mallory hagadorn dr. dana l. price department of ...faculty.salisbury.edu/~dlprice/su lab page/about...

Quick Guide for the Identification of

Maryland Scarabaeoidea

Mallory Hagadorn

Dr. Dana L. Price

Department of Biological Sciences

Salisbury University

This document is a pictorial reference of Maryland Scarabaeoidea genera (and sometimes species) that was created to expedite the identification of Maryland Scarabs. Our current understanding of Maryland Scarabs comes from “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland” (Staines 1984). Staines reported 266 species and subspecies using literature and review of several Maryland Museums. Dr. Price and her research students are currently conducting a bioinventory of Maryland Scarabs that will be used to create a “Taxonomic Guide to the Scarabaeoidea of Maryland”. This will include dichotomous keys to family and species based on historical reports and collections from all 23 counties in Maryland. This document should be cited as: Hagadorn, M.A. and D.L. Price. 2012. Quick Guide for the Identification of Maryland Scarabaeoidea. Salisbury University. Pp. 54. Questions regarding this document should be sent to: Dr. Dana L. Price - [email protected]

**All pictures within are linked to their copyright holder.

mailto:[email protected]

*All photographs are linked to their copyright holder. 2

Table of Contents

Families of Scarabaeoidea of Maryland……………………………………... 6

Geotrupidae……………………………………………………………………. 7

Subfamily Bolboceratinae……………………………………………… 7

Genus Bolbocerosoma………………………………………… 7

Genus Eucanthus………………………………………………. 7

Subfamily Geotrupinae………………………………………………… 8

Genus Geotrupes………………………………………………. 8

Genus Odonteus...……………………………………………… 9

Glaphyridae............................................................................................... 9

