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reptile family tree - peters 2018 1321 taxa, 231 characters · buriolestes ulbra-pvt280...

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  • Reptile Family Tree - Peters 2018 1321 taxa, 231 characters · Buriolestes ULBRA-PVT280 Pteroglossus Buceros Septencoarcias Pelecanus Catharacta Balaeniceps Fregata Pelagornis Puffinus
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Silvanerpeton Gephyrostegeus bohemicus Tijubina Bavarisaurus (= Homoeosaurus) macrodactylus Huehuecuetzpalli Macrocnemus BES SC 111 Litorosuchus LPV 30280 w/embryo Macrocnemus T2472 Dinocephalosaurus Langobardisaurus Tanytrachelos Triopticus Tanystropheus k and q Cosesaurus Kyrgyzsaurus Sharovipteryx Longisquama Schoenesmahl (formerly Bavarisaurus) Fraxinisaura Lacertulus Daohugou lizard IVPP V13747 Meyasaurus Carusia Hoyalacerta Homoeosaurus solnhofensis IPB R 535 Homoeosaurus tail IVPP V14386 Dalinghosaurus Eldeceeon 1990.7.1 Eldeceeon holotype Tulerpeton Westlothiana Casineria Brouffia Paleothyris Coelostegus Hylonomus Protorothyris MCZ1532 Eudibamus Aphelosaurus Petrolacosaurus Araeoscelis Spinoaequalis Galesphyrus Youngina BPI 3859 Acerosodontosaurus Thadeosaurus Tangasaurus Hovasaurus Claudiosaurus Adelosaurus Atopodentatus Largocephalosaurus Omphalosaurus Sinosaurosphargis Anarosaurus Youngina SAM K7710 Youngina TM 3603 Youngopsis rubidgei RC90 Youngopsis kitchingi TM1490 Youngoides RC91 Youngina AMNH 5561 Youngoides UC1528 Proterosuchus BPI 1 4016 Chasmatosaurus NMQR 1484 Teyujagua Proterosuchus RC96 Euparkeria SAM PK K5867 Euparkeria SAM PK K6047A Osmolskina Tasmaniosaurus Revueltosaurus Fugusuchus Garjainia Erythrosuchus Luperosuchus Shansisuchus Riojasuchus Ornithosuchus Vjushkovia Prestosuchus Batrachotomus Saurosuchus Smok Teratosaurus Sikannisuchus Postosuchus Decuriasuchus Pagosvenator Turfanosuchus Poposaurus Sacisaurus Silesaurus Lotosaurus Shuvosaurus Effigia Protorothyris CM 8617 Protorothyris MCZ 2149 Vaughnictis MFSN 19235 Scandensia Archaeovenator Anningia Orovenator Pyozia Broomia Milleropsis BPI-720 Erpetonyx Ascendonanus MNC-TA0924 Rauisuchus PVL 4597 Lewisuchus Tarjadia Parringtonia Gracilisuchus MCZ4116 Saltopus Scleromochlus Teleocrater Arizonasaurus Yarasuchus Qianosuchus Nundasuchus Ticinosuchus Aeotosauroides Aetosaurus Stagonolepis Wumengosaurus Thaisaurus Xinminosaurus Utatsusaurus Shastasaurus alexandrae Shastasaurus pacificus UCMP 9017 Grippia Macgowania Mikadocephalus YGMIRSPCV03108 Parvinatator Chaohusaurus Besanosaurus Qianichthyosaurus CMNH V1412 Qianichthyosaurus IVPP V11839 Phalarodon fraasi Contectopalatus 'Phalarodon' nordenskioeldii Mixosaurus Cymbospondylus petrinus Shonisaurus sikanniensis Guanlingsaurus GNG dq-50 Guanlingsaurus YGMR SPC V03107 Wimanius Platypterygius Guizhouichthyosaurus tangae IVPP V 11853 'Cymbospondylus' buchseri Shonisaurus popularis Ichthyosaurus communis Ophthalmosaurus Leptonectes tenuirostris Hauffiopteryx BRLSI M1399 Suevoleviathan GPIT 328/4/5 Suevoleviathan SMNS15390 Eurhinosaurus Chasmatosaurus QR 880C.500 Chasmatosaurus SAM PK 11208 Chasmatosaurus TM 201 Elaphrosuchus RC59 Diandongosuchus Teraterpeton Mesorhinosuchus Paleorhinus Parasuchus Chanaresuchus Proterochampsa PVSJ 77 Proterochampsa PVSJ606 Pseudochampsa PVSJ 567 Gualosuchus Cerritosaurus Tropidosuchus PVL 4601 Tropidosuchus PVL 4602 Lagerpeton