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    Evolution

    By Ranjitha .s

    Dept of orthodontics

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    Evolution is a process of change over a long

    period.

    It refers to the formation & development of life

    on earth.

    The idea that all living things evolved fromsimple organisms & changed through the ages

    to produce millions of species.

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    Era period Epoch Time First appearance of

    Cainozoic

    QuaternaryHolocene 1-2 million years ago Man

    Pleistocene

    Tertiary

    Pliocene

    70 million years ago

    Anthropoid

    Apes

    primates

    Miocene

    Oligocene

    Eocene

    Palaeocene

    Mesozoic

    Cretaceous 120 million yearsago

    200 million years

    ago

    Modernmammals

    Early

    mammals

    Jurassic

    Triassic

    palaezoic

    Permian

    500 million yearsago

    Reptiles

    Amphibians

    Fishes

    Carboniferous

    Devonian

    Silurian

    Ordovician

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    Charles Darwin

    On the Origin o f Species(1859)

    First to link biological diversity to evolution

    The Descen t o f Man, and Select ion in Relation

    to Sex(1871)

    Described human evolution

    Saw aesthetic factors (looks) as sexual

    attractors

    Thomas Huxley

    Evidence as to Mans Place in Nature(1863)

    First book to describe human evolution

    These scientists wrote about human evolution

    before human fossil evidence was ever discovered

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    FOSSIL EVIDENCEFOR EVOLUTION

    Paleontology

    Study of prehistoric life of all forms, typically using

    fossils

    Paleoanthropology

    Study of prehistoric human life (and human ancestry),

    typically using fossils

    Mary and Louis Leakey Lifetimes dedicated to finding fossil evidence of human

    ancestors in Africa

    Olduvai Gorge

    The Cradle of Mankind

    Great Rift Valley in eastern Africa (Tanzania)

    Erosion reveals layers of datable artifacts, bones, and fossils

    going back 2,000,000 years

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    Lamark

    In 1809 Jean-Baptiste Lamarkehypothesized that simple

    organisms emerge

    spontaneously and evolve to

    greater complexity.

    Lamarks hypothesis about

    acquired characteristics was

    disproved by Weisman, who

    concluded that changes in an

    individual during a lifetime do

    not affect its offspring.

    3 principles to his theory:1. Organisms constantly strive to

    improve themselves.

    2. Most used body structures

    develop and unused waste

    away.3. A structure is modified by use

    or disuse, the modification is

    inherited to the offspring.

    (Inheritance of acquired

    characteristics.)

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    HOMINID EVOLUTION

    Hominids= great apes

    Chimpanzees, gorillas, humans, and orangutans

    Numerous intermediary fossils have been found

    But scientists disagree on which are human ancestors and

    which are evolutionary dead ends

    Homo erectus

    Homo sapiensneanderthalensis

    Homo sapiens

    Homo habilis

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    Homo habi l is

    2.4 to 1.4 million years ago

    Fossils found in southern and eastern Africa

    Used simple bone and stone tools Nicknamed handy man

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    Homo erectus

    1.8 million years ago to

    70,000 years ago

    First human ancestor to walk

    fully uprightSome made complex stone

    tools

    ExamplePeking Man

    Descendants were humans

    and neanderthals

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    NEANDERTHALS

    Homo sapiens neanderthalensis

    400,000 to 30,000 years ago

    Lived in Europe and Asia

    Archaeogenetics analysis of ancient and modern DNA

    Comparison of human and Neanderthal DNA shows that

    humans are not descended from Neanderthals Genes reveal that Neanderthals had red hair and fair skin

    Fair skin developed to aid in the absorption of Vitamin D

    from the sun in areas far north of the equator

    Convergent evolution different species (such as humans

    and Neanderthals) developing same characteristic(s)

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    Homo neanderthalensis

    H i

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    Homo sapiens Homo (Latin) means

    man or human

    Sapiens (Latin) meanswise or intelligent

    Homo sapiens = wiseman or wise human

    Humans andNeanderthals bothconsidered sapiens

    Anatomically modernhumans in Africa by

    200,000 years ago Middle Paleolithic

    period

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    Comparative Hominid

    Anatomy Anatomically, we are very similar to apes and

    even to monkeys

    But we are not identical: Humans are fully

    bipedal, for example, while chimps are not. We have two arches in our foot; chimps do not.

