Bull. Kitakyushu Mm. Nat. Hist., 20: 1-15, pis. 1-2. March 31, 2001

Neoschwagerina (Permian fusulinacea) from limestone

pebbles in the Dobaru Formation, the lowest part

of the Lower Cretaceous Wakino Subgroup,

in Kitakyushu City, Japan

Yasuhiro Ota

Kitakyushu Museum and Institute of Natural History,3-6-1, Nishihonmachi, Yahata-higashi-ku,

Kitakyushu 805-0061, JAPAN(Received December 20, 2000)

Abstract Well-preserved Neoschwagerina aff. cheni lisinghaiana Shung with Parafusulinasp. and Chusenella sp. was recovered from limestone pebbles within the basal conglomerate in the Dobaru Formation, the lowest part of the Lower Cretaceous Wakino Subgroup, in Kitakyushu City, Japan. The conglomerate has also yielded well-preservedParafusulina species: Parafusulhia cf. kaerimizensis (Ozawa), Parafusulina sp. A andParafusulina sp. B (Ota, 2000). These occurrences support the interpretation thatthe provenance of this non-crystalline limestone was near the sedimentary basin ofthe Dobaru Formation in the Early Cretaceous time. These limestone pebbles withNeoschwagerina are considered to have been transported from limestone of theNeoschwagerina simplex Zone in the Tethyan region or with correlatives of Middle Permian age.

Introduction

Well-preserved fusulinaceans have recently been found in the basal conglomerate within the Dobaru Formation, the lowest part of the Lower Cretaceous WakinoSubgroup, in Kitakyushu City (Ota, 2000). The conglomerate with fusulinaceansis highly significant for deciphering the provenance of the limestone in KitakyushuCity and for inferring the paleoenvironment and sedimentological environment ofthe Wakino Subgroup in the Early Cretaceous time. The PaleozoicYobuno Group(Ota et al, 1992; Nakae, 1998), one of the Carboniferous to Permian groups in theAkiyoshi Terrane, is widely distributed in Kitakyushu City. However, most limestones in this group generally are crystalline, as are limestones of the Hirao Limestone Plateau, so well-preserved fusulinaceans have not been reported previouslyfrom this group. Ota and Yabumoto (1998) recognized that conglomerates inthe Dobaru and Takatsuo Formations of the Wakino Subgroup contain fusulinaceans derived from the underlying Paleozoic Yobuno Group. Ota (2000) firstreported well-preserved Parafusulina cf. kaerimizensis (Ozawa), Parafusulina sp. A and

2 Yasuhiro Ota

Parafusulina sp. B from the conglomerate in the Dobaru Formation near

Hazegatoge, in the Kokuraminami Ward (Kokura-minami-ku) of Kitakyushu City.Parafusulina cf. kaerimizensis is comparative to Parafusulina kaerimizensis, which wasfirst reported by Ozawa (1925a) from the Akiyoshi Limestone Group, SouthwestJapan. Parafusulina sp. A is closely similar to Parafusulina granumavenae reported

by Toriyama (1975) from the Rat Buri Limestone in Thailand. From the affinitiesof the Parafusulina species, the enclosing limestone cobble with Parafusulina species

is thought to have a close stratigraphic relationship with limestone of theParafusulina kaerimizensis Zone in the Akiyoshi Limestone Group and correlatives of

Middle Permian age.

A newly discovered Neoschwagerina species also occurs at the same locality (Fig.1), and at the same stratigraphic level as the Parafusulina, namely in limestonepebbles within the conglomerate in the Dobaru Formation, the lowest part of theLower Cretaceous Wakino Subgroup (Fig. 2).

