPI
TRICHODONTIDAE Trichodon trichodon Pacific Sand Fish PASF
EMBIOTOCIDAE Embiotocidae Surfperch Unidentified to species EMBIEmbiotoca lateralis Striped Perch STPERhacochilus vacca Pile Perch PIPEBrachyistius frenatus Kelp Perch KEPECymatogaster aggregata Shiner Perch SHPE
GOBIIDAE Gobiidae Goby Unidentified to species GOBIClevelandia ios Arrow Goby ARGO
STICHAEIDAE Lumpenus sagitta Pacific Snake Prickleback SNPR
PHOLIDAE Pholidae Gunnel Unidentified to species PHOLApodichthys flavidus Penpoint Gunnel PEGUPho/is ornata Saddleback Gunnel SAGU
SCORPAENIDAE Sebastes sp. Rockfish Unidentified to species SCORSebastes maliger Quillback Rockfish QURO
HEXAGRAMMIDAE Hexagrammidae Unidentified to species HEXAHexagrammos decagrammus Kelp Greenling KEGRHexagrammos stelleri Whitespotted Greenling WHGROphiodon e/ongatus Lingcod LING
COTTIDAE Cottidae Unidentified to species COTTOligocottus maculosus Tidepool Sculpin TISCEnophrys bison Buffalo Sculpin BUSCLeptocottus armatus Pacific Staghorn Sculpin STSC
6
Table 3 continued.
Family/Genus
BOTHIDAE
PLEURONECTIDAE
Scientific Name
Citharichthys sp.
PleuronectidaePlatichthys stellatusLepidopsetta bilineataPleuronectes vetulusPleuronichthys coenosus
7
Common Name
Sanddab
Flatfish Unidentified to speciesStarry FlounderRock SoleEnglish SoleCO Sole
SpeciesCode
SADA
PLEUSTARROSOENSOCOSO
Tab
le4.
Bea
chse
ine
data
from
Aga
mem
non
Cha
nnel
and
Sech
eltI
nlet
durin
g19
87.
DA
TE
ST
AT
ION
SE
TT
EM
P(O
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0520
11
--
0925
CHU
M11
619
8705
201
1-
0925
SOC
K23
1987
0520
11
--
0925
PASF
219
8705
201
2-
-09
33CH
UM
219
8705
201
3-
-09
43CH
UM
4419
8705
201
3-
-09
43PA
SF1
1987
0520
13
--
0943
PASL
1
1987
0520
21
--
0958
CHU
M36
1987
0520
21
-09
58H
EXA
119
8705
202
1-
-09
58TI
SC24
1987
0520
22
-10
10CH
UM
4719
8705
202
210
10PA
SL2
00
1987
0520
31
-10
45C
OH
O53
Esc
aped
smol
tsfr
omad
jace
ntne
tpen
.A
llfi
sh
>es
t.0.
5kg
wei
ght.
1987
0520
31
--
1045
CHU
M2
1987
0520
32
1101
CO
HO
1466
*E
scap
edsm
olts
from
adja
cent
netp
en.
All
fish
>es
t.0.
5kg
wei
ght.
1987
0520
33
1120
NC
0
1987
0520
41
--
1150
NC
019
8705
204
2-
1156
CHU
M61
1987
0520
43
--
1205
CHU
M1
1987
0520
51
-13
15CH
INSM
119
8705
205
1-
-13
15CH
UM
6619
8705
205
.2-
-13
40CH
UM
4919
8705
205
3-
-13
46CH
UM
6919
8705
205
3-
-13
46PA
SL10
1987
0520
53
--
1346
PASF
4
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(0
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0520
53
--
1346
can
619
8705
205
414
00N
C0
1987
0520
55
13.5
-14
01TH
SK26
1987
0520
56
--
1410
CHU
M22
1987
0520
56
--
1410
SHPE
13
1987
0520
61
1424
CHU
MSM
219
8705
206
114
24EM
SI1
1987
0520
62
-14
25CH
INSM
119
8705
206
214
25CH
UM
2419
8705
206
3-
1440
CHU
M5
1987
0520
71
-14
50CH
UM
6\0
1987
0520
71
1450
EMSI
16-
1987
0520
72
--
1458
CHU
M4
1987
0520
73
1503
CHU
M1
1987
0520
73
1503
EMSI
3
1987
0521
81
13.7
16.5
0913
CHU
M14
12.4
°can
d26
.5ps
uat
2mde
pth
1987
0521
81
0913
EMSI
2019
8705
218
1-
0913
SC
aRJU
219
8705
218
2-
0925
CHU
M5
1987
0521
83
-CH
UM
8119
8705
218
3-
-PI
PE1
1987
0521
83
--
SHPE
1519
8705
218
3-
-EM
SI1
1987
0521
91
--
0945
SHPE
7419
8705
219
213
.316
.509
58SH
PE33
Tab
le4
cont
inue
d.