Bergamodactylus MPUM 6009 Eudimorphodon Preondactyluis Dimorphodon Nyctiphruretus holotype Nycteroleter Nyctiphruretus PIN 4659/1 Barasaurus Kitchingnathus Santaisaurus Sauropareion Coletta Paliguana Sophineta Psilotrachelosaurus PIMUZ AIII 0192 SMF R 4710 Stereosternum Brazilosaurus holotype Mesosaurus Serpianosaurus Concavispina Xinpusaurus kohi Xinpusaurus suni Thalattosaurus Wachtlerosaurus Endennasaurus Clarazia Askeptosaurus Anshunsaurus Miodentosaurus Eusaurosphargis holotype Helveticosaurus Vancleavea Diandongosaurus Palatodonta Pappochelys Palacrodon Majiashanosaurus Paraplacodus Placodus gigas Henodus Placochelys Cyamodus Pachypleurosaurus Cartorhynchus Sclerocormus Qianxisaurus Keichousaurus Dianmeisaurus Dawazisaurus Hanosaurus Lariosaurus Corosaurus Nothosaurus Wangosaurus Simosaurus Anningsaura Pistosaurus Yunguisaurus Albertonectes Plesiosaurus Trinacromerum Brachauchenius Kronosaurus Proterosuchus SAMPKK10603 Doswellia Youngina BPI 2871 Elachistosuchus Tchoiria Ikechosaurus Champsosaurus Simoedosaurus Cteniogenys Philydrosaurus Lazarussuchus Hyphalosaurus Monjurosuchus GMV216 Monjurosuchus NMM3671 SMNS12591skull Terrestrisuchus Litargosuchus Erpetosuchus Hesperosuchus SMNS12352 partial skull Pedeticosaurus Yonghesuchus Dromicosuchus Sichuanosuchus Protosuchus Sphenosuchus Saltoposuchus Dibothrosuchus Dakosaurus Teleosaurus Pelagosaurus Dyoplax Metriorhynchus Sebecus Isisfordia Crocodylus niloticus Araripesuchus Simosuchus Mariliasuchus Armadillosuchus Caipirasuchus Baurusuchus Herrerasaurus Tchingisaurus Chometokadmon Gekko smithii Lialis Gekko gecko Eublepharis Norellius Ardeosaurus Eichstaettisaurus schroederi Eichstaettisaurus gouldi Jucaraseps Aphanizocnemus Adriosaurus Pontosaurus Primitivus Tetrapodophis Dinilysia Pachyrhachis Boa Crotalus Loxocemus Yabeinosaurus IVPP V13284 Xenopeltis Anilius Cylindrophis Uropeltis Anomochilus Leptotyphlops Bahndwivici FMNH PR2260 Aigialosaurus Tethysaurus Tylosaurus Plotosaurus Not Ascendonanus MNC TA1045 Saniwa ensidens FMNH Estesia Varanus komodensis Varanus griseus Shinisaurus Heloderma Lanthanotus Anniella Gobiderma Ophisaurus Kuroyuriella Myrmecodaptria Elgaria Cryptolacerta Chalcides ocellatus Chalcides guentheri Gymnophthalmus Vanzosaura Sirenoscincus Slavoia MGR-I-112 Sineoamphisbaena Crythiosaurus Spathorhynchus Dibamus Tamaulipasaurus Bipes Amphisbaena Barberenasuchus Eodromaeus Sakurasaurus Limnoscelis Orobates Tseajaia Tetraceratops Milleretta RC14 Milleretta RC70 Bashkyroleter Macroleter Emeroleter Lanthanosuchus Urumqia Bruktererpeton Thuringothyris MNG 7729 Thuringothyris MNG 10183 Reiszorhinus Cephalerpeton Opisthodontosaurus Concordia KUVP 8702a Concordia KUVP 96/95 Romeria primus Saurorictus Romeria texana Protocaptorhinus Paracaptorhinus Eocaptorhinus Captorhinus Labidosaurus Calanguban Acanthodactylus Eolacerta Lacerta Tupinambis Dorsetisaurus Slavoia holotype ZPAL MgR-I/8 Slavoia ZPAL MgR-III/77 Macrocephalosaurus Tianyusaurus Diadectes CM25471 Silvadectes absitus MNG 8853 Phonodus Bolosaurus Belebey Oradectes sanmiguelensis Diadectes UC706 Diadectes sideropelicus Procolophon Kapes Scolaparia NSM 99 GF82.1 Leptopleuron Pentaedrusaurus Hypsognathus Youngina BPI 375 Prolacerta AMNH 9502 Ixalerpeton Czatkowiella Malerisaurus Prolacerta BPI 1/471 Ozimek Protorosaurus NHMW 1943I4* Boreopricea Jaxtasuchus Protorosaurus NMK S 180 Protorosaurus