Since Yabe (1903) erected Neoschxvagerina, with Neoschwagerina craticulifera(Schwager) as its type species, over one hundred species of Neoschwagerina havebeen reported from the Tethyan region. The genus has been studied by manyauthors, including Hayden (1909), Deprat (1914), Ozawa (1925a, b, 1927),Dunbar and Condra (1927), Lee (1933), Gubler (1935), Anderson (1941),

KITAKYUSHU

Kokuraminami Ward

T^BnfcXVCWW •

Fig. 1. Sample location: HZ001 .(The base map is from 1/25,000 of Tokuriki published by Geographical Survey Institute in

Japan)

HZ001

Neoschwagerina (Permian fusulinacea) from Kitakyushu City, Japan

Ilillil

Neoschwagerina aff. cheni hsinghaiana SHENG

o50m ^

MUDSTONE

SANDOSTONE

CHERT

UMESTONE

Fig. 2. Schematic section of the Dobaru Formation with the fusulinacean limestone pebbles atHazegatoge [Modified from Oia and Yabumoio (1998) and Nakak (1998)].

4 Yasuhiro Ota

Thompson (1948), Yabe (1948), Hanzawa (1954), Miklukho-Maklay (1957,

1958), Kanmera (1957), Toriyama (1958), Minato and Honjo (1959, 1966),

Honjo (1959), Sada (1961, 1963), Leven (1963, 1967, 1980, 1981, 1992, 1993a, b,c, 1997), Hanzawa and Murata (1963), Nogami (1961, 1965), Skinner and

Wilde (1966a, b, 1967), Skinner (1969), Ozawa (1970), Yang (1985), Sheng el al.

(1988), Loeblich and Tappan (1988), Yang and Zheng (1993), Vachard el al.

(1995), Rauser el al (1996), Kobayashi (1997a, b), Zhou and Xie (1997), Zaw

(1999) and Yang and Yancey (2000).

The general features of the species from Kitakyushu City resemble those ofNeoschxoagerina cheni var. fisinghaiana, which was first reported by Sheng (1958) fromChinghai Province, North China. However, its size and number of volutions distinguish it from the latter species. Additional specimens and further studies areneeded to determine the specific name with confidence, so I treat the present species as Neoschwagerina aff. cheni hsinghaiana Sheng herein. According to Sada(1963), Neoschwagerina cheni Sheng represents the early adult stage of Neoschwagerinamargarilae of Deprat (1913), in which case the two species would be conspecific.Whether or not this is true remains unclear, but the present species has some affinities with Neoschwagerina cheni and Neoschwagerina cheni fisinghaiana. These speciesare widespread in the Tethyan region and fusulinaceans showing a great similarityto these two species occur on the Atetsu Limestone Plateau, Japan (Nogami, 1961),and within conglomerates in the northern part of Itsukaichi, Tokyo-to, Japan(Sakagami, 1958), the Maokou Limestone of Chinghai Province, China (Sheng,1958), the Qinghai - Xizang Plateau (Wang el al 1981), South China (Chen, 1956),and in Permian faunas of Pahang, Malaya (Ishii, 1966, Igo, 1966). Neoschwagerinamargarilae Deprat (including the forms of Neoschwagerina cheni sensu Sheng, 1958)occurs in even more localities within the Tethyan region. Therefore, the provenance of the Kitakyushu Citylimestone with Neoschxoagerina is verysimilar to that ofthe limestones of these many localities.

The Neoschxoagerina species from Kitakyushu City is characterized by having theprimitive morphological features of Neoschwagerina, including a small shell andrather thick spirotheca, septa and septula, having poorly developed axial and transverse septula, and by having its first to second volutions lenticular in shape as inMaklaya of Kanmera and Toriyama (1968). Therefore, the present species evidently is intermediate in form between Neoschwagerina and Maklaya, and seeminglyrepresents the most primitive form of Neoschwagerina. Furthermore, this specieshas more primitive features than Neoschwagerina craticulifera sensu Leven (1993a) orsensu Depart (1914), and was found together with Parafusulina sp. and Chusenellasp. at Hazegatoge, Kitakyushu City. Accordingly, the original stratigraphic level ofthe limestone with those species is considered to be the Neoschwagerina simplex Zonein the Tethyan region or its correlativesof Middle Permian age (Table 1).

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6 Yasuhiro Ota

Below is a description of Neoschxoagerina aff. cheni lisingfiaiana from Kitakyushu City.

