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    The History of Life on Earth

    When did life arise onEarth?

    The Earth is thought to

    be approximately 4.6billion years old, but life

    is believed to have

    occurred approximately 4

    billion years ago (bya)

    How did life begin???

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    The Origin of Life: Early Ideas

    Spontaneous Generation idea popular in the 1600-1700s

    living things come from the nonliving

    evidence: beetles and other insect larvae arise fromcow dung; frogs emerge from mud

    In 1688, the Italian Francisco Redi In 1668,Francesco Redi, an Italian physician, did anexperiment with flies and wide-mouth jars. Hedemonstrated that meat that was covered did

    not produce maggots

    This may have been the first true scientificexperiment

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    Francesco Redi experiment with

    flies and wide-mouth jars

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    The Origin of LifeSpontaneous generation

    Mid-1800sdisproved by Louis Pasteur andJohn Tyndall

    Broth in flask is boiled

    to kill preexisting

    microorganisms.

    Condensing water collects

    as the broth cools, sealing

    the mouth of the flask.

    If neck is later broken off,

    outside air can carry

    microorganisms into broth.

    no growth growth

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    Other Ideas: Life from aBiblical Creation?

    Christian Creationism statesthat the world, including alllife, was created about 6,000years ago in six literal daysby a God.

    But how does one accuratelyand fairly test for this?...

    Whats the observation,

    hypothesis, test?

    This idea does not really fit intothe confines of a Science

    course.

    Like the study of French Impressionist

    painters, Religion is not part of, nor

    adequately covered in, a Science course.

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    Origin of Life:Another idea

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    Biogenic-looking features in

    ALH84001 Martian meteorite

    http://ares.jsc.nasa.gov/astrobiology/biomarkers/images.html

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    In 1969, a meteorite (left-over bits from the origin of the

    solar system) landed near Allende, Mexico. The AllendeMeteorite (and others of its sort) have been analyzedand found to contain amino acids, the building blocksof proteins.

    This idea ofpanspermiahypothesized that life originatedout in space and came to earth inside a meteorite. Theamino acids recovered from meteorites are in a groupknown as exotics: they do not occur in the chemicalsystems of living things. The ET theory is nowdiscounted by most scientists, although the August1996 discovery of the Martian meteorite and its

    possible fossils have revived thought of life elsewherein the Solar System.

    Anyway.This only moves the problem to elsewhere!

    Extra-terrestrial Origins

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    The Lates t on Extra-terres tr ial

    Origins

    The Raelians Raelians believe that humanity

    was created from the DNA of

    superior alien scientists Follow the teachings of a

    former French magazinesportswriter and wannabe

    race-car driver ClaudeVorilhon, 56. He took thename "Rael" after he claimeda close encounter of the third

    kind.

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    Origin of Life: Current Theory

    Chem ical Evo lut ion

    .....The idea that long ago complex

    collections of chemicals formed the first

    cells.

    Life began in the oceans 4 bya from

    simple chemicals joining together in a

    primordial soup

    Complex chemicals evolved into living

    cells

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    What were the conditions like on Earth when life arose?

    Up to about 4 bya, asteroid impacts and volcanic eruptions resulted

    in the release of various gases that began to form an atmosphere

    It consisted mainly of CO2, with some nitrogen, water vapor and

    sulfur gases; hydrogen quickly escaped into space

    CO2 in the atmosphere trapped solar radiation, making the Earths

    surface rather warm

    Earth was cool enough to form a crust, and water vapor condensed

    to form oceans

    Oceans in turn helped to dissolve CO2 from the atmosphere and

    deposit it into carbonate rocks on the seafloor

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    What were the conditions like on Earth when life

    arose?

    Organic molecules were undoubtedly beingformed on the Earths surface

    Lightening and ultraviolet radiation from the Sun

    acted on the atmosphere to forms small traces of

    many different gases, including ammonia (NH3),

    methane (CH4), carbon monoxide (CO) and ethane

    Also, cyanide (HCN) probably formed easily in theupper atmosphere, from solar radiation and then

    dissolved in raindrops

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    The Origin of Life

    The possible origin of

    organic molecules

    a. 1953theStanley Miller

    experiment

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    What is the simplest living cell that one can imagine?

    A universal minimal cell

    must contain the following::

    Cell membrane

    Cytoplasm DNA and RNA

    Proteins

    Enzymes

    Ribozymes

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    The Origin of LifeEarly Speculations

    More circumstantial evidence

    accumulated

    Astronomers found simple organic

    compounds in meteorites

    They were convinced that Earths initialatmosphere could not have matched

    Oparin-Haldanes model

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    The Origin of LifeEarly Speculations

    More circumstantial evidence

    Fossils of ancient bacteria (3.5 billion years old)

    were found in Australia

    Suggested life may have evolved rapidly in less

    than a billion years

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    The Origin of LifeEarly Speculations

    What are the possible scenarios?

    When ocean tidal pool evaporates Salts get highly concentrated

    Could have happened in ancientoceans

    Concentrating aminos, may allow

    protein to form

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    The Origin of LifeEarly Speculations

    Phospholipids arrange themselves into bubbles

    Chemicals could be concentrated in bubbles (mightcontain protein, etc.)

    These bubbles would persist aided by natural selection

    If they burst, spew contents into air where otherreactions occur

    Over hundreds of millions of years, similar processescould have filled oceans with proteins, carbohydrates,phospholipids, nucleotides

    Th O i i f Lif

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    The Origin of LifeEarly Speculations

    Phospholipids arrange themselves intobubbles

    Eventually they reach a level ofcomplexity

    Called protocells (not living)

    Still cant reproduce, no DNA

    Th O i i f Lif

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    The Origin of LifeEarly Speculations

    Is DNA essential?