DA
TEST
ATI
ON
SET
TEM
PrC
)SA
LIN
ITY
(%0)
TIM
ESP
EC
IES
STA
GE
TOTA
LN
OTE
S19
8705
219
2-
-09
58BA
PI25
1987
0521
93
--
1010
SHPE
719
8705
219
3-
1010
SC
aRJU
419
8705
219
3-
-10
10BA
PI1
1987
0521
101
--
1028
SHPE
4819
8705
2110
1-
1028
CO
SO2
1987
0521
101
1028
RO
Sa
119
8705
2110
1-
1028
STSC
119
8705
2110
2-
1038
LIN
GJU
119
8705
2110
2-
-10
38SA
DA
JU1
1987
0521
102
--
1038
SHPE
619
8705
2110
2-
1038
ENSO
JU1
-19
8705
2110
3-
-10
48SH
PE3
0
1987
0521
111
-11
15CH
UM
4319
8705
2111
111
15SH
PE9
1987
0521
111
-11
15PA
HE
JU20
0*19
8705
2111
1-
1115
SC
aRJU
319
8705
2111
1-
1115
PIPE
419
8705
2111
1-
-11
15LI
NG
119
8705
2111
2-
-11
23B
USC
219
8705
2111
211
23S
CaR
119
8705
2111
211
23LI
NG
JU1
1987
0521
113
1128
SC
aRJU
319
8705
2111
313
.420
.211
28LI
NG
JU1
1987
0521
121
1239
SHPE
125*
1987
0521
121
-12
39CH
UM
319
8705
2112
1-
1239
cun
1Le
ngth
=12
cm
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(0
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0521
121
--
1239
SAD
A2
1987
0521
121
-12
39C
OSO
Adu
lt2
1987
0521
121
1239
PHO
L15
*19
8705
2112
1-
-12
39ST
SC2
1987
0521
122
14.4
21.9
1254
CHU
M1
1987
0521
122
-12
54ST
SC1
1987
0521
122
-12
54TH
SK2
1987
0521
122
-12
54PH
OL
JU20
*19
8705
2112
313
10W
HG
R2
1987
0521
123
--
1310
cun
1Le
ngth
=15
cm19
8705
2112
3-
-13
10SH
PE14
1987
0521
123
--
1310
STA
R1
1987
0521
123
1310
SAD
A1
......
......
1987
0521
131
-13
30N
C0
1987
0521
132
-13
35CH
UM
9519
8705
2113
213
35EM
SI6
1987
0521
132
--
1335
LIN
GJU
419
8705
2113
2-
-13
35SC
OR
JU3
1987
0521
133
16.8
17.8
1354
CHU
M26
1987
0521
133
16.8
17.8
1354
SCO
R1
1987
0521
133
16.8
17.8
1354
SHPE
1819
8705
2113
316
.817
.813
54PH
OL
1
1987
0521
141
--
1420
CHU
M1
1987
0521
142
-14
28ST
AR
50*
mos
tlyJU
1987
0521
14.2
-14
28ST
SC3
1987
0521
142
--
1428
CHU
M1
1987
0521
143
--
1440
CHU
M7
1987
0521
143
-14
40ST
AR
4
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(D
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0521
143
--
1440
STSC
1
1987
0521
151
--
1455
CHU
M1
1987
0521
152
--
1500
THSK
1019
8705
2115
2-
-15
00ST
AR
JU25
1987
0521
152
-15
00A
RG
O1
1987
0521
152
--
1500
STSC
JU15
0*19
8705
2115
3-
-15
08CH
UM
419
8705
2115
3-
-15
08TH
SK5
1987
0521
153
--
1508
STSC
1
1987
0611
81
-13
00N
C0
1987
0611
82
--
1305
NC
0.....