WMsN P47361 Pamelaria Zupaysaurus Megapnosaurus Stephanospondylus Provelosaurus Deltavjatia KPM232 Scutosaurus Bradysaurus NHMW Anthodon Pareiasuchus Pumiliopareia JZC Bu1803 - amber Liushusaurus Purbicella Euposaurus Iguana Trioceros Phrynosoma Basiliscus Anolis Pristidactylus Chlamydosaurus JKZ Bu267 -amber Tawa Staurikosaurus Segisaurus Zuolong Marasuchus Procompsognathus Dracoraptor Coelophysis Dilophosaurus Sinocalliopteryx Buriolestes CAPPA/UFSM 0035 Velociraptor Balaur Haplocheirus Shuvuuia Halszkaraptor Sinornithoides Sauronithoides Jinfengopteryx Anchiornis LPM B00169 Anchiornis PKUP V1068 Anchiornis IVPP V14378 Aurornis Caihong Buitreraptor Sinovenator Eosinopteryx Xiaotingia Elliotsmithia Apsisaurus Aerosaurus Varanops Varanodon Varanosaurus FMNH PR 1760 Varanosaurus BSPHM 1901 XV20 Archaeothyris Ophiacodon Haptodus Pantelosaurus Secodontosaurus Dimetrodon Sphenacodon Ianthodon Edaphosaurus Ianthasaurus Glaucosaurus Cutleria Stenocybus IVPP V18117 Kenyasaurus Galechirus Suminia Venjukovia Eodicynodon Dicynodon Heleosaurus Mycterosaurus FMNH UC169 Nikkasaurus Mesenosaurus PIN 158/1 Niaftasuchus Beipiaosaurus Rahonavis Jianchangosaurus Limusaurus Caudipteryx Khaan Juravenator Sinosauropteryx Compsognathus holotype Aorun Struthiomimus Gallimimus Ornitholestes Scriurumimus Microraptor IVPP V 13352 Microraptor M V1002 Sinornithosaurus holotype Sinornithosaurus NGMC 91 Fukuivenator Masiakasaurus Tianyuraptor Huaxiagnathus Zhenyuanlong Alioramus altai Gorgosaurus Raptorex Tyrannosaurus Reptile Family Tree - Peters 2018 1326 taxa, 231 characters Eoraptor Pampadromaeus Leyesaurus Saturnalia Pantydraco Efraasia Massospondylus adult Massospondylus embryo Yizhousaurus Chilesaurus Daemonosaurus Jeholosaurus Jesairosaurus Hypuronector Vallesaurus Avicranium Megalancosaurus Tridentinosaurus Coelurosauravus Mecistotrachelos Lanthanolania Icarosaurus Kuehneosaurus Xianglong Owenetta rubidgei Owenetta kitchingorum Candelaria Feeserpeton Australothyris Eocasea Delorhynchus Microleter Acleistorhinus Eunotosaurus Eorhynchochelys Oedaleops Eothyris Colobomycter vaughni Colobomycter pholeter Datheosaurus Ennatosaurus Casea Cotylorhynchus Placodus inexpectatus Hipposaurus Biarmosuchus Titanophoneus Procynosuchus Thrinaxodon Prozostrodon Probainognathus Chiniquodon Castorocauda Haldanodon Liaoconodon Pachygenelus Riograndia Chaliminia Oligokyphus Kayentatherium Tritylodon Spinolestes Jeholodens Gobiconodon Repenomamus robustus Repenomamus giganticus Yanoconodon Sinoconodon Brasilodon Haramiyavia Therioherpeton Megazostrodon Hadrocodium Dromiciops Asioryctes holotype Monodelphis domestica Marmosa Chironectes Docofossor Volaticotherium Alcidedorbignya Maotherium Maelestes Anagale Leptictis Rhynchocyon Harpagolestes macrocephalus Sinonyx Andrewsarchus Hemicentetes Tenrec Leptictidium Indohyus Onychodectes Ectoconus Pantolambda Barylambda Glyptodon Paramylodon Bradypus Peltephilus Fruitafossor Orycteropus Myrmecophaga Holmesina Chlamyphorus Dasypus Tamandua Cyclopes Meniscotherium Orthaspidotherium Titanoides Astrapotherium Phenacodus Thomashuxleya Gobiatherium Arsinoitherium Coryphodon Uintatherium Mesonyx Harpagolestes uintensis Ocepeia Homalodotherium Miocochilius Protypotherium Litolophus Embolotherium Eotitanops Metamynodon Aceratherium Hyracotherium Hyracodon Uintaceras Hyrachyus Mesohippus Pappoceras Equus Juxia Paraceratherium