pons, j.m. & vicens, e. field guide to pyrenean rudist bivalves. · 2008-04-23 · pons &...

Org. Divers. Evol. 6, Electr. Suppl. 16, part 3: 1 - 18 (2006) © Gesellschaft für Biologische Systematik http://senckenberg.de/odes/06-16.htm URN: urn:nbn:de:0028-odes0616-5

Org. Divers. Evol. 6, Electr. Suppl. 16, part 3 (2006)

Pons, J.M. & Vicens, E. Field Guide to Pyrenean Rudist Bivalves. Electr. Suppl. 16, part 3. − to: Malchus, N. & Pons, J.M. (2006): Abstracts and Posters of the “International Congress on Bivalvia” at the Universitat Autònoma de Barcelona, Spain, 22-27 July 2006. Org. Divers. Evol. 6(4): 327.

Pons & Vicens: Field Guide to Pyrenean Rudist Bivalves 2




Logo and cover design: Vicent Vicedo

Organising CommitteeDr. Cristian Altaba, Universitat de les Illes Balears

Dr. Jaume Gallemí, Museu de Geologia (MCNC), BarcelonaDr. Eulalia Gili, Universitat Autònoma de Barcelona

Dr. Gregori López, Museu de Geologia (MCNC), BarcelonaDr. Nikolaus Malchus, Universitat Autònoma de Barcelona, (president)

Dr. Ricard Martínez, Universitat Autònoma de BarcelonaDr. Jose Maria Pons, Universitat Autònoma de Barcelona, (vice-president)Dr. Carlos Saavedra, Instituto de Acuicultura-Torre La Sal, Castellón (CSIC)Dr. José Templado, Museo Nacional de Ciencias Naturales, Madrid (CSIC)

Dr. Francesc Uribe, Museu de Zoologia (MCNC), BarcelonaDr. Enric Vicens, Universitat Autònoma de Barcelona

Technical AssistenceJavier Aguilar, Sergi Albrich, Sandra Córcoles, Mercedes González, Ramón Mercedes,

Angéelica Oviedo, Yolanda Pichardo, Mònica Soler, Luis Troya, Vicent Vicedo,Raquel Vilallonga, Sheila Villalba

Scientific CommitteeDr. Cristian Altaba, Universitat de les Illes Balears

Dr. Rudiger Bieler, The Field Museum, Chicago, USADr. David Campbell, University of Alabama, USA

Dr. Joseph Carter, University of North Carolina, USADr. Antonio Checa, Universidad de Granada

Dr. Graciela Delvene, Museo Geominero, MadridDr. Gonzalo Giribet, Harvard University, USA

Dr. Serge Gofas, Universidad de MálagaDr. Elizabeth Harper, Cambridge University, UK

Dr. Gerhard Haszprunar, Zoologische Staatssammlung München, GermanyDr. Katrin Linse, British Antarctic Survey, Cambridge, UK

Dr. Paula Mikkelsen, American Museum of Natural History, NY, USADr. Diarmaid Ó'Foighil, Museum of Zoology, University of Michigan, USA

Dr. Juan Peña, Instituto de Acuicultura Torre la Sal, Castellón (CSIC)Dr. Carmen Salas, Universidad de Málaga

Dr. Eleftherios Zouros, University of Crete, Greece

Sponsors and Collaborators



OBJECTIVES

At Collades de Basturs, to observe rudists and rudist buildups in the neritic marlsand in the intercalated intervals each deposited from a carbonate shelf of the SantonianCollades Member. The succession consists of four small-scale sequences within the highstandsystems tract of a depositional sequence, topped by a small scale drowning sequence.

At Cellers-Moror area, to observe the last rudists and rudist buildups of the Pyreneanmarine Upper Cretaceous intercalated in the continental deposits of the Tremp Formation(Garumnian facies).

MOST RECENT REFERENCES

Sanders, D. and Pons, J.M. 2001. Stratigraphic architecture of a Santonian mixed siliciclastic-carbonate succession (Catalonian Pyrenees, Spain). Facies 44: 105-136.

Vicens, E., Ardèvol, L., López Martínez, N. and Arribas, M.E. 2004. Rudist biostratigraphyin the Campanian-Maastrichtian of the south-central Pyrenees, Spain. CourierForschungsinstitut Senckenberg 247: 113-127.

