phylloscopus plumbeitarsus - jst

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膁腺腯腾腳腰腮 Phylloscopus plumbeitarsus 膽臅腠腐腔腩膒臋 膸臂臊膩The Records of Two-barred Greenish Warbler Phylloscopus plumbeitarsus in Japan Yoshiki WatabeAbstract. Two-barred Greenish Warblers Phylloscopus plumbeitarsus were observed and photographed on Hegura Island (375105.1N, 1365506.2E), Ishikawa Prefec- ture, on three years: from 24th to 25th May in 2000 (3 individuals), on 20th May in 2001 (1 individual) and from 24th to 25th May in 2005 (3 or more individuals). They were distinguished from similar species (the Arctic Warbler P. borealis, Green Leaf-warbler P. nitidus, Greenish Warbler P. trochiloides, Pale-legged Leaf-warbler P. tenellipes, Sakhalin Leaf-warbler P. borealoides and the Large-billed Leaf-warbler P. magnirostris) by di#er- ences in the shape of the supercilium and wing bars, in the color of the legs, bill, and tip of the outer tail feathers, in their call, and in the wing structure. I also summarized the records of P. plumbeitarsus in Japan based on the data from the literature and personal communication. Eleven records (15 individuals or more) were obtained from 13th September 1998 to 29th May 2009 and seven records (11 individuals or more) occurred in mid and late May in 1999 to 2009. Key words: Two-barred Greenish Warbler Phylloscopus plumbeitarsus, Hegura Island. ῏ῌῑῌῐ膁腺腯腾腳腰腮 Phylloscopus plumbeitarsus臌膰膹膁腺腯腾腳腰腮腱膃腄腼 腣腚腢膪膨Phylloscopus trochiloides 腱膃腄腼腢膂腄膄腸腻腑腧 膺腭腴腭膀膅腲膃腷腽腸腹膜膘腬膧腨膈腤膴膉腟膵膋膼腭腴腭腠膚腓臃臁腘腩腱膃腄腼 腞腌腩腕腡腱膃腄腼腢膊膨腟腗腝膇腫腪腝6 腜腡膆膨腒腕腪腣腞腠膒膡腖腪腝腍腩腒 (Tic- ehurst 1938, Baker 1997, Clements 2007)腕腡腎腛 nitidus, trochiloides, plumbeitarsus 膻臉膨 腟腗腝膇腫腪腩膫膞腦腌腨腙腡膫膞腢 P. trochiloides 腡膵腠 viridanus, ludlowi, trochiloides, obscuratus 4 膆膨腒膑腥腧腪腩 (Williamson 1967, Dickinson 2003, del Hoyo et al. 2006)臅臄 膟腞腢腕腡膵腞膳臈腡膺腭腴腭膴膉腠膚腓臃臁腘腩plumbeitarsus 腟膛腏腧腪腩膙膲腬膮膯 膖腡臌膰膹腞 2000 膾腑腧 2005 膾腠腫腚腨膭膌膐膢腗腚腡腞臄膟腘腩plumbeitarsus 膠膔 Report Received 7 May 2010. 膱臆膘膥臉膏膓膬臄腵膅腶腄腁腋2520236 膱臆膘膥膵膋膕臀膤膗 1 膶臇 3 41 Sagamihara City Environmental Information Center, 1341 Fujimi, Chuo-ku, Sagamihara-shi, Kanagawa 2520236, Japan. 膣膍膷膎膦 (J. Yamashina Inst. Ornithol.), 42: 164174, 2011 Yamashina Institute for Ornithology 164

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Page 1: Phylloscopus plumbeitarsus - JST

������� Phylloscopus plumbeitarsus�� ����

�����

The Records of Two-barred Greenish Warbler Phylloscopus

plumbeitarsus in Japan

Yoshiki Watabe�

Abstract. Two-barred Greenish Warblers Phylloscopus plumbeitarsus were observed

and photographed on Hegura Island (37�51�05.1�N, 136�55�06.2�E), Ishikawa Prefec-

ture, on three years: from 24th to 25th May in 2000 (3 individuals), on 20th May in 2001

(1 individual) and from 24th to 25th May in 2005 (3 or more individuals). They were

distinguished from similar species (the Arctic Warbler P. borealis, Green Leaf-warbler P.

