exercise: designing finite state machines · 2016. 1. 22. · exercise: designing finite state...

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ORIGINAL ARTICLE The loss of dispersal on islands hypothesis revisited: Implementing phylogeography to investigate evolution of dispersal traits in Periploca (Apocynaceae) C. Garc ıa-Verdugo 1 | M. Mairal 2 | P. Monroy 1,3 | M. Sajeva 4 | J. Caujap e-Castells 1 1 Departamento de Biodiversidad Molecular y Banco de ADN, Jard ın Bot anico Canario Viera y ClavijoUnidad Asociada CSIC, Cabildo de Gran Canaria, Las Palmas de Gran Canaria, Spain 2 Real Jard ın Bot anico de Madrid, CSIC, Madrid, Spain 3 Fundaci on Canaria Amurga-Maspalomas, San Bartolom e de Tirajana, Spain 4 Dipartimento STEBICEF, Universit a di Palermo, Palermo, Italy Correspondence Carlos Garc ıa-Verdugo, Departamento de Biodiversidad Molecular y Banco de ADN, Jard ın Bot anico Canario Viera y Clavijo- Unidad Asociada CSIC, 35017 Las Palmas de Gran Canaria, Spain. Email: [email protected] Funding information PCT-MAC, Ministerio de Economia y Competitividad, Cabildo de Gran Canaria, Grant/Award Number: ENCLAVES (MAC/3/ C141), JCI-2012-15220 Editor: Sonya Clegg Abstract Aim: The loss of dispersal on islands hypothesis (LDIH) posits that wind-dispersed plants should exhibit reduced dispersal potential, particularly if island populations are old. In this study, we tested this hypothesis using a detailed phylogeographical framework across different geographical scales. Location: Mainland and island areas of the Atlantic and Mediterranean regions, including Macaronesia (Canary Islands and Cape Verde) and Mediterranean islands in the strait of Sicily. Methods: Forty-five populations of Periploca laevigata, a wind-dispersed shrub, were sampled. Plastid and nuclear microsatellite data were used to reconstruct spatio- temporal patterns of island colonization, and estimates of seed terminal velocity used as a surrogate for dispersal ability under both field and common garden condi- tions. Results: Our findings did not provide evidence of loss of dispersability in any island lineage. In all of the regions considered, dispersal ability was similar on island and mainland populations, or higher on islands. Contrary to LDIH expectations, lineages inferred as the oldest (western Canaries and Cape Verde) converged towards the most dispersive seed phenotype. This pattern was supported by data obtained under common garden conditions. Within the western Canarian lineage, successful dispersal was shown to be very rare among islands and extensive within islands, but dispersability did not vary significantly from older to more recent sublineages. Con- sidering all the study islands, we found a strong, positive correlation between dis- persal ability and estimates of within-island habitat availability. Main conclusions: This study suggests that dispersal ability can be favoured on islands, possibly because traits enhancing wind dispersal are positively selected when habitat availability is high. Our results challenge broad generalizations of the LDIH, but we discuss how overlooking species 0 phylogeographical history may give rise to misleading conclusions. KEYWORDS anemochory, dispersal ability, island colonization, Macaronesia, parallel evolution, seed dispersal DOI: 10.1111/jbi.13050 Journal of Biogeography. 2017;112. wileyonlinelibrary.com/journal/jbi © 2017 John Wiley & Sons Ltd | 1

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  • Exercise: Designing Finite State Machines Activity sheet to support Tenderfoot Unit 5: Theoretical Computers

    Source: Ideas and resources from Maths Maniacs www.mathmaniacs.org

    Hair Dryer: Airhead 2020

    Let's design a finite state machine that models the behaviour of a hair dryer - the top of the range Airhead 2020!

    Here are the specifications:

    Inputs: The inputs to the hair blower can be thought of as moving the switches in a given direction. So, we would have 4 inputs: Slide the left button up (let's call this "L-up"), slide the left button down (L-down), and similarly for R-up and R-down.

    Outputs: The machine is slightly different to the ones developed in the Fickle Fruit activites. The fruit activities had a singular output (a banana or apple) in response to an input request. We labelled the output on the transition, paired with the input.

    Here, the button will cause, say, air to be out-put, but not just one "chunk" of air. Instead, it makes more sense to think of the machine as being "in a continuous state of blowing air", until another input is received.

    We will therefore have NO outputs, but instead label the states to indicate what the hair blower is doing at any given time.

    States: As explained, the states of the machine should reflect exactly what the hair blower is doing at a given time. Thus, there would be one state called "blowing cool air fast", another one for "blowing hot air slow" and so on.

    Hint: make a state for each possible arrangement of the switches.

    Transitions: The machine should behave as you suspect. For example, if it was blowing cool air fast, and the input R-down were received, then it should next be in the state of blowing cool air slow.

    Note: In the state shown, if L-down were received, then since there is nothing cooler than "cool", the machine's behaviour would not change - it would stay in the same state. This would be like trying to force the button down further than it can go. (Or perhaps we could add a state for "broken"?).

    On the reverse side, or on a separate piece of paper, draw a state diagram for the Airhead

    2020