FIELD TRIP SCHEDULEThursday 27 July

7.00 Field trip starts from Hotel Serhs Campus, Bellaterra

7.00-9.15 Van ride Bellaterra – IsonaBellaterra - (A-18) - Manresa - (C-25) - Calaf - (C-1412 / C-1412a) - Ponts - (C-1412b) - Isona

9.15-9.45 Coffee pause in IsonaTo prepare us for the long first stop

9.45-10.00 Ride to STOP 1Isona – Abella de la Conca – Can Borrell

10.00-13.30 STOP 1. Collades de BastursSorry, this is really not a stop. We will wander on rudists up and dawn alongfour km. Solar protection for skin and eyes, a hat or a cap, and suitable footwearis highly recommended.

13.30-13.45 Ride back to Isona

13.45-14.45 Lunch

14.45-15.30 Ride to Cellers and MororIsona - (C-1412) - Tremp - (C-13) - Cellers - (LV-9124) - Moror

15:30-17:30 STOP 2. Moror /18:00 Gentle walk/ride visiting several outcrops of the Cellers-Moror area

18:00-20:30 Ride back to BellaterraMoror - (LV-9124) - Cellers (C-13 / C-147) - Balaguer - (LV-3025) - Agramunt- (L-303) - Cervera - (C-25) - Manresa - (A-18) - Bellaterra



THE GEOLOGICAL SCENERY OF OUR JOURNEY

Our field trip starts in Bellaterra, near Sabadell, and follows NNW the motorway untilManresa. From the continental Miocene deposits of the Vallès Depression we will cross theVallès fault, the metamorphic Palaeozoic, the folded marine Triassic, the overthrust of thelast two on the continental Palaeocene, this last, and the Lower Eocene in the Ebre Basin.

From Manresa the road runs NW gently on the Upper Eocene and Oligocene continentaldeposits of the Ebre Basin and then the road climbs to the Comiols Pass (1.101 m) arrivingto the Cretaceous. Unfortunately it is not the best period of the year to have a clear view ofthe Conca de Tremp with the Axial Area of the Pyrenees on the background. From here,the road goes dawn in the Tremp Basin and reaches Isona crossing from the last marineCretaceous rocks to the continental Garumnian facies.

The mountain road to Abella de la Conca first, and then to Can Borrell, shall drive uson the Cretaceous to the Santonian rudist-paradise of Collades de Basturs. STOP 1

After the lunch, we will cross towards S the Garumnian facies and the Palaeoceneof the Tremp Basin syncline until the uppermost marine Cretaceous beds of the Montsecto visit the last Pyrenean rudists near Cellers and Moror. STOP 2

Return shall be southwards, through the Terradets Gorge, crossing the MontsecRange, and then across the Marginal Ranges (Serres Marginals) until Balaguer. Theneastwards to Cervera and Manresa and from there back to Bellaterra.

Localities and units appearing in the figures 1 and 2 have been emphasized.

Servei Geològic de Catalunya, Institut Cartogràfic de Catalunya. 2002. Mapa Geològic deCatalunya 1:250.000.

Berastegui, X., Losantos, M., Muñoz, J. A., Puigdefàbregas, C. 1993. Tall geològic del Pirineucentral 1:200.000. Servei Geològic de Catalunya, Institut Cartogràfic de Catalunya.



Fig. 1 - Simplified geological map of Catalonia

Fig. 2 - Southern part of the ECORS profile

EC

OR

S



Fig. 3 - Aerial view of our field trip area

The photo was taken during a flight Barcelona-London. The Montsec Range appearsat the upper left corner. The Noguera Pallaresa river crosses the upper part of the photofrom the right margin (N) to the left (S). Sant Antoni Lake is distinguishable at right andCellers Lake and the Terradets Gorge at left. The centre of the photo is occupied by theTremp Basin syncline. The Sant Corneli-Abella-Boixols anticline appears at the lower rightmargin and the Isona anticline at the left bottom.

The Cellers-Moror area (STOP 2) at the left and the Collades de Basturs area(STOP 1) at the right are emphasized.



Fig. 5 - Ortho-photomap of the Cellers-Moror area

Fig. 4 - Ortho-photomap of the Collades de Basturs area



Fig. 6 - STOP 1 Collades de Basturs A - Panoramic view to west.B - Panoramic view to east.

A

B



Fig. 7 - STOP 1Collades deBasturs

A - Hippuritidbiostrome within acarbonate interval.Base is at right.View to west.

B - Detail of thetabulae developedin the inner shelllayer of thehippuritids. Baseis at right. View towest.

C - Detail of thehippuritidbiostrome toppedby the start of acoral-sponge-rudist biocon-struction. Base isat right. View towest.