Repository: All specimens used in this study are deposited in the Kitakyushu

Museum and Institute of Natural History, with the designation of KMNH IvP for theregistration numbers of the specimens. Sample numbers are given in brackets.

Acknowledgments: I thank Dr. Masamichi Ota, Counselor, Kitakyushu Museum and

Institute of Natural History, for invaluable suggestions and commentary throughoutthe course of this study. I also thank Dr. Kimiyoshi Sada, Professor Emeritus,Hiroshima University, who provided abundant helpful advice and suggestions during this study. I wish to thank Dr. Tsugio Shuto, Professor Emeritus, Kyushu University, who provided helpful suggestions on the type script. I specially thank Dr.Louie Marincovich, Earth Science Editing, who kindly reviewed the language in

my manuscript.

Systematic Paleontology

Superfamily FUSULINACEA von Moli.er, 1878Family NEOSCHWAGERINIDAE Dunbar and Condra, 1927

Subfamily NEOSCHWAGERININAE Dunbar and Condra, 1927Genus Neoschwagerina Yabe, 1903

Type-species: Schxoagerina craliculifera Schwager, 1883. F. von Richthofen'sCHINA, 4, p. 140-143, pi. 18, fig. 15-25. [Thompson, 1948 designated the figured specimen (Fig. 17, pi. 18, Schwager, 1883) as the holotype of Schxoagerinacraliculifera.]

Generic diagnosis: The genus Neoschwagerina has a medium to large and fusiformshell, with bluntly pointed to narrowly rounded poles. The spirotheca is composedof a tectum and keriotheca. The septulum occurs in two directions, i.e., in thedirection of the axis of coiling (axial septula) and in the direction of coiling (transverse or spiral septula). Primary transverse septula are contact with the paracho-mata only adjacent to the septa in primitive forms, but extend to the top of theparachomata in advanced forms. Secondary transverse septula are generallyabsent. Parachomata are broad and short in the primitive species and becomerelatively narrower and higher in the more-evolved species. There are foraminathroughout the shell.

Remarks: Leven (1993a, b) restudied Neoschwagerina craliculifera (Schwager, 1883)which is the type species of Neoschxoagerina. Regarding Neoschwagerina craliculiferasensu Deprat (1914), which is widely adopted as the morphotype of the typespecies of Neoschxoagerina, he pointed that Neoschxoagerina craliculifera sensu Deprat

Neoschxoagerina (Permian fusulinacea) from Kitakyushu City, Japan 7

(1914) is a primitive Neoschxoagerina because of its less-developed axial andtransverse (spiral) septula, and its lower stratigraphic level than those of the originalfigures of Schxoagerina craliculifera of Schwager (1883), one of which was chosenas the holotype by Thompson (1948). Therefore, he proposed a new name,Neoschxoagerina deprati nom. nov. Leven (1993a) for Neoschwagerina craliculiferasensu Depart (1914). Acceding with Leven (1993a), the original Neoschwagerinacraliculifera (Schwager, 1883) is a highly developed form among neoschwagerinidsand is transitional between Neoschwagerina haydeni Dutkevich and Khabakov andthe primitive Yabeina. Furthermore, Leven (1993a) mentioned that the form

specified by Nogami (1965) as the neotype of Neoschxoagerina craliculifera is ahighly developed neoschwagerinid and is fully comparable to the holotype ofNeoschxoagerina craliculifera (Schwager, 1883), but it is similar to the form in theinner nine or ten volutions of Neoschwagerina haydeni Dutkevich and Khabakov.Therefore, some specimens described and illustrated by Nogami, including theneotype, represent an early ontogenetic stage of ontogeny of Neoschxoagerina haydeniDutkevich and Khabakov, and it is questionable to synonymize it with the holotype of Neoschxoagerina craliculifera (Schwager, 1883). Toriyama and Kanmera(1979) also mentioned that Neoschxoagerina craliculifera sensu Nogami (1965) is considerablydifferent from the Neoschxoagerina craliculifera (Schwager, 1883,pi. 18, fig.17) designated as the lectotype by Thompson (1948), and they proposed the newspecific name of Neoschxoagerina schwageri with the type species of Neoschwagerinacraliculifera (Schwager) of Nogami (1965, pi. 9. fig. 9).