    Therefore we need to compare our ownanatomy with (a) other living primates and (b)fossil hominin forms.

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    Skull Morphology: Chimp and Human

    Note the following Larger brow ridge (supraorbital torus) of chimp compared to

    human

    Sloping forehead of chimp compared to human

    More prognathous (jutting) jaw of chimp compared to human

    Larger canine and gap (diastema) of chimp compared tohuman

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    Structure of the Brain:

    Diagram Frontal Lobe and Motor

    Cortex:

    Cognition: Thinking Ability

    Motor Abilities

    Parietal Lobe: Touch and

    Taste

    Temporal Lobe: Hearing

    Occipital Lobe: Vision

    Olfactory Bulb: Smell

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    Human and Chimp Skulls Compared:

    Brain Structure Compare the following

    Chimps brain is muchsmaller (400cc vs. 1400cc)

    It has reduced frontal lobe

    It has no Brocas or

    Wernickes area It does have Brodmanns

    area 10, where calls mayoriginatebut no speech

    It does have planumtemporale, where calls arereceivedbut not processedas language

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    What This All Means

    Our brains are larger than the chimps

    We have a well-developed frontal lobe

    We have well developed language areas:Brocasand Wernickes area

    The motor strip is more well developed among

    humans than among chimps

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    Comparative Primate Anatomy:

    Chewing Mechanism The next diagrams compare humandentition (structure

    of teeth) with that of the chimps

    Our overall dental arcade (arrangement of teeth) ismore rounded (arc-like) than the chimps

    Chimpanzee have a more rectangular dental arcade,with the back teeth more parallel

    Our teeth are much smaller than the chimps

    We have small canines (jagged teeth)

    Chimps have large canines, so large that they need agap (diastema) in the opposite jaw for them to fit

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    Chimp and Human Dentition

    Note the following:

    Dental Arcade: Humans are arc-like; apes, parallel back

    teeth

    Canines and Diastema (gap): Apes have larger caninesand gaps in opposite jaw to fit them; humans do not

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    Comparative Primate

    Anatomy: Hand Structure Our fingers are straight; that of the chimps and other

    apes are curved

    We have a much longer thumb than do the apes

    Importance: we are capable of a more precise grip thanthe apes

    This implies that we can make finer tools than those

    apes who can make and use tools

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    Power and Precision Grip

    Note the Following:

    Power grip: Fingers andthumbs wrap around the

    object Precision grip:

    Forefingers and thumbhold the object

    Importance: We can do

    finer work compared tononhuman primates

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    Comparative Primate Anatomy:

    Bipedalism vs. Quadrupedalism Homo sapiens is the only mammal capable of

    bipedalism, or the ability to stand and walk entirely ontwo feet.

    Kangaroos stand on two feet, but they hop rather thanwalk and their forepaws are too small for any function.

    Chimpanzees can walk on two feet, but not veryefficiently; they are closer to quadrupedalism, or theability to move around on four feet.

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    Advantages of Bipedalism

    Efficient locomotion

    Freeing of hands for many purpouses:

    Foraging and hunting/scavenging

    Tool making and use

    Care and provisioning of offspring

    Increased height for viewing across landscape:

    Tracking migrating herds

    Predator avoidance

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    Survival of the Fittest

    Individuals that are best suited to anenvironment (adaptations that enable fitness)

    survive and reproduce most successfully.

    Over time, natural selection results in

    changes in the inherited characteristics of a

    population. These changes increase a

    species fitness in its environment.

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    Natural Selection

    Natural Selection leads to organisms that are well

    suited in particular environments.

    It provides a scientific explanation for the historyof life on Earth as depicted in the fossil record and

    similarities evident within the diversity of existing

    organisms.

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    A-- AGNATHA

    B-- AUTOSTYLY

    C-- AMPHYSTYLY

    E,F-- HYOSTYLY

    G-- HOLOSTYLY

    Evolution of jaws

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    Comparative Primate Anatomy:

    Human and Chimpanzee In the next diagrams, the differences are significant to

    biological capacity for culture

    The area for brain of a chimp is more limited thanhuman brain because of its sloping forehead andtheheavy supraorbital torusthat covers much of the

    forehead The chimp jaw has a prognathismabsent in humans

    Chimps have largercanineteeth than humans; somuch so that there is a diastema(gap) for the

    opposite canine to fit.