Genus Lichnanthe vulpina…………………………………….. 9

Hybosoridae…………………………………………………………………… 10

Genus Germarostes…………………………………………… 10

Lucanidae………………………………………………………………………. 10

Subfamily Aesalinae…………………………………………………… 10

Genus Nicagus…………………………………………………. 10

Subfamily Luncaninae…………………………………………………. 10

Genus Dorcus…………………………………………………... 10

Genus Lucanus………………………………………………… 11

Genus Platycerus………………………………………………. 12

Subfamily Syndesinae…………………………………………………. 12

Genus Ceruchus……………………………………………….. 12

Passalidae……………………………………………………………………… 12

Genus Odontotaenius…………………………………………. 12

Scarabaeidae………………………………………………………………….. 13

Subfamily Aphodiinae…………………………………………………. 13

Tribe Aegialiini……………………………………………………… 13

Genus Aegialia………………………………………………… . 13

Genus Caelius………………………………………………..... 13

Tribe Aphodiini……………………………………………………… 14

Genus Acrossus………………………………………………... 14

Genus Agoliinus………………………………………………... 14

Genus Aphodius………………………………………………... 15

Genus Blackburneus…………………………………………... 15

Genus Calamosternus………………………………………… 15


Genus Chilothorax…………………………………………….. 16

Genus Colobopterus………………………………………….. 16

Genus Dialytellus……………………………………………… 16

Genus Dialytes………………………………………………… 17

Genus Flaviellus………………………………………………. 17

Genus Irrasinus……………………………………………….. 17

Genus Labarrus……………………………………………….. 18

Genus Lechorodius…………………………………………… 18

Genus Melinopterus………………………………………….. 18

Genus Oscarinus……………………………………………... 19

Genus Otophorus…………………………………………….. 19

Genus Oxyomus……………………………………………… 19

Genus Pardalosus……………………………………………. 20

Genus Planolinellus………………………………………….. 20

Genus Planolinus…………………………………………….. 20

Genus Pseudagolius……………………………………….... 21

Genus Stenotothorax………………………………………... 21

Genus Strigodius…………………………………………….. 21

Genus Teuchestes…………………………………………... 22

Tribe Didactyliini………………………………………………….. 22

Genus Aidophus……………………………………………… 22

Tribe Eupariini…………………………………………………….. 22

Genus Aphotaenius………………………………………….. 22

Genus Ataenius………………………………………………. 23

Genus Pseudataenius……………………………………….. 23

Genus Saprosites……………………………………………. 24

Tribe Psammodiini……………………………………………….. 24

Genus Pammodius…………………………………………... 24

Genus Rhyssemus…………………………………………... 25

Genus Pleurophorus…………………………………………. 25

Subfamily Cetoniinae………………………………………………… 26

Tribe Cetoniini……………………………………………………. 26

Genus Euphoria……………………………………………… 26

Tribe Cremastocheilini…………………………………………… 28

Genus Cremastocheilus…………………………………….. 28

Genus Psilocnemis………………………………………….. 28

Tribe Gymnetini…………………………………………………… 29

Genus Cotinis……………………………………………….... 29

Tribe Trichiini……………………………………………………… 29

Genus Gnorimella……………………………………………. 29

Genus Osmoderma………………………………………….. 29

Genus Trichiotinus…………………………………………… 30


Genus Trigonopeltastes…………………………………….. 31

Tribe Valgini………………………………………………………. 31

Genus Valgus……………………………………………….... 31

Subfamily Dynastinae………………………………………………... 31

Tribe Cyclocephalini……………………………………………... 31

Genus Cyclocephala…………………………………………. 31

Genus Dyscinetus……………………………………………. 32

Tribe Dynastini…………………………………………………….. 32

Genus Dynastes……………………………………………… 32

Tribe Oryctini……………………………………………………… 32

Genus Strategus……………………………………………… 32

Genus Xyloryctes…………………………………………….. 33

Tribe Pentodontini………………………………………………... 33

Genus Aphonus………………………………………………. 33

Genus Euetheola…………………………………………….. 34

Genus Tomarus………………………………………………. 34

Tribe Phileurini……………………………………………………. 35

Genus Phileurus……………………………………………… 35

Subfamily Melolonthinae…………………………………………….. 35

Tribe Diplotaxini…………………………………………………... 35

Genus Diplotaxis……………………………………………... 35

Tribe Hopliini………………………………………………………. 36

Genus Hoplia………………………………………………….. 36

Tribe Macrodactylini………………………………………………. 37

Genus Dichelonyx…………………………………………….. 37

Genus Macrodactylus……………………………………….... 38

Tribe Melolonthini…………………………………………………. 38

Genus Phyllophaga…………………………………………... 38

Genus Polyphylla……………………………………………… 40

Tribe Sericini………………………………………………………. 41

Genus Maladera………………………………………………. 41

Genus Nipponoserica………………………………………… 41

Genus Serica………………………………………………….. 41

Subfamily Rutelinae…………………………………………………... 42

Tribe Anomalini......................................................................... 42

Genus Anomala……………………………………………….. 42

Genus Popillia…………………………………………………. 44

Genus Strigoderma………………………………………….... 45

Tribe Rutelini………………………………………………………. 45

Genus Cotalpa………………………………………………… 45

Genus Parastasia…………………………………………….. 45

Genus Pelidnota………………………………………………. 46


Subfamily Scarabaeinae……………………………………………... 46

Tribe Ateuchini…………………………………………………….. 46

Genus Ateuchus……………………………………………..... 46

Tribe Canthonini…………………………………………………… 46

Genus Canthon………………………………………………... 46

Genus Melanocanthon……………………………………….. 47

Tribe Coprini……………………………………………………….. 47

Genus Copris………………………………………………….. 47

Genus Dichotomius…………………………………………... 48

Tribe Onthophagini……………………………………………….. 48

Genus Onthophagus………………………………………….. 48

Tribe Phanaeini…………………………………………………… 49

Genus Phanaeus……………………………………………... 49

Trogidae……………………………………………………………………….. 50

Subfamily Troginae……………………………………………………. 50

Genus Omorgus……………………………………………….. 50

Genus Trox…………………………………………………….. 51

References…………………………………………………………………….. 53


Families of the Scarabaeoidea of Maryland

Geotrupidae Glaphyridae Hybosoridae

Geotrupes hornii Lichnanthe vulpina Germarostes globosus

Lucanidae Passalidae

Lucanus capreolus Odontotaenius disjunctus

Scarabaeidae Trogidae

Canthon viridis Trox spinulosus


GEOTRUPIDAE

Subfamily Bolboceratinae

Genus: Bolbocerosoma

Species: Bolbocerosoma farctum

Dichotomous Key: Scarab Beetles of South Carolina

Key to Tribe Bolboceratini

Page 51

http://bugguide.net/node/view/6814/bgimage?from=38

Genus: Bolbocerosoma

Species: Bolbocerosoma tumefactum



Page 51

http://bugguide.net/node/view/429013/bgimage

Genus: Eucanthus

Species: Eucanthus lazarus

Similar to: E. impressus



Page 51; Skip to 9(5)


Back to Table of Contents

http://bugguide.net/node/view/6814/bgimage?from=38




Genus: Eucanthus

Species: Eucanthus subtropicus





Subfamily Geotrupinae

Genus: Geotrupes

Species: Geotrupes splendidus

Similar to: G. balyi, G. semiopacus


Key to Tribe Geotrupini



Genus: Geotrupes

Species: Geotrupes blackburnii

Similar to: G. egeriei










Genus: Geotrupes

Species: Geotrupes hornii

Similar to: G.ulkei (Cited in Literature)





Genus: Odonteus (Bolboceras)

Species: Odonteus ( Bolboceras) liebecki

Simlar to: O. cornigerus (cited in Literature)

O. darlingtoni, O. falli, O. thoracicornis


Key to Tribe Bolboceratini, Page 51; skip to

2(1) O. falli-The Scarabaeoid Beetles of

Nebraska Key to Odonteus, Page 38


GLAPHYRIDAE

Genus: Lichnanthe

Species: Lichnanthe vulpina

Dichotomous Key:

Species description in Scarab Beetles of South

Carolina, Page 58







HYBOSORIDAE

Ceratocanthinae

Genus: Germarostes (Cloetous)