Manyauthors have discussed the evolution of the subfamilyNeoschwagerininaeand two main lineages are recognized: the first lineage is Misellina - Shengella -Cancellina, and the second is Misellina - Maklaya - Neoschxoagerina. Thompson(1948) and Kanmera (1957) treated derivatives of Neoschxoagerina as theNeoschxoagerina - Yabeina - Lepidolina lineage, and Leven (1993b) treated them asthe two lineages Neoschxoagerina - Yabeina and Neoschxoagerina - Colania. Ozawa(1970) treated the lineage Cancellina - Colania - Lepidolina as a separate stock.Yang (1985) treated Lepidolina as a synonym of Yabeina and proposed theNeoschxoagerina stock with a branch of Yabeina (Colania was also treated as a

synonym of Yabeina and some species of Colania were assigned to Yabeina orNeoschxoagerina).

The important criteria for distinguishing between these genera are asfollows. Misellina differs from Shengella and Maklaya by the absence of septula.Shengella differs from Cancellina by its primitive and imperfect septula and lack ofaxial septula. Maklaya can be discriminated from Cancellina and Shengella by itssubspherical to ellipsoidal shell and relatively thicker spirotheca and septa, andfrom Neoschxoagerina by the absence of axial septula and its poorlydeveloped transverse septula and smaller shell. Cancellina is distinguished from Neoschxoagerina byhaving relatively thinner spirotheca and septa, thinner and shorter septula, and the

8 Yasuhiro Ota

narrower parachomata that are in contact with the transverse septula. The validityof Colania and Lejndolina has been questioned, but Colania may be a descendant ofNeoschxoagerina (Lee, 1933, Leven, 1993b) or Cancellina (Ozawa, 1970), and evolvedthrough Yabeina to Lepidolina (Lee, 1933) or to lepidolina (Ozawa, 1970). Colaniais intermediate in shell structure between Neoschwagerina and Yabeina, and its inner

volutions with rather thick spirotheca composed of a tectum and keriothecaresembles closely Neoschwagerina. Its outer ones with a thin and dense layer of atectum and keriotheca are similar to those of Yabeina. Neoschxoagerina can be distin

guished from Yabeina by its smaller shell, fewer volutions, lesser development oftransverse and axial septula and thicker spirotheca. Lepidolina is a highly evolvedfusulinacean with a large and highly inflated fusiform shape, and has a much thinner spirotheca, septula lacking alveoli, and highly developed secondary septula, sothat the differences between Lepidolina and Neoschxoagerina are clear.

Neoschwagerina aff. cheni fisinghaiana Sheng (1958)

Compare:1958. Neoschxoagerina cheni fisinghaiana Sheng. Acta Palaeontologica Sinica, 6(3),p. 288, 289, pi. 2, figs. 5, 6, pi. 3, figs. 4, 5.

Material: Axial sections: KMNH IvP 400, 351 [HZ00100P-0001-1], KMNH IvP 400,

352 [HZ00100P-0002-1], KMNH IvP 400, 353 [HZ00100P-0002-2] and KMNH IvP400, 354 [HZ00100P-0007-1]. Sagittal sections: KMNH IvP 400,355 [HZ00100Q-0006-1] and KMNH IvP 400,356 [HZ001000^0006-2].