Ceratotherium Anisodon grande Pakicetus Maiacetus Mammalodon Coronodon Aetiocetus Zygorhiza Mystacodon Physeter Monodon Delphinapterus Tursiops Orcinus Hippopotamus Anthracobune Cambaytherium Cornwallius Janjucetus Paleoparadoxia Neoparadoxia Desmostylus Behemotops RBCM sp. Eschrichtius Isanacetus Balaenoptera Tokarahia Yamatocetus Cetiotherium Sitsqwayk Caperea Eubalaena Arganaceras Sphodrosaurus Sclerosaurus Odontochelys Trionyx Perochelys Ocepechelon Bunostegos Elginia mirabilis Elginia wuyongae Niolamia Meiolania Proterochersis Proganochelys Kayentachelys Baena Kallokibotion_UBB ToK-2 Arvinachelys Glyptops Mongolochelys Chubutemys Chelonia Santanachelys Dermochelys Macrochelys temminckii Terrapene carolina Foxemys Carettochelys Pelomedusa Araripemys Chelus Saichangurvel Magnuviator Pleuraspidotherium Periptychus Nambaroo ?Hapalodectes IVPP V2385 70 Anthracodromeus Reptilia Utegenia Balanerpeton Caerorhachis Amphibamus Platyrhinops Dendrerpeton Cochleosaurus Tersomius Acheloma Platyhistrix Stegops Dissorophus Broiliellus Cacops Ecolsonia Fedexia Tambachia Georgenthalia Micromelerpeton Doleserpeton Apateon Gerobatrachus Andrias Celtedens Rana Utaherpeton Tuditanus Kirktonecta Sauropleura Ptyonius Batropetes Rhynchonkos Asaphestera Micraroter Euryodus primus Pantylus Scincosaurus Batrachiderpeton Keraterpeton Diceratosaurus Diplocaulus Diploceraspis Acherontiscus Microbrachis Adelospondylus Adelogyrinus Chinlestegophis Brachydectes Lysorophus Eocaecilia Dermophis Cheirolepis Eusthenopteron Gogonasus Osteolepis Pholidogaster Colosteus Panderichthys Tiktaalik Greererpeton Koilops Spathicephalus Trimerorhachis Dvinosaurus Batrachosuchus Gerrothorax Neldasaurus Trematosaurus Metoposaurus Rhineceps Wantzosaurus Trematosaurus long rostrum Ossinodus Acanthostega MGUH-VP-8160 Deltaherpeton Ventastega Crassigyrinus Pederpes Whatcheeria Baphetes Megalocephalus Sclerocephalus smns90055 Iberospondylus Eryops Nigerpeton Saharastega Ichthyostega Proterogyrinus Eoherpeton Neopteroplax Anthracosaurus Eucritta Discosauriscus Seymouria Makowskia Seymouria.tadpole Guaibasaurus Miohippus Pectodens Ancodus Lambdotherium Sus Cainotherium Anthracotherium Archaeotherium Danjiangia Ectocion ralstonensis Notostylops Procavia Dusisiren Prorastomus Paleomastodon Gomphotherium Elephas Moeritherium Amynodon Note: This tree does not support DNA topologies over long phylogenetic distances. Certain dental traits are convergent and do not define clades. Heptodon Lophiodon Tapirus Protapirus Parotosuchus Oestocephalus Lethiscus Rileymillerus Phlegethontia CGH129 Phlegethontia AMNH6966 Acroplous Eusaurosphargis juvenile Fostercooperia Junggarsuchus Carnufex Pseudhesperosuchus Trialestes Asioryctes long rostrum Ariekanerpeton Kotlassia Pseudocrypturus Apteryx Rhynchotus Eudromia Struthio Patagopteryx Casuarius Rhea Cariama Phoenicopterus Sylviornis Sagittarius Falco Pandion Buteo Torgos Apus Parargornis Tyto Aegotheles Llallawavis Yanornis Changzuiornis Jiangchangornis Longicrusavis Jinguofortis Juehuaornis Rhamphocephalus Hongshanornis Dingavis Cryputurus Megapodius Apsaravis Ichthyornis Gansus Hesperornis Coccyzus Monias Ardea Ardeotis Eozygodactylus Butorides Anhima Pterocles Fulica Crex Aepyornis Palaelodus Passer Lonchura Tetrao Gallus Pavo Eogranivora Meleagris Crax Opisthocomus Macrocephalon Dinornis Ara macao Melopsittacus Nestor Gastornis