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    EVOLUTIONIts the inheritance of the

    heritable endowmentAlso defined as development of species

    from earlier forms

    Factors responsible for evolution are:-1. Genetic

    2. Environmental

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    SIMPLIFIED CLASSIFICATION

    OF LIVING VERTEBRAES CLASS EXAMPLES

    1. Agnatha Hagfishes,lampreys

    2. Chondrichthyes Sharks,rays3. Osteichthyes Bony fishes

    4. Amphibia Frogs,toads

    5. Reptilia Lizards,crocodiles,snakes

    6. Aves Birds

    7. mammalia

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    MAMMALIA

    SUBCLASS

    PROTHERIA METATHERIA EUTHERIA

    e.g. platypus e.g.red kangaroo e.g,placental mammals

    SUBCLASS

    PROTHERIA METATHERIA EUTHERIA

    e.g. platypus e.g.red kangaroo e.g,placental mammals

    HUMAN EVOLUTION

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    HUMAN EVOLUTION

    Homo habi l is

    2.4 to 1.4 million years ago

    Fossils found in southern and eastern Africa

    Used simple bone and stone tools

    Nicknamed handy man

    Homo erectus 1.8 million years ago to 70,000 years ago

    First human ancestor to walk fully upright

    Some made complex stone tools

    ExamplePeking Man Descendants were humans and neanderthals

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    Development of Conceptof Occlusion

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    Development of Concepts of occlusion

    It can be divided into

    Fictional Period (prior to 1900)

    Hypothetical Period (1900 to 1930)

    Factual Period (1930 to present)

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    Fict ional Period

    Fuller Clark & Imrie talked of antagonism,meeting or gliding of teeth.

    Talbotsaid its a historical event long since

    passed in the decline of species, and possibleonly with an atavism throwback to our primitiveancestors.

    The creation of a normal standard of occlusion ,a basis on which to compare departure fromnormal was lacking.

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    Hypo thet ical Period

    According to

    Edward H. Angel

    Mathew Cryer and Calvin Case

    Lischer and Paul Simon

    Milo Hellman

    Ed d H A l

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    Edward .H. Angle

    In 1907Angle put a hypothesis that:

    Occlusion is the basis of the science of orthodontia. The shapes of thecusps, crowns and roots , and even the very structural material of the teeth

    and attachments are all designed for the purpose of making occlusion the one

    grand object. we shall define occlusion as being normal relations of the

    occlusal inclined planes of the teeth when the jaws are closed.

    He described the illustration Old glory. which according

    to him was ideal occlusion.

    Angle stated Apollo Belverdos straight profile as ideal

    The best eg. of his hypothesis was key of Occlusion

    His hypothesis was suggestive of static occlusion

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    Old GlorySkull

    Apollo Belverdo

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    Mathew Cryer and Calvin caseHe opposed Angle

    How could one mix prognathic denture and

    orthognathic profile?

    How bimaxillary protrusion considered normal and

    individual variation was not considered?

    Then they were proved right by Turners illustrations

    Cases concept of occlusion was static

    Normal occlusion( it suits Apollos face better)

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    Wheelers illustrations for normal

    occlusion

    PrognathicRetrognathic

    Lischer And Paul Simon

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    Lischer And Paul Simon

    They tried to relate concept of occlusion by

    relating teeth with rest of the face andcranium.

    Lischer introduced mastication as requisite of

    occlusion

    Simon outlined the anthropometric approach

    Milo HellmanHe showed racial variation in normal

    occlusion

    His study was also confined to static occlusion

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    Factual Period

    Dividing line b/w static and dynamic occlusion.