.19
8706
118
313
10CH
UM
22N
1987
0611
91
-13
40N
C0
1987
0611
92
--
1345
cun
1L
engt
h=
25cm
1987
0611
93
-13
50N
C0
1987
0611
101
1440
NC
019
8706
1110
2-
-14
45N
C0
1987
0611
103
1450
NC
0
1987
0611
111
-15
10CH
UM
719
8706
1111
2-
1520
NC
019
8706
1111
3-
-15
37CH
UM
42
1987
0611
121
--
1600
CHU
M24
1987
0611
122
--
1610
CHU
M1
1987
0611
123
--
1615
NC
0
Tab
le4
cont
inue
d.
DA
TEST
ATI
ON
SET
TEM
P(O
C)SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SST
AG
ETO
TAL
NO
TES
1987
0611
131
--
1620
NC
019
8706
1113
2-
1630
NC
019
8706
1113
3-
-16
40N
C0
1987
0611
141
1650
NC
019
8706
1114
2-
1655
CHU
M1
1987
0611
143
1700
NC
0
1987
0706
21
1355
PASL
619
8707
062
1-
1355
CO
TT2
1987
0706
22
1410
CO
TT1
1987
0706
22
1410
SALM
0.....
.19
8707
062
314
20TH
SK50
w19
8707
062
314
20EM
SIJU
3
1987
0706
11
1445
NC
019
8707
061
214
46N
C0
1987
0706
13
1455
THSK
119
8707
061
314
55CH
UM
2
1987
0706
51
-15
10CH
UM
119
8707
065
1-
1510
THSK
319
8707
065
2-
1520
NC
0R
ock
caug
htin
sein
e19
8707
065
3-
-15
25C
OTT
20*
1987
0706
53
-15
25PA
SL
1987
0706
61
1540
CHU
M3
1987
0706
62
--
1550
CHU
M37
1987
0706
62
--
1550
THSK
4
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(0
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0706
63
--
1605
CHU
M23
1987
0706
63
--
1605
THSK
3
1987
0706
71
--
1615
CHIN
1SN
N19
8707
067
1-
1615
THSK
119
8707
067
2-
1625
CO
HO
1SN
N19
8707
067
2-
1625
CHU
M17
1987
0706
73
-16
35CH
IN2
SNN
1987
0706
73
-16
35CH
UM
119
8707
067
316
35TH
SK15
1987
0706
73
--
1635
EMBI
1
1987
0706
41
1650
CO
HO
1SN
N.....
...j:
::..
1987
0706
41
-16
50SH
PE1
1987
0706
42
-16
55N
C0
1987
0706
43
-17
10PA
SL4
1987
0706
31
1725
PASL
119
8707
063
217
30N
C0
1987
0706
33
--
1740
CO
HO
FR1
1987
0707
81
-10
00SH
PE11
1987
0707
81
1000
PIPE
719
8707
078
1-
-10
00Q
UR
O2
1987
0707
81
--
1000
BAPI
219
8707
078
1-
1000
con
119
8707
078
.2-
-10
05PA
HE
419
8707
078
2-
-10
05Q
UR
O2
1987
0707
82
-10
05SH
PE2
1987
0707
82
--
1005
PIPE
1
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(O
C)S
AL
INIT
Y(%
0)T
IME
SP
EC
IES
ST
AG
ET
OT
AL
NO
TE
S
1987
0707
82
--
1005
PASL
4019
8707
078
2-
-10
05ST
PE1
1987
0707
82
-10
05SA
DA
119
8707
078
3-
-10
18C
OSO
119
8707
078
310
18PI
PE7
1987
0707
83
--
1018
BU
SC1
1987
0707
83
--
1018
BAPI
619
8707
078
3-
1018
PASL
2619
8707
078
310
18co
n4
1987
0707
83
1018
PEG
U2
1987
0707
91
--
1033
GO
BI1
1987
0707
91
--
1033
SHPE
3V
l19
8707
079
1-
1033
QU
RO
219
8707
079
110
33PI
PE1
1987
0707
92
-10
45SH
PE57
1987
0707
92
1045
PIPE
519
8707
079
2-
1045
SAG
U1
1987
0707
92
--
1045
QU
RO
219
8707
079
311
00Q
UR
O7
1987
0707
93
1100
SHPE
719
8707
079
3-
-11
00PI
PE2
1987
0707
101
--
1120
BAPI
519
8707
0710
1-
1120
SAD
A2
1987
0707
101
-11
20SH
PE26
0*19
8707
0710
.1
-11
20PI
PE1
1987
0707
101
-11
20Q
UR
O1
1987
0707
102
--
1130
SHPE
419
8707
0710
2-
-11
30BA
PI4
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(0
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0707
102
--
1130
SA
DA
419
8707
0710
2-
-11
30TH
SK1
1987
0707
103
--
1135
SAD
A13
1987
0707
103
--
1135
AR
GO
219
8707
0710
3-
1135
con
1
1987
0707
111
-12
15Q
UR
O5
1987
0707
111
-12
15SH
PE35
0*19
8707
0711
112
15PI
PE7
1987
0707
112
1220
PASL
2519
8707
0711
2-
1220
BU
SC1
1987
0707
112
-12
20Q
UR
O5
1987
0707
112
1220
PIPE
1.....