Aramus Rallus Gavia Aechmophorus Thalasseus Charadrius Balearica Jacana Himantopus Eurypyga Grus Chroicocephalus Eocypselus Archilochus Megaceryle Jabiru Cinclus Troglodytes Melanerpes Sitta Uria Aptenodytes Geococcyx Psophia Musophaga Menura Semioptera Paradisaea Corvus Turdus Quiscalus Ciconia Hirundo Cyanocitta Rhynochetos Palaechthon Cynocephalus Chriacus Onychonycteris Rousettus Myotis Choeronycteris Zhangheotherium Metachcheiromys Manis Archaeopteryx siemensii Thermopolis Archaeopteryx albersdoerferi Daiting Archaeopteryx siemensii Berlin Liaoning embryo bird Archaeopteryx London Chongmingia Cathayornis Sulcavis Pengornis Chiappeavis Yanornis STM9-52 Iberomesornis Protopteryx Serikornis Archaeopteryx recurva (Eichstätt) Jianianhualong Liaoningvenator Sapeornis IVPP V13276 Sapeornis DNHM-F3078 Confuciusornis GMV-2132 Archaeopteryx #11 Confuciusornis MBAv1168 Changchengornis Zhongornis Archaeornithura Vegavis Ostromia - Haarlem Oedicnemus Falcarius Deltatherium Bishops Caluromys Ectocion cedrus UM 8615 Micromeryx Giraffa Camelus 98 Miocaperea Chonecetus Archaeopteryx 12 private Archaeopteryx bavarica (Munich) Jeholornis prima Jeholornis curvipes Hong Kong specimen Mei Scansoriopteryx Yi Epidexipteryx Omnivoropteryx Archaeovolans (=Yanornis) Draco volans Moloch Lyriocephalus Megachirella BRSUG 29950-12 Pleurosaurus Marmoretta Gephyrosaurus Ankylosphenodon Paleopleurosaurus Planocephalosaurus Heleosuchus Kallimodon Sphenodon Brachyrhinodon Clevosaurus Sphenotitan Leptosaurus Vadasaurus Sapheosaurus Noteosuchus Trilophosaurus Azendohsaurus Shringasaurus Eohyosaurus Mesosuchus Priosphenodon Rhynchosaurus articeps Bentonyx Hyperodapedon Colobops Saurosternon Palaegama Polychrus JZC Bu154 in amber Archaeopteryx (Wellnhoferia) grandis Buriolestes ULBRA-PVT280 Pteroglossus Buceros Septencoarcias Pelecanus Catharacta Balaeniceps Fregata Pelagornis Puffinus Macronectes Morus Fraterculus Rynchops Pinguinus Psilopogon Cyrilavis Phaethon Upupa Scopus Phalacrocorax Threskiornis Platalea Anas Anser Mergus Presbyornis Pumiliornis Columba Caloenas Pezophaps Raphus Aptornis Didunculus Coragyps Vultur Goura Lepidosaurmorpha Archosauromorpha Juramaia Chaoyangodens Brasilitherium Kuehneotherium Cifelliodon Tachyglossus Akidolestes Ornithorhynchus Ukhaatherium Morganucodon Eomaia Agilodocodon Acristatherium Cronopio Didelphis Ambolestes Arctocyon not Amphicyon Thylacinus Thylacosmilus Oxyaena Vincelestes Hyaenodon Ernanodon Didelphodon Masrasector Borhyaena Perameles Macrotis Anebodon Notoryctes Interatherium Mesotherium Procoptodon Dendrolagus Proargyrolagus Macropus Eurygenium Toxodon Pyrotherium Vombatus Phascolarctos Paedotherium Groeberia Vintana Balbaroo Phalanger Petaurus Thylacoleo Sinodelphys Theosodon Macrauchenia Chalicotherium? goldfussi Paludidraco Cymatosaurus Lactodens ?Hapalodectes IVPP V5235 Notharctus Indri Rooneyia Darwinius Tarsius Aegyptopithecus Proconsul Pan Gorilla Ardipithecus Homo sapiens Ptilocercus Tupaia Macroscelides Chrysochloris Necrolestes Apatemys Trogosus Rhyncholestes Scutisorex Limnogale Microgale cowani Potamogale Micropotamogale Zalambdalestes Solenodon Henkelotherium Gomphos Oryctolagus Brachyerix Erinaceus Echinops Thomomys Heterocephalus Ceratogaulus Coendou Maiopatagium Neoreomys Chinchilla Cavia Hydrochoerus Shenshou Vilevolodon Rattus Mus Paramys Ratufa Paulchoffatia Ignacius