Description: The shell of Neoschxoagerina aff. cheni var. fisinghaiana is medium in sizeand fusiform, with a straight coiling axis and bluntly pointed polar ends. Lateralslopes are slightly concave. Two mature specimens (HZ00100P-0001-1 andHZ00100P-0002-1) having thirteen and a half volutions are 4.00 to 4.05 mm longand 1.95+ to 2.75 mm wide. The form ratio ranges from 1.45 to 2.08 (specimenHZ00100P-0002-1 is slightly depressed). The average form ratio of the two specimens is 1.77. The other two axial specimens (HZ00100P-0002-2 and HZ00100P-0007-1) have almostequal to the above mature specimen in general shape. Thesespecimens are thought to be incomplete and immature specimens of the presentspecies, and data related to this are as follows. Specimen HZ00100P-0002-2 hastwelve and a half volutions, is 3.35+ mm long, 2.40+ mm wide, and has a form ratioof 1.40±. Specimen HZ00100P-0007-1 has nine volutions and is 2.05 mm long and1.40 mm wide, with a form ratio of 1.47. In the present species, the inner first tosecond or third volutions are discoidal and coil slightly askewto later volutions, andsucceeding volutions have almost the same axial profile as the mature shell. Theform ratios of the first and second volutions of specimen HZ00100P-0001-1 are 0.60

Neoschwagerina (Permian fusulinacea) from Kitakyushu City,Japan 9

and 0.50, respectively. The form ratios of the first and second volutions in

specimen HZ00100P-0002-1 are 0.87 and 0.80, respectively. Average ratios of thehalf-length to the radius vector of the third to thirteenth and a half volutions fortwo mature specimens are 2.15, 2.39, 2.40, 2.29, 2.14, 2.21, 2.07, 2.19, 2.04, 1.86,1.70, and 1.59, respectively. The proloculus is minute and spherical with anoutside diameter of 0.04 mm, and its wall is seemingly homogeneous, measuring0.010 mm thick. The volutions, except for the inner two, are closely and regularlycoiled and expand gradually. Average radius vectors of the third to thirteenth anda half volutions for two mature specimens are 0.11, 0.15, 0.20, 0.27, 0.37, 0.47, 0.60,0.73, 0.87, 1.04, 1.20 and 1.33 mm, respectively. Unfortunately, the detailed structure of the spirotheca can not be seen owing to poor preservation, but the twolayers composed of the tectum and the rather thick layer are visible. It is difficultto confirm the exact thickness of the spirotheca, but average thicknesses for the firstto thirteenth and a half volutions for two mature specimens are about 0.010, 0.010,0.015, 0.020, 0.020, 0.020, 0.030, 0.035, 0.030, 0.050, 0.050, 0.045, 0.040 and 0.035

mm, respectively. The septa are somewhat thick and unfluted. The short butbroad fan-shaped primary transverse septula are well developed throughout theshell and extend down into the space between each pair of foramina. Theparachomata are relatively large and well developed throughout the growth of theshell. They generally connect with the primary transverse septula at about two-thirds of the height of the chamber and are fused to each other. The secondarytransverseseptula can not be observed, even in the outer volutions. Sagittal sectionsare poorly preserved, and it is difficult to recognize the detailed structure of eachchamber. Axial septula are poorly developed, short and rare, although at least oneaxial septulum per chamber can be observed sporadically in some outer volutions.

Remarks: The present species is similar to Neoschwagerina cheni fisinghaiana Shengfrom limestone of Chinghai Province in the upper Yangtze Valley, China.However, it differs from the latter species in that the size is smaller, the number ofvolutions is fewer, the spirotheca and septula are thicker, and the axial septula arepoorly developed. Neoschxoagerina craliculifera sensu Leven resembles the presentspecies in some general features. However, the former has short and wide secondary transverse septula, whereas the latter lacks secondary transverse septula.Neoschxoagerina craliculifera sensu Deprat (1914) is also similar to the presentspecies, but the former has a larger shell with numerous volutions. As alreadypointed out by Sada (1963), Neoschxoagerina cheni Sheng, 1958 is quite similarto Neoschxoagerina margarilae of Deprat (1913) and he treated the species asconspecific. The present species resembles Neoschxoagerina cheni sensu Sheng(1958) or Neoschxoagerina margarilae Deprat, 1913 in some features, but the formerspecies differs from the latter one by having a smaller shell and somewhat thickerspirotheca, septa and septula, and having both transverse and axial septula poorly

10 Yasuhiro Ota

developed. The present species can be easily distinguished from Neoschxoagerinamargarilae Deprat, 1913 by the presence or absence of rudimentary secondary transverse septula.