    1931 Broadbent introduced Cephalometry.

    With this advanced study tech the factual

    period become functional period.By now 3 components of occlusion are set up1. Interdigitation of teeth

    2. Status of controlling musculature

    3. TMJ integrity

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    Ardipithecus ramidus

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    Australopithecus afarensis

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    Australopithecus africanus

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    Homo ergaster

    Homo ergasteris one of the more problematic of species

    designations

    Each researcher that sees ergasteras valid sees different

    specimens as belonging or not belonging

    Most researchers see too little difference between ergaster

    and erectus to form the basis of a species As a general rule of thumb, one can consider most

    attributed ergasterspecimens to be early erectus

    geographically confined to Africa

    H t

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    Homo ergaster

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    Evo lut ion o f den t it ion

    E l ti f D titi

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    Evolution of Dentition

    The mammalian dentition is 100, 000 million yrs old

    The theories behind mammalian dentition are

    Theory o f Concrescence derived by fusion of 2

    or more primitive conical teeth. Theory of tr i tuberculy each tooth derived from

    single reptilian tooth by secondary differentiation of

    tubercule and root.

    Theory o f mu lt i tuberculy derived from reductionand condensation of primitive multituberculate teeth.

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    Non mammalian vertebratesPolyphyodonty(many dentitions)

    In mammalsDiphyodonty (2 dentitions )

    During growthof the animal increase in the

    jaw size is associated with tooth size

    Evolution of Dentition

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    Dentition off ishes( Agnatha)

    Eg - Sea lamprey & hag fishes

    No true calcified teethArranged circumferentially

    Horny teeth are also seen on the tongue

    Chondr ichthyes (carrtilagenous fishes)

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    Chondr ichthyes(carrtilagenous fishes)

    Large no. of teeth are presentHomodont & polyphyodont

    Teeth are covered by enameloid

    lower jaw of shark

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    Osteichthyes

    (bony f ishes)

    Haplodont conical shape

    prehension

    Polyphyodont

    Teeth- vomer, palatine bones roof of the mouth

    & tongue

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    Amphib ia

    Small, homodont, polyphyodont prehension

    In the f rog small teeth on the upper jaw &

    no teeth on the lower jaw

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    Repti les

    Homodont & polyphyodont Tend to be tricuspid or cone shaped

    Egg teeth in embryos ofl izards & snakes

    - used to break the shell

    Dentition of mammals

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    Dentition ofmammals

    Heterodont ( 4 types )Restricted to 2 rows

    Ability to masticate improves digestive

    efficiency for high rate of metabolism

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    Other features

    TMJ articulation

    Salivary glands

    Prismatic enamel Diphyodonty

    Secondary palate

    Significant muscle development

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    Dentition ofprother ia

    Spiny ant eater edentulous

    Duck b i l led platypu s 3 functional posterior teeth ineach jaw quadrant

    No. of additional teeth develop but do not erupt but

    rapidly resorb

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    Metatheria

    (Marsupials)

    Eg. Red Kangroo

    Dental formula 3/1, 0/0, 1/1, 4/4 .Feed on various plant.

    Anterior teeth allow the food to be firmly grasped.

    Marsupials have no. of dental features which

    distinguish them from placental animals.

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    Eutheria

    Eg - hedge hog

    Insectivoral

    Incisors form a forcep like mechanism to pick preys.

    Molars primitive tritubercular with sharp cusps

    Suitable for crushing the shells of insects

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    Rodent ia

    Eg - rat

    Heterogenous , Monophyodont

    Continuously growing incisors

    DF 1/1, 0/0, 0/0, 3/3

    Rat & mice are widely used as experimental animals inDental sciences

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    Carnivora

    Incisors are small and arranged in line.

    Enlarged canines offensive weapons

    Incisors and canine tearing flesh

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    Primates

    Classified in to

    -Pros imi i or Lemu r type

    -Anthropoidea

    Mainly herbivorous sometimesinsectivorous

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    Pros im ii or Lemur type

    DF 2/2, 1/1, 3/3, 3/3

    incisors & canines are procumbent and form dental

    comb.

    mid line diastema .Upper canine is blade like

    Upper molar tritubercular

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    Anthropoidea(monkey, apes &Man)

    Eg -Macaca monkey

    DF 2/2, 1/1, 2/2, 3/3

    Canines are long & prominent ( esp. in male )

    Lateral diastema

    Chimpanze

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    Chimpanze

    Resembles permanent teeth of man

    Homosapiens

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    Homosapiens

    Modern man exis ted 100,000 yr s. ago