Plesiadapis Daubentonia Carpolestes Megaconus Rugosodon Xianshou Kryptobaatar Ptilodus Yubaatar Taeniolabis Mirischia Colius Pharomachrus Brachiosaurus Camarasaurus Diplodocus Rapetosaurus Pteropus Vulpavus Protictis Nandinia Procyon Ailurus Prohesperocyon Herpestes Talpa Miacis Hyopsodus Enaliarctos Desmatophoca Pithanotaria Zalophus Proteles Canis Amphicyon major Hesperocyon Panthera Crocuta Amphicyon galushi Mustela Ailuropoda Puijila Neotherium Ursus Gulo Arctodus Stylinodon Amphicynodon Psittacotherium Palaeosinopa Phoca Odobenus Diplovertebron Romeriscus Bystrowiella Solenodonsaurus Laosuchus Chroniosaurus PIN3585/124 Chroniosaurus Chroniosuchus Haya Pisanosaurus Scelidosaurus Emausaurus Minmi Agilisaurus Stegoceras Kulindadromaeus Tianyulong Heterodontosaurus Lesothosaurus Scutellosaurus Liaoningosaurus Stegosaurus Changchunsaurus Dryosaurus Edmontosaurus Laquintasaura Aquilops Hexinlusaurus Yinlong Psittacosaurus mongoliensis Liaoceratops Auroraceratops Leptoceratops Stenopelix Triceratops adult Triceratops juvenile Chasmosaurus adult Chasmosaurusjuvenile 72 86 97 96 88 71 100 87 99 69 94 98 82 70 95 98 62 64 64 79 74 96 62 100 100 100 <50 Compsognathus CNJ79 Proceratosaurus Scipionyx Deinocheirus Xiongguanlong Suchomimus Spinosaurus Dilong Guanlong Megaraptor Yutyrannus Sinraptor Majungasaurus Allosaurus Sinosaurus Monolophosaurus Acrocanthosaurus Eustreptospondylus Ceratosaurus Tanycolagreus 83 92 100 67 96 76 78 <50 62 92 70 97 71 80 62 67 77 Anchiornis BMNHC PH804 Barlochersaurus Llanocetus Eohupehsuchus Parahupehsuchus Hupehsuchus 88 100 74 87 82 89 95 95 87 75 79 98 87 100 97 87 100 100 92 72 75 85 72 94 83 85 100 100 100 98 100 72 84 89 100 94 98 60 100 91 85 100 73 66 94 <50 99 80 <50 95 99 60 63 82 99 77 91 78 87 84 93 94 85 94 82 100 91 94 99 97 97 99 96 100 95 84 80 93 84 60 95 97 72 92 90 99 85 98 100 89 97 98 99 93 90 97 98 72 99 91 98 57 65 54 57 80 62 81 88 95 89 89 100 100 100 98 89 55 92 81 70 90 75 77 96 95 64 51 100 <50 91 93 78 56 80 55 85 78 82 73 64 60 95 62 98 62 80 79 98 97 66 84 81 76 89 55 71 62 60 94 99 90 71 95 89 91 87 81 93 96 91 100 83 57 92 90 88 94 95 95 76 75 88 84 100 79 98 62 57 80 65 89 72 75 78 55 100 95 90 67 93 99 86 92 76 61 69 96 96 99 83 100 77 94 98 87 <50 79 83 68 100 100 100 97 86 98 77 99 97 99 98 83 83 100 92 57 64 79 90 59 81 <50 <50 85 92 81 100 100 99 96 99 86 93 98 97 88 94 97 100 99 97 94 64 100 99 87 96 92 66 73 77 99 97 81 100 91 100 97 88 98 100 100 90 90 98 96 97 99 91 82 84 84 <50 87 51 93 100 95 96 88 71 67 65 69 67 91 73 73 79 100 100 95 94 100 79 100 97 98 100 95 86 62 60 86 100 98 98 89 89 96 78 55 65 100 90 75 79 83 77 100 69 93 86 100 93 70 94 99 100 99 99 92 99 99 97 70 80 98 99 100 96 74 66 86 96 99 88 86 <50 98 95 95 75 67 72 98 71 98 92 96 58 98 52 69 98 98 98 99 94 91 85 68 80 82 71 88 <50 57 53 84 <50 96 95 82 83 99 68 87 81 66 91 100 57 65 70 83 55 84 66 80 67 99 78 69 93 67 95 75 99 83 78 85 77 95 88 100 91 96 <50 80 80 50 90 76 93 63 <50 91 82 82 61 100 91 81 84 84 67 51 86 90 84 89 82 76 80 97 97 82 100 85 99 82 87 97 62 62 52 71 82 83 94 100 81 99 94 Archaeothyris Haptodus Dimetrodon Edaphosaurus Varanosaurus Ophiacodon Cutleria Stenocybus IVPP V18117 Galechirus Galepus Galeops Patranomodon Suminia Eodicynodon Dicynodon Venjukovia Otsheria Anomocephalus Tiarajudens Hipposaurus Ictidorhinus Herpetoskylax Lemurosaurus Proburnetia Burnetia