In general, the following evolutionary trends of the Neoschwagerininae are recognized: the shell became larger and the number of volution increased; thespirotheca, septa and septula became thinner; the secondary transverse and axialseptula appeared in progressively earlier volutions as the shell developed; theparachomata generally become slender (e.g. Kanmera, 1957, Ozawa, 1970). Inlight of these evolutionary trends, the Neoschxoagerina aff. cheni fisinghaiana Shengfrom Kitakyushu City is considered to be a primitive species, due to its small shellwith few volutions, its rather thick spirotheca and primary transverse septula, theabsence of secondary transverse septula, and its very slightly developed axialseptula. In addition, the present species is characterized by having discoidalvolutions in the inner one to two volutions that are askew to later volutions. These

characteristics are present in species assigned to Maklaya Kanmera and Toriyama(1968). Therefore, the present species is considered to be intermediate in structure between Maklaya and Neoschwagerina and to be the most primitive form ofNeoschwagerina.

Occurrence: Limestone pebbles in conglomerate from the Lower CretaceousDobaru Formation of Hazegatoge in Kitakyushu City. Fossils associated with it areParafusulinasp. and Chusenella sp.

Table 2. Measurements (in Millimeters) of Neoschwagerina aff. chenifisinghaiana Sheng

Sample Kumbor L. W. S. Prol.

D.

Prol.

Wall

BZ00100P-

0001-1

4.00 2.75 1.45 0.04 0.010

BX00100P-

0002-1

4.05 1.95+ 2.08- 0.04 0.010

Max.

Min.

Avar.

4.05

4.00

4.03

2.75

1.95+

2.35+

1.45

2.08-

1.77-

0.04

0.04

0.04

0.010

0.010

0.010

BX00100P-

0002-2

3.35+ 2.40+ 1.401 0.05 0.010

BX00100P-

0007-1

2.05 1.40 1.47 0.04 0.010

Neoschwagerina (Permian fusulinacea) from Kitakyushu City,Japan 11

Table 2 (continued).

Radius vector (mm.)

1 2 3 4 5 6 7 8 9 10 11 12 13 14

HZ00100P- 0.05 0.07 0.13 0.17 0.23 0.30 0.42 0.52 0.65 0.80 0.95 1.11 1.28 1.42

0001-1

azooioop- 0.05 0.06 0.08 0.12 0.17 0.24 0.32 0.42 0.54 0.65 0.78 0.97 1.11 1.23

0002-1

Knx. 0.05 0.07 0.13 0.17 0.23 0.30 0.42 0.52 0.65 0.80 0.95 1.11 1.28 1.42

Kin. 0.05 0.06 0.08 0.12 0.17 0.24 0.32 0.42 0.54 0.65 0.78 0.97 1.11 1.23

Aver. 0.05 0.07 0.11 0.15 0.20 0.27 0.37 0.47 0.60 0.73 0.87 1.04 1.20 1.33

1 2 3 4 5 6 7 e 9 10 11 12 13 14

BZ00100P- 0.03 0.05 0.08 0.13 0.18 0.27 0.35 0.45 0.S8 0.73 0.91 1.09 1.17

0002-2

BZ00100P- 0.03 0.07 0.12 0.16 0.22 0.33 0.38 0.52 0.65

0007-1

Ratio of Hl./Rv.