nitidus, Greenish Warbler P. trochiloides, Pale-legged Leaf-warbler P. tenellipes, Sakhalin

Leaf-warbler P. borealoides and the Large-billed Leaf-warbler P. magnirostris) by di#er-

ences in the shape of the supercilium and wing bars, in the color of the legs, bill, and tip

of the outer tail feathers, in their call, and in the wing structure. I also summarized the

records of P. plumbeitarsus in Japan based on the data from the literature and personal

communication. Eleven records (15 individuals or more) were obtained from 13th

September 1998 to 29th May 2009 and seven records (11 individuals or more) occurred in

mid and late May in 1999 to 2009.

Key words: Two-barred Greenish Warbler Phylloscopus plumbeitarsus, Hegura

Island.

������ ������� Phylloscopus plumbeitarsus� ����

�������� !" #$%&'() Phylloscopus trochiloides� !"&� *!+,-./0121� 345 � 67,89:;<=>?@ABCDEF121�GHIJK�� !"LM�N O� !"&P(BQRSTUR� 6VW(XOU$L��YZUR[�X (Tic-

ehurst 1938, Baker 1997, Clements 2007)� O\] nitidus, trochiloides, plumbeitarsus&� ^_(BQRSTU�`abM=� c`a& P. trochiloidesC� viridanus, ludlowi, trochiloides,

obscuratus 4W(Xde/U� (Williamson 1967, Dickinson 2003, del Hoyo et al. 2006)N �fgL&� OCLhi0121@A�GHIJK�� plumbeitarsusBjk/U�lm;nop���L 2000q./ 2005q�T%=rstuQ%LfgK�N plumbeitarsus&� vw

Report �� �

Received 7 May 2010.

� xy:z_{|}f~4�!� �252�0236 xy:zCD���� 1�� 3� 41��� Sagamihara City Environmental Information Center, 1�3�41 Fujimi, Chuo-ku, Sagamihara-shi, Kanagawa

252�0236, Japan.

����� (J. Yamashina Inst. Ornithol.), 42: 164�174, 2011 �Yamashina Institute for Ornithology

164

Page 2: Phylloscopus plumbeitarsus - JST

��������� (Irwin 2001, 2005)����� ����������� ������ Two-barred Greenish Warbler P. plumbeitarsus ����������� ��� �!"#�$��� %&'()*+,-./01������� 23�4 5 2000� 6720028� 9: (1941)#����; (1942)�<=�� >?� P. plumbeitarsus 2�@� P. t.

plumbeitarsus8 �� 2A�8 ��)*+,-./��B��C�DE��������F=��0)*+,-./���� G�� plumbeitarsus0���������� nit-

idus, trochiloides��������H���������� 48 km��E 37I51J05.1K�D 136I55J06.2K 2LMN��8 �O�� 2PQ�R�P�S� TP�U!VW XY8� "Z� 0.552 km2� #[ 12.4 m�$%�\ ]���� 2&�^_` _`a 20028���'�b�(c�)*d� �e]f�gh�i+�� �5b��jk�l�m� n�o�p���.qrs Pinus thunbergii�t,������u��-v�wx� yz�� 2000.� 2001.� 2005.� 5{�|/�}B���010~���2��� �� ������3��|/0G��� ������3��4m����}B��2���

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)*+,-./ Phylloscopus plumbeitarsus�����¹�|/ 165

Page 3: Phylloscopus plumbeitarsus - JST

�� 3 2005� 5�2005� 5 � 23��� 27��������� 24� 1 ��� 25� 3 �������

��� �������� �24� !"#$%� �24�� 25� &'�� 24� �()*�+,�������-. 15 m ������/0+,12�� ��� 24�! 25� �()*�+,�"#$%��� .3�-.� 2.5 m� ��� 50% �� ��45� �6 �789� Artemisia sp., :;� Polygonaceae� <=� Carex sp. >?�@�) 1 m �.+�������� 100% &'�� ��A��BCD� 5 m &'�� �E� 24�)�F� 25�)G,�� ��HI��J&'�� K�DL�M��N� �O� 24� 8 D 55�)P�P����O7 m/s &'��

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����f5 2�^_`ab�cgF��� h+�'�)� Q� 2 ����f5 �iU4j()2>�'��k� lm+��'mF���>�'�� ��� �n1���!�2o,���pqrs">t\u�vw�#\�x y\u�>5� z{|cab�c P. cor-

onatus �}4!#\u)~>5���,�F�\ ���,�F�� Q� 3 � 1 �����)�o� �\u�vw1��4��)W'�� ���S$%����S!�U}1����&\ ��� Q� 1 ��'(��)y\ ��4��p2�� )*�U}1����qrs">y&\ ���� )*�Y���+�)�~WF� ,gF��)p'!p-)�2��)�5� ,���.f�Bp-)�2��p*�+,�)� ,gFM�p'!p-)�2���*�+,>�'�� ��� ^_`ab�c�g� /�(������ 0�,4�!�>5�M�.f����121���)*)�f�12�)� 1��12>�'�� )*�����\ ��4�!pW'�� �����\�,gFM��5� ,gF����,!qr2�-��3�W4�!)����4���� ��,�S  ��Y�y&\(�)W4�!)����4����5�[���42#\ �Q� 1 ! 3 ����\u�vw�#\ �Q� 2 &'�� �¡6!

l¡6�¢£�&\�� ¤51�3> 2��5v�Y0�12�� �� 2��5v�^_`ab�cgF��� -)�5� ¥�¦ab�c P. inornatus §¨©ª«ab�c P. proregulus

�e�12��  7���[��¥�¦ab�c§¨©ª«ab�c�g�>¬\(��*�+,o� s" ���� $�qr­$�&\()>�'�� 5[��(����®¯�� �S��°�8�� X1�l¡6¢£ &\���¬\()WF� l¡69±( ²U4l¡6¢£&\(�:+���B³´� 1/3 �� l¡6¢£&\(�Bp:2(���;�<µgFp:5���� ��� l¡6¢£�&\(�³= p = p¶�� ³=�-)��'��B:K7���Q� 3 ��� (Fig. 3C) g4!� ´�� 7>,� P7 �� P6 ! P8 �·,gFC�'�� ·��k� ¸7��! 7���� 7>�K7��)?±�1���� P9 �¢£�� Q� 3 � 2005� 5 � 24�¹º� 1>��� (Fig. 3C) �� P3 ¢£.f W4g� ���� ³����4 7��!·���?±�124K7���:+���� ^_`ab�c �i�1h+5� ����»¼)½@&'�Q� 1 ! 3 �� 1/2 �� 2/3A�&'�� P10 �

B ( � -166

Page 4: Phylloscopus plumbeitarsus - JST

� 1. 2000�5�25������ ������������ Phylloscopus plum-

beitarsus.

Fig. 1. Two-barred Greenish War-

bler Phylloscopus plumbei-

tarsus photographedonHe-

gura Island, Ishikawa Pre-

fecture, 25th May, 2000.

� 2. 2001�5�20������ ������������ Phylloscopus plum-

beitarsus.

Fig. 2. Two-barred Greenish War-

bler Phylloscopus plum-

beitarsus photographed on

Hegura Island, Ishikawa

Prefecture, 20th May,

2001.

� 3. 2005�5�24� (C)�25�(A, B)����� ������������Phylloscopus plumbeitarsus.

Fig. 3. Two-barred Greenish Warbler Phyllosco-

pus plumbeitarsus photographed on

Hegura Island, Ishikawa Prefecture,

24th (C) and 25th (A, B) May, 2005.

������� Phylloscopus plumbeitarsus������� ! 167

Page 5: Phylloscopus plumbeitarsus - JST

����������� �������� P10�� ������������������� P6�� P8 �� 3������� !"#$ %&����'�� (�)*+����,-��.����� /0�12*+ 3��4"%&567' (Fig. 3B) 8� ��+$�+�9$:;< ��=�������������>'?1 @��A0 B��CDEF0�G� 2�HI�� J�=� G� 1$ 3��HI��=?���JK�JA0� 3��L�� ������������M=N �1����O�4� PQRST �� PQRUST $�#�"V� W�XYD�Motacilla alba�O�4�ZZ[?2? G� 3��\] Accipiter gularis�^_L�`��J�a4J PQR�T $2bV5�'��c�����������d�� G� 3�HI�efC gh�i�jkl ��Lm n�`Zn�o�Ipqr�s"$4�tu�ps"#$ !=� 8�9��`��ps"#$�J�=�� v��� wx�yz'?{� O@�� 1 m|B�}�"#$�J�=��

������#L��HI� ���������K� ~�$*0 c� ���������['I@! 9�.�J�0�p���"0� IB! 9�.�J#0� ��J$�$��� !� � 1�=?2? � 9����#0% J�=�#$�� ��������Phylloscopus�.�$���L" (Baker 1997, del Hoyo et al. 2006)� ������������b� �p��I@! 9�.���0�p���"0�&�� J B�'p��IB! 9�.��#0� 2(���J)� !"#$�����������������U���;L��� ��S����� P. tenellipes ������ P. borealoides W������� P. magnirostris �2�L�$���L" (del Hoyo et al. 2006)��������� $�� *  +��h¡�.,�!"#$ $� *  �¢%�£'J2#$ (�)��-¤'?2"��)��� �����"(�)��_���¥2./�!"#$ �� *  /2�0�p���¦0 /§�0 �¦0 /200J;$1¨�L"0� /20©�!"#$ O�4 ‘drzt’, ‘drz-it’, ‘dzrt’, ‘drrt’, ªF«¬ªFF«¬ ªSR«¬ J;$1G�L"#$�� gTicehurst 1938, Alstrom, & Olsson 1987, Harris

et al. 1989, Svensson 1992, Leader 1993, Baker 1997, Beaman & Madge 1998, Svensson et al. 1999&2­a® 2000 ¯3°9� 2004l (4�HI$±J"���S������ $� #0 ���S��0² +�@$o ³´05p��"#$)p6µ�`¶�)$(�)����) 5�� 6�'�-¤'?��J2#$B� 70�¢%p·2?�0�!"#$ �� ³¸0 /270 /2¹5�!"�0/2ºE�0J;$1G�L"0©�!"#$ O�4 »�8J ‘tik tik’, ‘pit pit’, ‘pink’,

‘tsic’J;$1G�L"#$�� g¼Q½�´ 1996, Baker 1997, del Hoyo et al. 2006 9:9�2009l (4�HI$±J"�������� &;��h< A¦�0�=|B�"�0$0> ±J"#$ $� # ���S��0#0�!" o? ³´05p��"#$ B� ¢%p·2?�0�!"

@ % A v168

Page 6: Phylloscopus plumbeitarsus - JST

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P. nitidus� P. trochiloides� ^_`KH�L � ab�cY�RS c��defgR�hi���� T./jk�l�mn�o�./pqrP��s�mn�t��u�vwxKH�L,uyrPz�* 55�65% 0^_`KH�L� P. trochiloides�{| P. t. viridanusA ��� 55

}70% (P. nitidus)������ P10�s�~�)\t���� P6./ P8R�� 3�,T�����\ P5�T��� �,�/��.��� ��������� ��vwxKH�L)\�*� +���������b.����z��Q�b������ ���vwxKH�L)\���*�*jk���� G����L,�S���(�i�����./� B��CD���i� (Svensson 1992, Baker 1997, Beaman & Madge 1998, van der Vliet et al. 2001, del Hoyo

et al. 2006)F�����./� B��CD � P. nitidus, P. trochiloides� ^_`KH�L����.�����k/��F

�������P. nitidus, P. trochiloides� ^_`KH�L��� �+���\��rSFP. nitidus D�������������\� ab�  V� D+��c���¡���� P9

�WX ¢£ P4WX� P5WX�¤¥,����./ (Svensson 1992, Baker 1997, van der

Vliet et al. 2001, del Hoyo et al. 2006)� B��CD� E��FP. trochiloides, 4{|�¦§/�P��S§� {|¨�,��rSF {| P. t. viridanus