Biarmosuchus Eotitanosuchus Cyonosaurus Promoschorhynchus Scymnognathus Gorgonops Aelurognathus Lycosuchus Procynosuchus Thrinaxodon Chiniquodon Pachygenelus Megazostrodon IVPP V18119 Microurania IVPP V18120 Sinophoneus Deuterosaurus Estemmenosuchus uralensis Estemmenosuchus miribilis Phthinosuchus Archaeosyodon Australosyodon Syodon Titanophoneus Anteosaurus magnificus Anteosaurus abeli Biseridens Jonkeria Struthiocephalus Struthionops Moschops Ulemosaurus 84 88 86 68 79 81 89 70 98 85 100 89 98 94 51 93 100 68 85 100 55 60 99 93 100 98 83 92 82 97 97 81 95 90 100 99 100 66 69 68 77 87 100 76 74 79 86 89 89 92 88 81 98 95 92 57 66 87 98 68 86 63 86 86 88 99 99 98 100 75 98 98 99 97 81 98 100 96 98 94 95 93 85 96 88 100 100 100 93 66 74 94 98 70 55 70 93 93 63 99 74 85 63 98 85 72 50 90 73 85 91 97 81 88 73 91 81 74 84 100 100 100 99 83 86 96 96 100 89 9 4 <50 <50 97 100 71 87 100 100 95 97 67 72 87 91 79 95 83 78 72 85 67 98 81 90 100 58 98 73 79 75 69 87 99 76 80 71 66 80 81 71 100 99 86 67 97 95 75 95 90 97 100 55 56 94 90 88 95 <50 96 76 66 89 100 <50 98 87 98 90 100 91 67 97 59 <50 100 94 78 85 99 53 55 56 92 64 61 <50 99 <50 78 <50 81 75 90 100 100 95 90 93 85 98 96 90 51 89 95 79 95 75 73 73 94 96 94 100 72 100 92 99 <50 100 76 67 55 99 78 80 88 51 89 82 72 91 70 100 86 87 90 84 77 97 91 98 100 81 83 98 96 88 99 56 65 100 73 85 86 68 76 <50 58 83 95 77 85 55 77 95 79 87 94 96 99 98 98 97 <50 100 58 93 76 98 95 93 <50 54 50 96 95 98 91 86 86 77 78 95 99 72 80 63 58 58 58 85 90 77 71 95 69 87 96 82 98 79 89 <50 99 95 92 96 83 <50 82 82 88 98 73 99 100 96 97 <50 92 93 85 78 76 71 77 99 100 98 75 95 60 64 88 65 65 95 85 96 72 86 99 81 100 89 94 96 69 100 92 99 99 91 95 96 66 Basal Therapsida 59 taxa 75 skull only characters 82 87 95 96 97 94 100 76 <50 95 97 95 62 78 <50 74 88 99 96 100 96 96 91 85 85 100 97 94 93 89 88 93 87 92 97 96 83 82 74 99 100 83 75 100 88 100 91 81 52 98 98 100 83 66 99 100 87 71 64 79 100 55 92 69 73 92 89 60 83 73 84 80 85 95 85 98 100 52 <50 <50 98 88 91 84 92 98 100 100 89 97 100 73 51 70 <50 72 93 65 81 98 82 95 98 95 82 76 76 79 <50 77 96 81 95 69 83 87 87 93 94 94 97 80 84 94 72 89 68 91 96 71 95 94 94 <50 <50 71 86 88 77 96 99 99 80 98 70 87 70 89 100 96 87 96 63 87 83 89 73 65 83 93 66 86 57 80 98 97 78 90 68 77 78 62 88 60 57 85 92 99 56 98 92 57 90 82 Bootstrap scores are based on testing overlapping subsets due to limitations in computing power. 94 99 94 95 88 99 100 92 99 90 97 87 89 100 100 81 99 99 100 60 75 79 68 64 100 90 62 80 87 80 91 88 100 97 88 62 97 90 99 91 79 100 98 89 100 99 90 77 96 100 100 82 74 95 <50 79 57 90 79 93 95 60 96 65 74 64 90 75 67 87 95 93 98 87 73 85 90 87 78 63 71 63 95 64 97 100 61 94 61 88 74 64 82 81 100 64 57 95 98 98 67 85 67 80 93 95 91 89 <50 88 97 96 98 88 88 75 97 75 98 <50 54 56 99 93 80 96 96 78 64 89 92 93 94 <50 85 <50 72 <50 77 78 <50 96 <50 51 77 91 83 92 <50 69 85 85 68 81 100 96 62 88 67 84 81 94 100 99 94 95 <50 85 85 85 99 90 <50 53 72 96 88 90 98 97 <50 93 85 52 78 78 100 93 78 <50 67 100 97 98 67 100 97 100 97 93 <50 99 <50 67 70 77 98 95 <50 76 Priacodon Trioracodon Tinodon Shuotherium Ausktribosphenos Amphitherium Docodon 81 99 81 91 57 88 93 97 100 78 57 100 76 96 100 67 100 96 83 77 88 <50 55 81 99 85 96 95 96 <50 95 57 88 81 <50 61 99 62 88 100 100 100 82 77 78 85 94 97 99 58 59 76 Milosaurus 83 100 70 65 80 57 81