A

B

C



Fig. 8 - STOP 1 Collades de BastursA - Depositional geometry in the upward-shoaling part of a carbonate interval.

View to east.B - Bioclastic rudstones to grainstons interval toplapping the bioconstruction.

View to east.C - Coral-sponge-rudist bioconstruction. View to east.

A

B C



Fig. 9 - STOP 2 Cellers-Moror. Correlated sections alongthe Cellers-Moror area.We hope you learned enough Catalan to understandthe legend, designs may help.

Fig. 10 - Geological map of the Cellers-Moror area.



Fig. 11 -STOP 2Cellers-Moror

A - Biradiolites chaperi horizon

B - Detail of A

D - Detail of C

C - Biradiolites moroi horizon

E -Hippuritellacastroi s.s.horizon withPraeradiolitesboucheroni

F - Hippuritella castroi s.s. horizonwith Monopleura moroi

G - Detail of F



Fig. 12 - Shell morphology of the hippuritids (Hippurites radiosus)A - Transverse section of the right valve close to the commissure.B - View of the left valve.C - Detail of the left valve.D - Radial section of both valves.E - Tangential section of the left valve.F- Radial section of both valves.



Fig. 13 - Shell morphology of the radiolitidsShell growth in normal bivalves and in radiolitid rudist bivalves. A, hemiperiferal growth(upper) and holoperiferal growth (lower) in bivalve shells. B, scheme of the radial section ofboth valves in a radiolitid, vertical and radial components of shell growth have been indicatedwith arrows in both valves. C, adaptation of Taylor et al. (1969, modified from Beedham1958) figure of Anodonta cygnea to explain shell layers and shell growth in bivalves and tocompare with radiolitids, the outer shell layer part of a growth lamella has been shaded inboth valves, most details of the soft parts are irrelevant to our subject but have been retainedas in the original figure. D, main sections used to study the outer shell layer structure inradiolitids, three growth lamellae have been represented on some of the sections, flat andhorizontal on the radial section at left, concentrically folded (dipping of the lamellae changesfrom the inner to the outer margin) on the radial section at right, and radially folded on thetangential section, a transverse section should cut the growth lamellae with different angleat different parts of the section in the last two cases. E, acute and flat costae and sinusesat the outer surface of both valves and zigzag commissure, produced by up- and down- radialfolds of the growth lamellae, in Biradiolites lameracensis. F, concentric folds of the growthlamellae and void space between growth rings (filled with dark sediment) on the radial sectionof both valves in Praeradiolites subtoucasi. Scale bars in E and F represent 10 mm.



Fig. 14 - Shell morphology of the radiolitidsRadial structures (bands, sinuses or folds) in radiolitids. A-H, transverse sections of rightvalves. A, Biradiolites chaperi. B, Radiolites aurigerensis. C, Bournonia excavata. D, Sauvagesiasharpei. E, Biradiolites lameracensis. F, Durania arnaudi. G, Praeradiolites subtoucasi. H,Darendeella anatolica. Arrows indicate, at the inner shell margin, the position of the radialstructures. Scale bars represent 10 mm (do note F has its own bar).



fig. 15 - Shell morphology of the radiolitidsNon compact structure with continuous radial ridges on the growth lamellae. Biradiolites

fig. 16 - Shell morphology of theradiolitidsSimplified model of the noncompact structure withcontinuous radial ridgesCenter right.

R A D I A L S E C T I O N

TANGENTIAL SECTION

moroi. A, transverse section of the right valve, cutting obliquethe growth lamellae. B, section nearly parallel to the gently folded

growth lamellae. C, section nearly parallelto the folded growth lamellae. D, tangential

section. E-F, radial sections.Scale bar represents 10 mm.



fig. 19 - Shell morphology of theradiolitidsGrowth micro-rhythms in thegrowth lamellae. Radial sectionof right valve, Rosellia sp. Scalebar represents 0.5 mm.

fig. 17 - Shell morphology of theradiolitidsNon compact normal cellularstructure on the growth lamellae.Radiolites angeiodes. Radialstructures are a widening of theinner-marginal groove incorrespondence with upwardsradial folding of the growthlamellae (sinus). A-B, radialsections of both valves; theventral radial structure is cut inB. C, transverse section of rightvalve. Scale bar represents 10mm.

fig. 18 - Shell morphology ofthe radiolitidsSimplified model of thenon compact normalcellular structure.

R A D I A L S E C T I O N

TANGENTIAL SECTION

pons, j.m. & vicens, e. field guide to pyrenean rudist bivalves. · 2008-04-23 · pons &...

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