.19
8707
0711
2-
-12
20SH
PE3
0'\
1987
0707
11PA
HE
312
3535
1987
0707
113
1235
KEG
R1
1987
0707
113
-12
35SH
PE6
1987
0707
113
--
1235
PIPE
119
8707
0711
3-
1235
QU
RO
4
1987
0707
121
--
1304
QU
RO
619
8707
0712
1-
1304
SNPR
119
8707
0712
1-
1304
SHPE
135
1987
0707
121
-13
04SA
GU
319
8707
0712
113
04PI
PE7
1987
0707
121
--
1304
STSC
119
8707
0712
113
04H
EXA
119
8707
0712
1-
-13
04PE
GU
219
8707
0712
2-
-13
15CH
UM
319
8707
0712
2-
-13
15PI
PE9
Tab
le4
cont
inue
d.
DA
TE
ST
AT
ION
SE
TT
EM
P(0
C)
SA
LIN
ITY
(%0)
TIM
ES
PE
CIE
SS
TA
GE
TO
TA
LN
OT
ES
1987
0707
122
--
1315
SHPE
1519
8707
0712
2-
1315
BU
SC2
1987
0707
122
1315
SAD
A1
1987
0707
122
--
1315
SAG
U4
1987
0707
122
--
1315
PIPE
419
8707
0712
2-
-13
15ST
SC4
1987
0707
122
--
1315
AR
GO
119
8707
0712
3-
1325
AR
GO
319
8707
0712
3-
1325
SHPE
1019
8707
0712
3-
1325
con
319
8707
0712
3-
-13
25PL
EUJU
119
8707
0712
313
25SA
PI3
.......
1987
0707
123
-13
25SA
DA
1-.
...]
1987
0707
131
-13
45SH
PE8
1987
0707
131
1345
STPE
219
8707
0713
213
50SH
PE8
1987
0707
132
--
1350
PIPE
219
8707
0713
2-
-13
50ST
PE2
1987
0707
133
-14
00CH
UM
219
8707
0713
3-
-14
00ST
PE1
1987
0707
133
--
1400
SHPE
119
8707
0713
3-
-14
00SA
PI1
1987
0707
133
--
1400
con
1
Table 5. Organisms found in stomach content analysis.GROUP FAMILY/GENUS LIFE STAGE CODE
INSECTA Insect Adult INSAInsect Larvae INSLInsect Pupae INSP
COPEPODA Calanoid CALNHarpacticoid HARP
BARNACLES Cirripedia Adult CIRACyprid Larvae CIRL
CLADOCERA Cladocera CLAD-
AMPHIPODA Gammaridea GAMMHyperiidea HYPECaprellidea CAPR
DECAPODA Crab Zoea BRAZShrimp Zoea DECZ
MYSIDACEA Mysidacea MYSI
CUMACEA Cumacea CUMA
MOLLUSCA Clam Larvae BIVLBRYOZOAN Bryozoan Larvae BRYZTUNICATE Tunicata TUNIPOLYCHAETE Polychaete PLCH
MITES Hydracarina HYDRMites
FISH Fish Larvae TLST
DECAPODA Decapod Larvae DECL
MOLLUSCA Gastropod Larvae GASL
EUPHAUSIIDS Euphausiidea EUPH
EGGS Unidentifed Eggs EGGS
18
Tabl
e6.