1 2 3 4 5 6 7 8 9 10 11 12 13 14

BZ00100P- 1.00- 1.00- 1.54 1.94 1.97 2.00 1.83 1.98 1.77 1.98 1.72 1.67 1.52 1.48

0001-1

BX00100P- 1.00- 2.00- 2.75 2.83 2.82 2.58 2.44 2.43 2.37 2.40 2.35 2.05 1.87 1.69

0002-1

(tax. 1.00- 2.00- 2.75 2.83 2.82 2.58 2.44 2.43 2.37 2.40 2.35 2.05 1.87 1.69

Kin. 1.00- 1.00- 1.54 1.94 1.97 2.00 1.83 1.98 1.77 1.98 1.72 1.67 1.52 1.48

Aver. 1.00- 1.50- 2.15 2.39 2.40 2.29 2.14 2.21 2.07 2.19 2.04 1.86 1.70 1.59

1 2 3 4 5 6 7 8 9 10 11 12 13 14

BX00100P- 1.67? 2.007 2.50 2.39 2.61 2.15 2.23 2.18 1.98 1.75 1.65 1.56 1.56

0002-2

BX00100P- 1.00 1.00 1.33 1.88 1.82 1.67 1.92 1.79 1.66

0007-1

Thickness of spirotheca (mm.)

1 2 3 4 5 6 7 8 9 10 11 12 13 14

axooioop- 0..010 0,.010 0..010 0..020 0.,020 0.020 0..030 0.040 0..020 0.050 0.050 0.050 0.050 0.040

0001-1

bzooioop- 0..010 0,.010 0..020 0..020 0,.020 0.020 0.,030 0.030 0..040 0.050 0.050 0.040 0.030 0.030

0002-1

Max. 0..010 0..010 0..020 0..020 0,.020 O.020 0.,030 0.040 O..040 0.050 0.050 O.OSO 0.050 0.040

Hill. 0..010 0..010 0..010 0..020 0..020 0.020 0.,030 0.030 0,.020 0.050 0.050 0.040 0.030 0.030

Aver. 0..010 0..010 0..015 0..020 0..020 0.020 0,,030 0.035 0,.030 0.050 0.050 0.045 0.040 0.035

1 2 3 4 5 6 7 8 9 10 11 12 13 14

BZ00100P- 0..010 0..010 0..010 0..020 0..020 0.020 0..030 0.040 0..020 0.080 0.07? 0.070 0.070 0.040

0001-1

BX00100P- 0..010 0..010 0,.020 0..020 0.,020 0.020 0.,030 0.030 0. 040 0.057 0.050 0.040 0.030 0.030

0002-1

Septal count

BXOOlOOq- - - - 14? 19 20? 32 35 38 41? 44?0006-1

BXOOlOOq- - - 5? 15 15? 21? 22 28 31 36? 390006-2

12 Yasuhiro Ota

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Chine et d'Indochine. Mem. Serv. Geol. de I'lndochine, 3, Fasc. I, EtudedesFusulinidesduJapon deChine et d'Ifidochine el classification des calcaires afusulinesIII. Mem.: 1-45, pis. 1-8.

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Neoschwagerina (Permian fusulinacea) from limestone

pebbles in the Dobaru Formation, the lowest partof the Lower Cretaceous Wakino Subgroup,

in Kitakyushu City, Japan

Yasuhiro Ota

Plates 1-2

Explanation of Plate 1

Figs. 1-6. Neoschxoagerina aff'. cheni fisinghaiana Sheng.

1-4. Axial sections: la, KMNH IvP 400, 351 (HZ00100P-0001-1),

x20; lb, Enlarged part of the same specimens as Fig. la,x40; 2, KMNH IvP 400, 352 (HZ00100P-0002-1), x20; 3,

KMNH IvP 400, 353 (HZ00100P-0002-2), x20 and 4, KMNH

IvP 400, 354 (HZ00100P-0007-1), x20.

5, 6: Sagittal sections: 5, KMNH IvP 400,355 (HZ00100Q0006-1),x20 and 6, KMNH IvP 400, 356 (HZ00100Q0006-2), x20.

Ota, Y. Neoschwagerina (Permian fusulinacea) from Kitakyushu City,Japan Plate I

•, • ••••-

Explanation of Plate 2

1. Fragment of Chusenella sp., KMNH IvP 400, 357 (HZOOlOOP-0014-1),x20.

2. Tangential section of Parafusulina sp., KMNH IvP 400, 358(HZ00100P-0004-l),xlO.

Ota, Y. Neoschwagerina (Permian fusulinacea) from Kitakyushu City,Japan Plate 2