Species: Germarostes (Cloetous) globosus

Similar to: Germarostes (Cloetous) aphodioides

Dichotomous Key:

The Scarabaeoid Beetles of Nebraska,

Key to Germarostes, Page 130



LUCANIDAE

Subfamily Aesalinae

Genus: Nicagus

Species: Nicagus obscurus

Dichotomous Key:

For Species Description see the link below

http://www-

museum.unl.edu/research/entomology/Guide/Scaraba

eoidea/Lucanidae/AES/NIC/obscurus.html

Subfamily Lucaninae

Genus: Dorcus

Species: Dorcus brevis (Cited in Literature)

Dichotomous Key:

For Genus Description see Link Directly Below

http://www-

museum.unl.edu/research/entomology/Guide/S

carabaeoidea/Lucanidae/LUC/DOR/brevis.html

http://bugguide.net/node/view/60176




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Lucanidae/AES/NIC/obscurus.html



http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Lucanidae/LUC/DOR/brevis.html





Genus: Dorcus

Species: Dorcus parallelus

Dichotomous Key: Species Description in


Page 108

.


Genus: Lucanus (Pseudolucanus)

Species: Lucanus capreolus

Dichotomous Key:

The Scarabaeoid Beetles of Nebraska

Key to Lucanus, 0

http://www.flickr.com/photos/pcoin/5914932906/in/set-

72057594074593321

Genus: Lucanus

Species: Lucanus elaphus

Dichotomous Key:


Key to Lucanus, Page 110

http://www.flickr.com/photos/entomopixel/5098352100

/



http://www.flickr.com/photos/pcoin/5914932906/in/set-72057594074593321

http://www.flickr.com/photos/pcoin/5914932906/in/set-72057594074593321

http://www.flickr.com/photos/entomopixel/5098352100/



Genus: Platycerus

Species: Platycerus virescens

Similar to: Platycerus piceus



Page 106


Subfamily Syndesinae

Genus: Ceruchus

Species: Ceruchus piceus



Page 105


PASSALIDAE

Genus: Odontotaenius (Popilius)

Species: Odontotaenius disjunctus



Page 25







SCARABAEIDAE

Aphodiinae

Tribe: Aegialiini

Genus: Aegialia

Species: picture just representative of genus

Similar to: Aegialia blanchardi (cited in literature)

***Species found in Maryland***

Dichotomous Key:

Scarab Beetles of South Carolina

Key to Aegialia, Page 25

http://www-

museum.unl.edu/research/entomology/Guide/Scarabaeoidea/S

carabaeidae/Aphodiinae/AphodiinaeTribes/Aegialiini/Aegialia/A

egialia.html

Genus: Caelius (Aegialia)


Similar to: Caelius humeralis (Aegialia humeralis)

(Cited in the Literature)


Dichotomous Key:


Key to Aegialia, Page 25

http://www-


carabaeoidea/Scarabaeidae/Aphodiinae/Aphod

iinaeTribes/Aegialiini/Caelius/Caelius.html


http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aegialiini/Aegialia/Aegialia.html




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aegialiini/Caelius/Caelius.html





Tribe Aphodiini

Genus: Acrossus (Aphodius)

Species: Acrossus rubripennis (Aphodius rubripennis)

Dichotomous Key:

Aphodiini of the United States and Canada,

Key to Acrossus, page 254


Genus: Acrossus (Aphodius)

Species: Acrossus rufipes (Aphodius rufipes)

Dichotomous Key:


Key to Acrossus, page 254

http://typhaeus.chez-alice.fr/galaph.htm

Genus: Agoliinus (Aphodius)


Similar to: Agoliinus leopardus (Aphodius leopardus)

Agoliinus manitobensis (A. manitobensis)


Dichotomous Key:


Key to Agoliinus, Page 125

http://www-


carabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Agoliinus

/Agoliinus.html



http://typhaeus.chez-alice.fr/galaph.htm

http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Agoliinus/Agoliinus.html





Genus: Aphodius

Species: Aphodius fimetarius



Page 106


Key to Aphodius, Page 147


Genus: Blackburneus (Aphodius)

Species: Blackburneus aegrotus (A. campestris)

Similar to: Blackburneus rubeolus (A. rubeolus)

B. lentus (A. lentus) B. stercorosus (A. stercorosus)

Dichotomous Key:

Aphodiini of the United States and Canada

Key to Blackburneus, page 351



http://bugguide.net/node/view/496691/bgpage

Genus: Calamosternus (Aphodius)

Species: Calamosternus granarius (A. granarius)



Page 258






http://bugguide.net/node/view/496691/bgpage



Genus: Chilothorax (Aphodius)

Species: Chilothorax distinctus (Aphodius distinctus)



Page 258




Genus: Colobopterus (Aphodius)

Species: Colobopterus erraticus (Aphodius erraticus)



Page 92




Genus: Dialytellus (moved from Tribe Eupariini)

Species: Dialytellus dialytoides

Similar to: Dialytellus tragicus (D. humeralis)

Dichotomous Key:


Key to Dialytellus, Page 78

http://www-


eoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/A

phodiini/Dialytellus/Dialytellus.html




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Dialytellus/Dialytellus.html