Dat
aof
chum
stom
ach
anal
ysis
.F
ish
Site
Dat
eL
Wt
Gu
t%
II
IC
HC
CC
GH
CB
DM
CB
BT
PH
TD
GE
E(m
m)
(g)
Co
nte
nt
Gu
tN
NN
AA
II
LA
YA
RE
,yU
IR
U0
YL
EA
UG
We
igh
tW
eig
ht
SS
SL
RR
RA
MP
PA
CS
MV
YN
LD
SC
SP
G(g
)A
LP
NP
AL
DM
ER
ZZ
IA
LZ
IY
RT
LL
HS
11
Ma
y75
4.4
0.07
41.
690
00
150
00
00
00
90
00
00
00
00
06
00
21
May
723.
80.
124
3.27
40
00
00
00
20
016
00
00
00
00
00
10
0
31
Ma
y55
1.6
0.03
62.
241
00
02
00
03
00
00
00
932
020
04
00
01
00
41
May
501.
10.
041
3.69
50
00
00
00
40
00
11
00
00
00
00
00
0
51
Ma
y37
1.6
0.03
32.
094
00
08
00
025
00
00
00
00
00
00
00
00
61
May
491.
50.
040
2.69
10
05
10
00
10
01
00
00
025
00
00
200
0
71
May
501.
20.
064
5.36
30
03
120
00
10
00
00
04
460
00
00
00
0
81
May
390.
60.
049
8.20
50
04
90
00
111
00
00
00
02
00
00
00
0
91
May
540.
90.
074
8.20
120
01
180
00
620
00
00
00
00
00
00
00
010
1M
ay47
1.0
0.06
06.
001
00
33
00
048
00
00
00
00
00
00
00
00
111
May
410.
70.
048
6.81
31
02
200
00
452
00
00
00
00
00
00
00
012
4M
ay
450.
80.
043
5.35
120
00
00
00
41
00
00
00
00
00
00
00
013
4M
ay59
2.5
0.20
48.
144
00
84
00
00
00
00
00
010
000
00
00
00
0
144
May
571.
80.
037
2.04
80
01
40
00
10
00
00
00
00
00
00
00
30.....
154
Ma
y49
1.3
0.07
75.
900
00
21
00
00
00
00
00
02
120
00
00
020
0\0
164
May
491.
10.
040
3.64
41
10
90
00
90
00
00
00
00
01
00
00
2017
4M
ay
571.
80.
049
2.72
10
06
260
00
02
00
00
10
00
00
00
00
018
4M
ay
491.
30.
105
8.04
10
01
10
00
40
00
00
030
1000
00
00
00
010
019
4M
ay
481.
10.
033
3.03
80
03
40
00
10
00
00
00
03
00
00
00
3020
4M
ay
481.
20.
067
5.59
30
00
00
00
00
00
00
00
04
00
00
00
5021
4M
ay47
1.1
0.06
35.
759
00
04
00
04
00
00
00
00
60
00
00
050
224
May
450.
90.
052
5.77
193
01
220
00
250
00
00
00
00
00
00
00
023
6M
ay74
3.6
0.02
00.
560
00
00
00
00
00
00
00
00
00
00
00
00 I
246
Ma
y72
3.5
0.09
92.
837
00
00
00
00
116
00
00
00
00
01
05
00
256
Ma
y77
4.4
0.08
71.
980
00
00
00
00
07
10
00
00
00
00
00
00
266
Ma
y44
0.8
0.03
03.
791
01
01
00
00
12
00
00
00
00
00
00
00
276
Ma
y66
2.8
0.00
70.
250
00
00
00
00
01
00
00
00
00
01
00
00
286
May
561.
60.
061
3.83
02
02
120
00
00
60
00
00
100
00
02
00
00
296
May
521.
10.
037
3.38
10
00
20
00
00
180
00
00
04
00
00
00
0
306
Ma
y56
.1.
60.
041
2.57
00
02
20
00
03
00
00
00
025
00
00
70
031
6M
ay62
2.4
0.08
93.
7013
00
07
00
03
119
00
00
120
00
00
00
10
032
6M
ay68
3.1
0.04
61.
480
00
31
00
00
00
00
00
00
00
02
00
00
336
May
672.
80.
198
7.07
00
027
200
00
00
00
00
00
70
00
01
01
00
346
May
632.
30.
010
0.43
00
00
00
00
00
110
00
04
03
00
00
00
0
356
Ma
y83
5.5
0.17
23.