    Scripps Institute, 1993 found smallmolecules of synthetic RNA that within an

    hour began making copies of itself & the

    copies made more copies

    Then copies began to change - evolve-

    acquiring new chemical characteristics,

    but not alive

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    The Origin of LifeEarly Speculations

    Is DNA essential? Protocells might qualify as the first cells if they have

    RNA that:

    Can make copies of itself & evolve

    Could synthesize enzymes capable of breaking downother organic compounds

    Could synthesize enzymes capable of building and

    maintaining cell membranes

    Later DNA could have evolved as method ofconveniently & safely

    Storing vital chemical info contained in cell RNA

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    The Origin of LifeEarly Speculations

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    The First Cells

    Age of microbes3.5 billion years ago

    1. The earliest living cellsanaerobic

    prokaryotes

    2. Photosynthetic bacteria and the

    evolution of an oxygen-rich environment

    3. Development of aerobic metabolism

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    II. The first cells

    The rise of eukaryotesabout

    1.4 billion years ago

    1. Endosymbiotic hypothesis

    2. The origin of the nucleus

    1. Anaerobic, predatory

    prokaryotic cell engulfs

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    prokaryotic cell engulfs

    an aerobic bacterium.

    2. Descendants of engulfed

    bacterium evolve into

    mitochondria.

    3. Mitochondria-containing

    cell engulfs a photosynthetic

    bacterium.

    4. Descendants of photosynthetic

    bacterium evolve into chloroplasts.

    aerobic

    bacterium

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    First Cell Types

    Heterotrophic cells

    Incapable of producing their own food

    Autotrophs Can produce chemicals to store energy

    Chemoautotrophs

    Store energy found in certain inorganic chemicals

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    First Cell Types

    Most organisms found free oxygen intolerable

    In oceans

    Organisms that built simple and complex organic

    compounds

    Removed CO2 from the atmosphere

    More advanced autotrophs removed most of the rest

    & replaced it with oxygen

    The excess oxygen changed forever chemical natureof atmosphere to todays

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    Further Evolution of First Cells

    First cells, prokaryotes, were always simple

    in structure

    2 - 1.5 billion years ago A new cell appearedeukaryotes

    Had membranes to isolate certain chemical

    reactions

    Cellular life then evolved into what we know

    today

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    Archaea & Bacteria Domains

    Directly related to oldest organisms on

    earth

    Have had lots of time to evolve & differentiate

    Thrive nearly everywhere

    Depths of oceans & Earth, all surfaces

    M lti ll l i

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    Multicellular organisms

    A. Advantages of multicellularity

    B. Challenges of multicellularity

    C. The first multicellular organisms

    1. Plantsprimitive marine algae

    2. Animalsmarine invertebrates

    D. The transition to land

    1. Advantages of terrestrial living

    2. Challenges of terrestrial living

    III M lti ll l i

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    III. Multicellular organisms

    The transition to land

    The evolution of land plants

    a. The first land plants

    1) Mosses and ferns 2) Continued water dependency

    b. Conifersthe invasion of dry habitats

    c. Flowering plants

    1) The dominant plant form today

    2) Pollination by insects

    III M lti ll l i

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    III. Multicellular organisms

    D. The transition to land

    The evolution of terrestrial animals a. Arthropods

    b. Lobefin fish to amphibians

    c. Amphibians to reptiles 1) The age of the dinosaurs

    2) Reptiles and maintenance of body temperature

    d. Birds 1) Insulating feathers retain body heat

    2) Evolution of feathers for flight

    III M lti ll l i

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    III. Multicellular organisms

    The evolution of terrestrial animals

    e. Mammals

    1) Insulating hair retains body heat

    2) Live births and mammary glands

    IV H man e ol tion

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    IV. Human evolution

    A. Primate evolution

    1. Grasping handsprecision grip and

    power grip

    2. Binocular and color vision with

    overlapping fields of view

    3. Large brainallows fairly complex social

    systems

    H.habil is

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    Ardip i thecusramidus

    Austra lop i thecusafarensis

    H. heidel-bergensis

    A. boisei

    A. afr icanus

    A. robustus

    H.sapiens

    H. erectu s

    Homo ergaster

    H. neanderth alensis

    IV Human evolution

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    IV. Human evolution

    Hominid evolution

    1. The evolution of dryopithecinesbetween

    20 and 30 million years ago

    2. Australopithecinesthe first true hominids

    a. Appeared 4 million years ago as evidenced by

    fossils

    b. Walked upright

    c. Large brains

    IV Human evolution

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    IV. Human evolution

    3. Homo habilis2 million years ago

    a. Larger body and brain

    b. Ability to make crude stone and bone tools

    4. Homo erectus1.8 million years ago

    a. Face of modern human

    b. More socially advanced c. Sophisticated stone tools aided in hunting

    d. Used fire

    IV Human evolution

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    IV. Human evolution

    5. Homo sapiens200,000 years ago

    a. Neanderthals evolved 100,000 years ago

    1) Similar to humansmuscular, fully erect,

    dexterous, large brains

    2) Developed ritualistic burial ceremonies

    b. Cro-Magnons evolved 90,000 years ago

    1) Direct descendants of modern humans

    2) Were artistic and made precision tools