� ab�¢£�����©rP�\� ª« ¢£9~�� 1B¬­�¤~�,��®¯�°[±S�� 9~�WX�Y�N ²�,hr��.��� ²�� ³�.��\� 9~�WXY�N�t�����´�)\°¢£µ��� (Ticehurst 1938, Svensson 1992, Baker 1997,

van der Vliet et al. 2001)� B��CD� E��F {| P. t. ludlowi ª«� 1B��*� P9�P3)\µ��� (Ticehurst 1938, Williamson 1967, Ali & Ripley 1997, Grimmett et al. 1998)� B��CD� E��F {| P. t. trochiloides 1B�ª« uy#��¬� 2B[ ³�.��\� P9�¢£ P3)\µ��� (Ticehurst 1938, Svensson 1992, Ali & Ripley 1997)� B��CD� E��F {| P. t. obscuratus P9� P2�¶t. P3�¶t��� P2� P3�¤¥�t������� B��CD���{|���·¸¹ ���� D+��cY������ (Tic-

ehurst 1938, Williamson 1967)� B��CD� ººE��F^_`KH�L D+��Y������� ab��(»kP�©i�CD 9¼������� ¤~��9~��WX�cY�.Y����S§ 2B�ª«������ 9~�WX�Y�N ²�,hi���� 9~�WXY�N�t�����´�)\°¢£9����� P9

^_`KH�L Phylloscopus plumbeitarsus�½B,�­��� 169

Page 7: Phylloscopus plumbeitarsus - JST

� P2���� P5������� P2� P4�� ������ (Ticehurst 1938, Svensson

1992, Baker 1997, van der Vliet et al. 2001)� ��������� ���� ����� !"��#��$%& ��'����()*+���� ,���� �-��� ./%)��� 0�#���&1&2�� (Dickinson 2003, del Hoyo et al.

2006), P. nitidus � P. trochiloides 345��#�������6�� ��7�8���9:� ;��<=� ��#��>���������� ! P. plumbeitarsus�?@�A��

��������5� P. trochiloides��������BCDE (2000)��AF���,�2�G���:�� � ��H��G�CDE (2005)� P. trochiloides �!IJ 2"G�� ����� !�!IJ 6"G��K#LA��;=� MANAG��O:=P$;=%Q;=: � &R��A��MSG�TUV� P$;� Table 1 ���W�� �X� ��H��G�CDE(2005)�� Table 1 �Y;�G�Z�� [\]'�^(_��X` 2001) 5a 5� 1

������� !G����� bc (2002)�de�*+fg�YLA=: � ;�;�&R�hi$,���� ��������� !2���� jk�� !2� �?@��W� Table 1��YL��<��bc (2002)������ !��;�jk�� !���;�:���;�lm-�./no�� pqLA=: de���01K2�<r�;�:J� &Rs2jk�� !de �lm-t3�*4�u�v@ de�wx� � Gy z{|%}~ �:u��jk�� !��������� !��� �?@�A �� ��2r �g���� ��8�5;:� bc (2002)�� �6�7�8��8����:�n2�8��) n2� ��T����� !�n��;=: �;=: �� �9!I���A=:�:� 6:Gy�� !I�;=� van der Vliet et al. (2001)���5�6� P. trochiloidesZ��n2���� �����n�v@� Leader (1993) T�9;����������/:n2� �;=: � ;�;���� Leader (1993)��6���1 :n2� ��TGy;OO� ��� ����n���<�:n�2��� ����� !���n2� �;=: � bc (2002)�pqLA=: de2��6n����/n2��� ����� !"�:���2� 7�8T;�;rA�:�� jk�� !���6:� bc (2002)���;=:�:�� de��<=�>n2������ n2�������� !��>jk�� !����;=: � ��� bc (2002)�� ?@��A�¡)¢£¤¢��:�;=: �� de���¥�����>¡)¢£¤¢�� �u�v@� jk�� !��;=: � L��� ���v@ ¦�B§�M���C¨;=: ©�B§�LD� P9�L� P10ª�©�m-��C¨E�� bc (2002)�#W=: �u�� ����� !��>«;�jk�� !�6:� ©�B§� 7Fv@ ���¬¥�=?@ �� ��������� !��>jk�� !�?@ Zu�­®2� ������ !G�� 13"�<�� Mu¯� No. 3u¯ 2000) 5a 26�G��

No. 6, No. 7 3"�*+fg�012�<�� �°;� No. 3 5a 26�G������±����2�²�����³LA=: �W� �A�*���°<�7�8�� � ;��<=� *+fg�G1;=: ����� !G�� 11" 15��hJ ´Hµ;���G�LA�¶¬�� 1�I��ª>·:��¸T¹J;=¬º;�» °<�� �A�u¯� ¼

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