130
00
30
00
01
00
00
00
00
00
00
00
00
Tabl
e6
cont
inue
d.F
ish
Site
Dat
eL
Wt
Gu
t%
II
IC
HC
CC
GH
C8
DM
C8
8T
PH
TD
GE
E(m
m)
(g)
Co
nte
nt
Gu
tN
NN
AA
II
LA
YA
RE
'Y
UI
RU
0Y
LE
AU
GW
eig
ht
We
igh
tS
SS
LR
RR
AM
PP
AC
SM
VY
NL
DS
CS
P.•G
(g)
AL
PN
PA
LD
ME
RZ
ZI
AL
ZI
YR
TL
LH
S
366
Ma
y90
7.3
0.15
62.
130
00
10
00
00
00
00
00
30
00
00
00
00
378
Ma
y37
0.4
0.00
71.
635
00
01
00
03
00
00
00
00
00
10
00
00
388
May
370.
50.
012
2.36
50
00
20
00
00
00
00
00
00
00
00
00
0
398
May
340.
30.
004
1.20
10
00
10
00
10
00
00
00
00
00
00
00
0
408
May
380.
50.
009
1.78
90
00
00
00
00
00
00
00
00
00
00
00
0
418
May
480.
90.
025
2.72
180
00
80
00
20
00
00
060
00
00
00
00
0
428
May
380.
50.
020
4.06
31
00
00
00
10
00
00
00
00
00
00
00
4
438
Ma
y37
0.5
0.00
30.
669
01
011
00
00
00
00
00
00
00
00
00
00
448
Ma
y48
1.0
0.02
72.
6637
02
00
00
00
00
00
00
10
00
00
00
00
458
Ma
y38
0.6
0.02
03.
3327
00
00
00
01
00
00
00
00
00
00
00
00
468
Ma
y43
0.8
0.03
44.
2626
01
00
00
00
00
00
00
00
00
00
00
00
4711
Ma
y75
4.4
0.00
70.
150
00
00
00
00
00
00
00
30
110
00
00
06
N48
11M
ay
846.
90.
056
0.80
120
00
00
00
00
00
00
00
00
00
00
00
00
4911
Ma
y66
3.1
0.02
30.
732
00
218
00
00
00
00
00
10
00
00
00
06
5011
May
795.
40.
045
0.83
100
03
00
00
00
00
00
00
00
00
00
00
0
5111
May
784.
90.
008
0.17
120
00
00
00
00
00
01
00
02
00
00
00
052
13M
ay
541.
70.
066
3.88
340
01
00
00
13
00
00
06
00
00
00
00
053
13M
ay
440.
90.
027
3.02
151
04
00
00
03
00
00
012
00
00
20
00
354
13M
ay
511.
30.
034
2.62
290
01
00
30
02
00
00
010
00
00
00
00
00
5513
Ma
y38
0.5
0.01
83.
5020
00
28
00
02
20
00
00
200
00
00
00
00
5613
Ma
y46
1.0
0.02
82.
7817
10
12
00
00
00
00
00
90
30
00
00
00
5713
Ma
y53
1.6
0.04
52.
7918
00
31
03
00
80
00
00
10
00
01
00
0,0
5813
May
471.
20.
022
1.83
40
00
00
10
00
00
00
020
00
00
10
00
00
5913
May
541.
50.
057
3.79
250
02
30
20
028
00
00
030
00
00
80
00
060
13M
ay50
1.4
0.05
03.
5722
00
16
03
00
20
00
00
300
00
00
00
00
061
13M
ay
531.
70.
011
0.62
130
00
00
00
00
33
00
00
00
00
00
00
062
15M
ay
501.
10.
063
5.74
70
00
30
150
02
00
10
00
21
00
01
00
00
6315
Ma
y45
1.0
0.05
95.
860
30
10
01
200
20
00
00
02
00
00
00
00
064
15M
ay
481.
00.
029
2.86
40
00
00
010
00
00
00
00
00
00
00
00
00
6515
Ma
y46
1.0
0.06
46.
435
10
02
01
100
30
01
00
00
00
00
200
00
066
8Ju
n77
5.0
0.11
22.
230
00
00
00
00
30
00
00
00
00
00
00
300
067
8Ju
n86
6.4
0.30
94.