Genus: Dialytes (moved from Tribe Eupariini)

Species: Dialytes truncatus

Similar to: Dialytes striatulus, D. ulkei

Dichotomous Key:


Key to Dialytes, Page 82


Genus: Flaviellus (Aphodius)

Species: Flaviellus phalerioides (A. phaleroides)

Dichotomous Key:


Key to Flaviellus, Page 335

http://www-


carabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Flaviellus

/Flaviellus.html

Genus: Irrasinus (Aphodius)

Species: Irrasinus stupidus (Aphodius stupidus)

Dichotomous Key:


Key to Irrasinus, Page 368




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Flaviellus/Flaviellus.html






Genus: Labarrus (Aphodius)

Species: Labarrus pseudolividus (Aphodius lividus)

Dichotomous Key:


Key to Labarrus, Page 261


Genus: Lechorodius (Aphodius)

Species: picture just representative for genus

Similar to: Lechorodius terminalis (A. terminalis)

Lechorodius lutulentus (A. lutulentus)


Dichotomous Key:


Key to Lechorodius, Page 371

http://www-



phodiini/Lechorodius/Lechorodius.html

Genus: Melinopterus (Aphodius)

Species: Melinopterus femoralis (Aphodius femoralis)

Similar to: Melinopterus prodromus (A. prodromus)

Dichotomous Key:


Key to Melinopterus, Page 320

http://www-



phodiini/Melinopterus/Melinopterus.html



http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Lechorodius/Lechorodius.html




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Melinopterus/Melinopterus.html





Genus: Oscarinus (Aphodius)


Similar to: Oscarinus silvanicus (Aphodius silvanicus)

Oscarinus rusicola (A. ruricola)


Dichotomous Key:


Key to Oscarinus, Page 165

http://www-

museum.unl.edu/research/entomology/Guide/Scarab

aeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes

/Aphodiini/Oscarinus/Oscarinus.html

Genus: Otophorus (Aphodius)

Species: Otophorus haemorrihoidalis

(A. haemorrihoidalis)



Page 89




Genus: Oxyomus

Species: Oxyomus sylvestris (O. silvestris)

(cited in literature)



Page 87

http://www-



phodiini/Oxyomus/Oxyomus.html


http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Oscarinus/Oscarinus.html





http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Oxyomus/Oxyomus.html





Genus: Pardalosus (Aphodius)

Species: picture just representative for the genus

Similar to: Pardalosus serval (Aphodius serval)


Dichotomous Key:


Key to Pardalosus, Page 377


Genus: Planolinellus (Aphodius)

Species: Planolinellus vittatus (Aphodius vittatus)



Page 122



http://www-



phodiini/Planolinellus/Planolinellus.html

Genus: Planolinus (Aphodius)

Species: Planolinus tenellus (Aphodius tenellus)

(Cited in Literature)



Page 117






http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Planolinellus/Planolinellus.html






Genus: Pseudagolius (Aphodius)

Species: Pseudagolius bicolor (Aphodius bicolor)

Dichotomous Key:


Key to Pseudagolius, Page 311


Genus: Stenotothorax (Aphodius)

Species: Stenotothorax badipes (A. badipes)

Dichotomous Key:


Key to Stenotothorax, Page 189

http://www-


carabaeoidea/Scarabaeidae/Aphodiinae/Aphod

iinaeTribes/Aphodiini/Stenotothorax/Stenototho

rax.html

Genus: Strigodius (Aphodius)

Species: Strigodius robinsoni (A. robinsoni)



Page 237



http://www-



phodiini/Strigodius/Strigodius.html



http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Stenotothorax/Stenotothorax.html





http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Aphodiini/Strigodius/Strigodius.html





Genus: Teuchestes (Aphodius)

Species: Teuchestes fossor (Aphodius fossor)



Page 94



http://www.galerie-

insecte.org/galerie/view.php?adr=image%2Fdos6%2F

temp%2Fteuchestes_fosso.jpg

Tribe Didactyliini

Genus: Aidophus (Aphodius)

Species: Aidophus parcus (Aphodius parcus) (Cited in

Literature)



5


Tribe Eupariini

Genus: Aphotaenius (moved from Tribe Psammodiini) Species: Aphotaenius carolinus

Dichotomous Key:

The Scarbaeoid Beetles of Nebraska

Species Description, Page 227

http://www-


eoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/E

upariini/Aphotaenius/Aphotaenius.html


http://www.galerie-insecte.org/galerie/view.php?adr=image%2Fdos6%2Ftemp%2Fteuchestes_fosso.jpg




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Eupariini/Aphotaenius/Aphotaenius.html





Genus: Ataenius

Species: Ataenius strigatus

Similar to: Ataenius apicalis, A. brevis, A. exiguous,

A. fattigi, A. glaseri, A. imbricatus, A. insculptus (cited

in literature), A. ovatulus, A. spretulus, A. wenzelii

***Black/Really Dark Brown color***

Dichotomous Key:


Key to Ataenius, Page 229


Genus: Ataenius

Species: Ataenius gracilis

Similar to: Ataenius alternates (cited in literature),

A. abditus, A. cylindrus, A. glaseri, A. miamii

***Lighter Brown Color***

Dichotomous Key:


Key to Ataenius, Page 229


-Species A. figulator and A. simulator found in Staines but not in current taxonomic classification-

Genus: Pseudataenius (Pseudoataenius)


Similar to: Pseudataenius contorus (cited in literature),

P. walterhorni (cited in literature)




Page 36







Genus: Saprosites (Moved from Tribe Psammodiini) Species: pictures just representative for genus

Similar to: Saprosites ventralis




Page 37

http://www-

museum.unl.edu/research/entomology/Guide/Scarabae

oidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Eup

ariini/Saprosites/Saprosites.html

Tribe Psammodiini

Genus: Psammodius Species: Psammodius basalis

Link Below for Psammodius Description from the

Generic Guide to New World Scarab Beetles

http://www-


eoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/P

sammodiini/Psammodius/Psammodius.html

Genus: Psammodius

Species: Psammodius pierotti (P. sulcicollis)

(cited in literature)



http://www-




http://coleoptera.ksib.pl/kfp/search.php?img=6954


http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Eupariini/Saprosites/Saprosites.html




http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Psammodiini/Psammodius/Psammodius.html










Genus: Psammodius

Species: Psammodius laevipennis



http://www-




http://www.entomologiitaliani.net/public/forum/phpBB3

/viewtopic.php?t=29232

-Species P.bidens and P. interruptus found in Staines but not in current taxonomic classification-

Genus: Rhyssemus

Species: picture just representative for the genus

Similar to: Rhyssemus scaber


Dichotomous Key:


Key to Species of Psammodiini, Page 44


Genus: Pleurophorus Species: Pleurophorus caesus Dichotomous Key:

Species Description in The Scarabaeoid Beetles of Nebraska, Page 244

Link below for addition Species Description

http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Psammodiini/Pleurophorus/Pleurophorus.html






http://www.entomologiitaliani.net/public/forum/phpBB3/viewtopic.php?t=29232

http://www.entomologiitaliani.net/public/forum/phpBB3/viewtopic.php?t=29232







Genus: Pleurophorus

Species: Pleurophorus atlanticus (cited in literature)

Dichotomous Key:

Genus Description in The Scarabaeoid Beetles

of Nebraska, Page 244

http://unibio.unam.mx/irekani/bitstream/123456789/25

622/1/9310.jpg

Cetoniinae

Tribe Cetoniini

Genus: Euphoria

Species: Euphoria areata (Stephanucha areata)


Scarab Beetles of South Carolina, Page 116


Genus: Euphoria

Species: Euphoria flugida

Dichotomous Key:


Key to Euphoria, Page 469



http://unibio.unam.mx/irekani/bitstream/123456789/25622/1/9310.jpg





Genus: Euphoria

Species: Euphoria herbacea

Dichotomous Key:




Genus: Euphoria

Species: Euphoria inda

Dichotomous Key:






Genus: Euphoria

Species: Euphoria sepulchralis

Dichotomous Key:











Tribe Cremastocheilini

Genus: Cremastocheilus

Species: Cremastocheilus canaliculatus

Similar to: Cremastocheilus castaneae, C. variolosus

Dichotomous Key:


Key to Species of Cremastocheilini , Page 118


Genus: Cremastocheilus

Species: Cremastocheilus harrisii

Dichotomous Key:




Genus: Psilocnemis

Species: Psilocnemis leucosticte (cite in literature)

Dichotomous Key:









Tribe Gymnetini

Genus: Cotinis

Species: Cotinis nitida



Page 466


Tribe Trichiini

Genus: Gnorimella

Species: Gnorimella maculosa


The Scarab Beetles of South Carolina

Page 122

http://www.chebucto.ns.ca/environment/NHR/Images

3/Gnorimella_maculosa1.jpg

Genus: Osmoderma

Species: Osmoderma eremicola

Similar to: Osmoderma scabra

Dichotomous Key:


Key to Osmoderma, Page 121

http://www.entomology.umn.edu/museum/links/course

files/JPEG%20images/Coleoptera%20web%20jpeg/P

olyphaga/Scarabaeoidea/Scarabaeidae/Cetoninae/Os

moderma.jpg



http://www.chebucto.ns.ca/environment/NHR/Images3/Gnorimella_maculosa1.jpg

http://www.chebucto.ns.ca/environment/NHR/Images3/Gnorimella_maculosa1.jpg

http://www.entomology.umn.edu/museum/links/coursefiles/JPEG%20images/Coleoptera%20web%20jpeg/Polyphaga/Scarabaeoidea/Scarabaeidae/Cetoninae/Osmoderma.jpg





Genus: Trichiotinus

Species: Trichiotinus affinis

Similar to: Trichiotinus piger

Dichotomous Key:


Key to Trichiotinus, Page 122


Genus: Trichiotinus

Species: Trichiotinus bibens

Dichotomous Key:




Genus: Trichiotinus

Species: Trichiotinus lunulatus

Dichotomous Key:









Genus: Trigonopeltastes

Species: Trigonopeltastes delta


Scarabs Beetles of South Carolina

Page 122


Tribe Valgini

Genus: Valgus

Species: Valgus canaliculatus

Similar to: Valgus squamiger

Dichotomous Key


Key to Valgus, Page 502



Dynastinae

Tribe Cyclocephalini

Genus: Cyclocephala

Species: Cyclocephala borealis

Similar to: Cyclocephala immaculata

Dichotomous Key:


Key to Cyclocephala, Page106








Genus: Dyscinetus

Species: Dyscinetus morator

Similar to: Dyscinetus trachypygus

Dichotomous Key:


Key to Dyscinetus, Page 443


Tribe Dynastini

Genus: Dynastes

Species: Dynastes tityus



Page 113



Tribe Oryctini

Genus: Strategus

Species: Strategus antaeus

Dichtomous Key:

Scarab Beetles from South Carolina

Key to Strategus, Page 113










Genus: Xyloryctes

Species: Xyloryctes jamaicensis



Page 460

Species Description in


Page 112



Tribe Pentodontini

Genus: Aphonus (moved from Tribe Oryctini)

Species: Aphonus castaneus

Dichotomous Key:


Key to Aphonus, Page 111

http://www.pbase.com/tmurray74/image/125304612






Genus: Aphonus (moved from Tribe Oryctini)

Species: Aphonus densicauda

Dichotomous Key: Aphonus Discription in


Page 111


Genus: Euetheola

Species: picture just a representative for genus

Similar to: Euetheola rugiceps




Page 112


Genus: Tomarus(Ligyrus) (moved from Tribe Oryctini)

Species: Tomarus gibbosus

Dichotomous Key:


Key to Tomarus, Page 455







Genus: Tomarus(Ligyrus) (moved from Tribe Oryctini)

Species: Tomarus relictus

Dichotomous Key:


Key to Tomarus, Page 455


Tribe Phileurini

Genus: Phileurus

Species: Phileurus valgus (P. castaneus)

Dichotomous Key:


Key to Phileurus, Page 115

Species Description in The Scarabaeoid

Beetles of Nebraska, Page 463


Melolonthinae

Tribe Diplotaxini

Genus: Diplotaxis

Species: Diplotaxis frondicola

Similar to: Diplotaxis fulva,

D. subcostata (cited in literature)

Dichotomous Key:


Key to Diplotaxis, Page 73







Genus: Diplotaxis

Species: Diplotaxis harperi

Similar to: Diplotaxis atlantis,

D. bidentata (cited in literature), D. liberta,

D. punctatorugosa (cited in literature),

D. sordid, D. tristis

Dichotomous Key:


Key to Diplotaxis, Page 73


Tribe Hopliini

Genus: Hoplia

Species: Hoplia trifasciata

Similar: Hoplia equina, H. modesta

Dichotomous Key:


Key to Hoplia, Page 96


Genus: Hoplia

Species: Hoplia trivialis

Similar to: Hoplia equina

Dichotomous Key:


Key to Hoplia, Page 96







Tribe Macrodactylini

Genus: Dichelonyx

Species: Dichelonyx albicollis

Similar to: Dichelonyx fuscula

Diochotomous Key:


Key to Dichelonyx, Page 94

D. albbicollis-Scarabaeoid Beetles of Nebraska


http://www-

museum.unl.edu/research/entomology/Guide/Scarabaeoi

dea/Scarabaeidae/Melolonthinae/Melolonthinae-Generic-

pages/Macrodactylini/Dichelonyx/Dichelonyx.html

Genus: Dichelonyx

Species: Dichelonyx diluta

Dichotomous Key:




Genus: Dichelonyx

Species: Dichelonyx elongatula (Dichelonyx elongata)

Similar to: Dichelonyx subvittata

Dichotomous Key:





http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Melolonthinae/Melolonthinae-Generic-pages/Macrodactylini/Dichelonyx/Dichelonyx.html







Genus: Macrodactylus

Species: Macrodactylus angustatus

Dichotomous Key:


Key to Macrodactylus, Page 95


Genus: Macrodactylus

Species: Macrodactylus subspinosus

Dichotomous Key:


Key to Macrodactylus, Page 95

Species Description in The Scarabaeoid

Beetles of Nebraska, Page 319


Tribe Melolonthini

Genus: Phyllophaga

Species: Phyllophaga crenulata

Similar to: Phyllophaga aemula

Dichotomous Key:


Key to Phyllophaga, Page 75

P. crenulata Scarabaeoid Beetles of Nebraska








Genus: Phyllophaga

Species: Phyllophaga implicita (cited in literature)

Similar to: Phyllophaga balia, P. ephilida, P. fraterna

P. forsteri, P. fusca, P. futillis, P. hirticula, P. ilicis,

P. inversa, P. marginalis, P. rugosa,

Dichotomous Key:






Genus: Phyllophaga

Species: Phyllophaga tristis

Similar to: Phyllophaga glaberrima (cited in the

literature), P. gracilis, P. prununculina (cited in

literature), P. quercus

Dichotomous Key:








Genus: Phyllophaga

Species: Phyllophaga hornii

Similar to: Phyllophaga anxia, P. diffinis, P. drakii,

P. foxii, P. hirsuta, P. latifrons, P. luctuosa, P. mican,

P. subtonsa, P. sylvatica

Dichotomous Key:





http://www-


eoidea/Scarabaeidae/Melolonthinae/Melolonthinae-

Generic-

pages/Melolonthini/Phyllophaga/Phyllophaga-

species/P-hornii/P-hornii.html

--Could not find species descriptions on P. barda & P. vilifrons--

Genus: Polyphylla

Species: Polyphylla variolosa

Dichotomous Key: Genus Description in


Page 385

Genus Description in the Scarab Beetles of

South Carolina, Page 92



http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Melolonthinae/Melolonthinae-Generic-pages/Melolonthini/Phyllophaga/Phyllophaga-species/P-hornii/P-hornii.html








Tribe Sericini

Genus: Maladera

Species: Maladera castanea



Page 72


Genus: Nipponoserica (Serica) Species: Nipponoserica peregrine For Species information visit the link below

http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Melolonthinae/Melolonthinae-Generic-pages/Sericini/Nipponoserica/Nipponoserica.html

Genus: Serica Species: Serica intermixta Similar to: Serica georgiana, S. mystaca, S. parallela, S. vespertina Dichtomous Key:


Key to Serica, Page 66













Genus: Serica

Species: Serica iricolor

Dichotomous Key:




Genus: Serica

Species: Serica sericea

Similar to: Serica blatchleyi, S. loxia (cited in

literature), S. opposita (cited in literature)

Dichotomous Key:



http://www-


carabaeidae/Melolonthinae/Melolonthinae-Generic-

pages/Sericini/Serica/Serica.html

--Could not find species descriptions on S. atracapilla (atricapilla), S. floridana, & S. imitans—

Rutelinae

Tribe Anomalini

Genus: Anomala

Species: Anomala binotata

Dichotomous Key:

The Scarbaeoid Beetles of Nebraska

Key to Anomala, Page 414

http://bugguide.net/node/view/393759/bgimage 4 black spots 2 on each elytra



http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Melolonthinae/Melolonthinae-Generic-pages/Sericini/Serica/Serica.html






Genus: Anomala

Species: flavipennis

Similar to: Anomala marginata, A. semilivida

Dichotomous Key:



A. flavipennis-The Scarabaeoid Beetles of

Nebraska, Key to Anomala, Page 414


Genus: Anomala

Species: Anomala innuba, A. oblivia

Dichotomous Key:



A. innuba-The Scarabaeoid Beetles of

Nebraska, Key to Anomala, Page 414


Genus: Anomala

Species: Anomala lucicola

Similar to: Anomala umbra

Dichotomous Key:









Genus: Anomala

Species: Anomala orientalis

Dichotomous Key:




Genus: Anomala

Species: Anomala undulata

Dichotomous Key:






Genus: Popillia

Species: Popillia japonica



Page 427


Page 103







Genus: Strigoderma

Species: Strigoderma pygmaea

Similar to: Strigoderma arbicola

Dichotomous Key:


Key to Strigoderma, Page 102

S. arbicola-Scarabaeoid Beetles of Nebraska

Page 429


Tribe Rutelini

Genus: Cotalpa

Species: Cotalpa lanigera

Dichotomous Key:


Key to Cotalpa, Page 405


Key to Tribe Rutelini, Page 103


Genus: Parastasia

Species: Parastasia brevipes

Dichotomous Key:









Genus: Pelidnota

Species: Pelidnota punctata

Dichotomous Key:




Species Description, Page 409


Scarabaeinae

Tribe Ateuchini

Genus: Ateuchus

Species: Ateuchus histeroides

Similar to: A. lecontei (cited in literature)

Dichotomous Key:


Key to Ateuchus, Page 16

A. histeroides-The Scarabaeoid Beetles of

Nebraska Species Description, Page 268


Tribe Canthonini

Genus: Canthon

Species: Canthon vigilans

Similar to: C. chalcites, C. pilularius,

C. (Boreocanthon) probus

Dichotomous Key:


Key to Canthon, Page 283







Genus: Canthon

Species: Canthon viridis

Similar to: C. pilularius

Dichotomous Key:


Key to Canthon, Page 283


Genus: Melanocanthon

Species: Melanocanthon bispinatus



Page 15


Tribe Coprini

Genus: Copris

Species: Copris minutis

Similar to: C. fricator

Dichotomous Key:


Key to Copris, Page 270









Genus: Dichotomius

Species: Dichotomius carolinus



Page 274

Species Description in Scarab Beetles of South

Carolina, Page 17


Tribe Onthophagini

Genus: Onthophagus

Species: Onthophagus concinnus

Dichotomous Key:

Beetle Genus Onthophagus—Howden, Cartwright

Key to Onthophagus, Page 10


Genus: Onthophagus

Species: Onthophagus hecate

Similar to: O. pennsylvanicus, O. striatulus, O. taurus

O. tuberculifrons

Dichotomous Key:













Genus: Onthophagus

Species: Onthophagus nuchicornis

Dichotomous Key:




Genus: Onthophagus

Species: Onthophagus orpheus

Dichotomous Key:






Tribe Phanaeini

Genus: Phanaeus (Moved from Tribe Coprini)

Species: Phanaeus vindex

Dichotomous Key:


Key to Phanaeus, Page276

http://scarabaeoidea.lifedesks.org/pages/1228






TROGIDAE

Troginae

Genus: Omorgus (Trox)

Species: Omorgus (Trox) monachus

Similar to: Omorgus (Trox) asper, O. (Trox) tytus

Dichotomous Key:


Key to Omorgus, Page 92


Key to Species of Troginae (Trogidae), Page 61



Species: Omorgus (Trox) scabrosus

Dichotomous Key:





Species: Omorgus (Trox) suberosus

Dichotomous Key:









Genus: Trox

Species: Trox scaber

Similar to: Trox aequalis, T. affinis, T. capillaris,

T. hamatus, T. laticollis, T. sordidus, T. terrestris,

T. tuberculatus

Dichotomous Key:


Key to Trox, Page 69

T. affinis & T. terrestris- Scarab Beetles of

South Carolina, Key to Troginae (Trogidae),

Page 61


Genus: Trox

Species: Trox spinulosus

Similar to: Trox foveicollis

Dichotomous Key:








Genus: Trox

Species: Trox striatus

Similar to: Trox unistriatus

Dichotomous Key:




Genus: Trox

Species: Trox variolatus

Dichotomous Key:








References

BugGuide. Iowa State University Entomology. http://bugguide.net/node/view/15740 Borowiec, Lech. “Psammodius sulcicollis (Illiger, 1801)”. Iconographia Coleopterorum Poloniae. http://coleoptera.ksib.pl/kfp/search.php?img=6954 Cartwright, O.L. and H.F. Howden. “Scarab Beetles of the Genus Onthophagus Latreille North of Mexico (Coleoptera: Scarabaeidae)”. Proceedings of the United States National Museum, 114(3467): 1-135, 11 figures, 9 plates. Dabry, Julien. “Acrossus rufipes”. Portraits d’éboueurs. Last Visited July 25, 2012. http://typhaeus.chez-alice.fr/images/Acrossus_rufipes.jpg Gordon, R. D. and P. E. Skelley. 2007. A monograph of the Aphodiini inhabiting the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). Memoirs of the American Entomological Institute 79: 580 p. Guzman, David. “Lucanus elaphus”. Flickr from Yahoo. Last Visited July 25, 2012. http://www.flickr.com/photos/entomopixel/5098352100/ Harpootlian, P.J. 2001. Scarab Beetles (Coleoptera: Scarabaeidae) of South Carolina. Biota of South Carolina. Volume 2. Clemson University Public Service Publishing. Jameson, M.L. and B.C. Ratcliffe. “Generic Guide to New World Scarab Beetles”. University of Nebraska State Museum Division of Entomology. http://museum.unl.edu/research/entomology/Guide/Guide-introduction/Guideintro.html Majka, Christopher. “Gnorimella maculosa (Knoch)”. Empty Mirrors Press. Last Visited July 25, 2012. http://www.chebucto.ns.ca/environment/NHR/Gnorimella_maculosa.html Manci, Cosmin-Ovidiu. “Psammodius laevipennis (A. Costa, 1844—Aphodiinae)”. Forum Entomologi Italiani. Murray, Tom. “Aphonus castaneus”. PBase. Last Visited July 25, 2012. http://www.pbase.com/tmurray74/image/125304612 Picture of Pleurophorus atlanticus taken from http://unibio.unam.mx/irekani/bitstream/123456789/25622/1/9310.jpg Last Visited July 25, 2012 Picture of Teuchestes fossor taken from http://www.galerie-insecte.org/galerie/view.php?adr=image%2Fdos6%2Ftemp%2Fteuchestes_fosso.jpg Last Visited July 25, 2012. Price, Dana L. “Phanaeus vindex (MacLeay)”. eScarab LifeDesks. Last Visited July 25, 2012. http://scarabaeoidea.lifedesks.org/pages/1228




http://typhaeus.chez-alice.fr/images/Acrossus_rufipes.jpg


http://museum.unl.edu/research/entomology/Guide/Guide-introduction/Guideintro.html

http://www.chebucto.ns.ca/environment/NHR/Gnorimella_maculosa.html







Ratcliffe, B.C. and M.J. Paulsen. 2008. The Scarabaeoid Beetles of Nebraska (Coleoptera: Scarabaeoidea). University of Nebraska State Museum Bulletin 22: 1-570. Staines, C. L., “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland”. 1984. Maryland Entomologist 2(4): 79-89.


mallory hagadorn dr. dana l. price department of ...faculty.salisbury.edu/~dlprice/su lab page/about...

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