832
00
10
00
00
20
00
00
00
00
01
00
1000
068
8Ju
n83
6.3
0.27
74.
400
00
20
00
00
00
00
00
00
00
00
00
900
069
8Ju
n80
5.5
0.33
16.
023
00
00
00
00
00
00
00
00
00
00
00
1100
0
Tab
le6
cont
inue
d.F
ish
Site
Da
teL
Wt
Gu
t%
II
IC
HC
CC
GH
CB
DM
CB
BT
PH
TD
GE
E(m
m)
(g)
Con
tent
Gu
tN
NN
AA
II
LA
YA
RE
,YU
IR
U0
YL
EA
UG
We
igh
tW
eig
ht
SS
SL
RR
RA
MP
PA
CS
MV
YN
LD
SC
SP
G(g
)A
LP
NP
AL
DM
ER
ZZ
IA
LZ
IY
RT
LL
HS
708
Jun
846.
10.
022
0.37
01
00
00
00
00
00
00
00
00
00
10
020
0
718
Jun
897.
60.
191
2.51
00
00
00
00
012
00
00
00
00
00
00
030
00
728
Jun
673.
10.
165
5.31
121
11
00
00
00
00
00
00
00
00
00
020
00
738
Jun
703.
40.
214
6.28
00
012
00
00
00
00
00
00
00
00
02
030
00
748
Jun
551.
70.
023
1.33
60
05
40
02
00
00
00
00
00
00
02
07
0
758
Jun
743.
90.
006
0.15
00
04
300
09
00
00
00
00
00
00
10
04
0
762
Jul
774.
50.
053
1.17
130
01
00
00
13
00
00
00
00
00
00
00
0
772
Jul
601.
90.
031
1.61
52
10
00
00
00
04
00
00
00
00
00
00
0
782
Jul
866.
30.
026
0.41
20
02
00
00
00
00
00
00
00
00
00
00
0
792
Jul
795.
00.
022
0.44
00
00
00
00
00
00
00
00
00
00
00
00
0
802
Jul
948.
80.
094
1.07
30
00
011
00
00
00
00
00
00
00
00
00
0
816
Jul
153
47.6
1.95
34.
100
00
00
00
02
120
00
00
00
00
00
00
110
N82
6Ju
l12
725
.00.
222
0.89
00
00
00
00
00
00
00
00
00
00
00
00
0-'
837
Jul
110
14.4
0.28
31.
960
00
00
00
00
00
00
00
00
00
00
00
00
847
Jul
9810
.20.
014
0.14
00
00
00
00
02
00
00
00
00
00
50
00
0
857
Jul
906.
90.
079
1.15
270
01
04
00
10
01
00
00
00
00
10
00
0
867
Jul
101
11.5
0.08
90.
770
00
00
00
00
20
00
00
00
00
00
00
00
877
Jul
109
14.3
0.55
33.
860
00
00
00
00
10
00
00
00
00
01
01
00
8812
Jul
897.
80.
166
2.12
00
093
00
00
00
00
00
00
00
00
00
01
089
12Ju
l85
6.1
0.09
81.
616
00
170
00
00
00
00
00
00
00
01
00
00
9012
Jul
110
15.4
0.10
20.
660
00
60
10
00
00
171
40
00
00
00
00
00
9112
Jul
908.
00.
182
2.27
186
023
02
00
00
00
01
00
00
00
00
00
0
SECHELTINLET
N
A
10
13
',", 14, .~
15.' f""'4/
>11\
#-~
~»,.. ,~4:. 12 ~,~. (,'~ .
\
AGAMEMNONCHANNEL
~<~?J,.
<:fd'~
1r....."
\:-..
Figure 1. Location of beach seine sampling sites in Agamemnon Channel and SecheltInlet. Refer to Table 1 for site descriptions.
22
12 I I
10
8
>.ucQ)
6- 6~u.
4
2
o
n=48
30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105 110 115 120 125 130 135 140 145 150
Length (mm)
Figure 2a. Length frequency data for chum salmon fry caught in Agamemnon Channel(data for May and July combined, n=48).
8
7
6
5
~cQ)
6- 4~u.
3
2
o
n=43
30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105 110 115 120 125 130 135 140 145 150
Length (mm)
Figure 2b. Length frequency data for chum salmon fry caught in Sechelt Inlet (data forMay, June and July combined, n=43).
23