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Page 1: Reptile Family Tree - Peters 2018 1321 taxa, 231 characters · Buriolestes ULBRA-PVT280 Pteroglossus Buceros Septencoarcias Pelecanus Catharacta Balaeniceps Fregata Pelagornis Puffinus

SilvanerpetonGephyrostegeus bohemicus

TijubinaBavarisaurus (= Homoeosaurus) macrodactylusHuehuecuetzpalli

Macrocnemus BES SC 111

LitorosuchusLPV 30280 w/embryo

Macrocnemus T2472Dinocephalosaurus

Langobardisaurus

TanytrachelosTriopticus

Tanystropheus k and q

CosesaurusKyrgyzsaurusSharovipteryxLongisquama

Schoenesmahl (formerly Bavarisaurus)FraxinisauraLacertulus

Daohugou lizard IVPP V13747

Meyasaurus

Carusia

Hoyalacerta

Homoeosaurus solnhofensisIPB R 535 Homoeosaurus tail

IVPP V14386Dalinghosaurus

Eldeceeon 1990.7.1Eldeceeon holotype

Tulerpeton

WestlothianaCasineriaBrouffia

PaleothyrisCoelostegus

Hylonomus

Protorothyris MCZ1532

EudibamusAphelosaurusPetrolacosaurusAraeoscelisSpinoaequalisGalesphyrusYoungina BPI 3859AcerosodontosaurusThadeosaurusTangasaurusHovasaurusClaudiosaurusAdelosaurusAtopodentatus

LargocephalosaurusOmphalosaurus

Sinosaurosphargis

Anarosaurus

Youngina SAM K7710

Youngina TM 3603Youngopsis rubidgei RC90Youngopsis kitchingi TM1490Youngoides RC91Youngina AMNH 5561Youngoides UC1528

Proterosuchus BPI 1 4016

Chasmatosaurus NMQR 1484Teyujagua

Proterosuchus RC96

Euparkeria SAM PK K5867Euparkeria SAM PK K6047A

OsmolskinaTasmaniosaurusRevueltosaurusFugusuchusGarjainiaErythrosuchusLuperosuchusShansisuchus

RiojasuchusOrnithosuchus

VjushkoviaPrestosuchusBatrachotomusSaurosuchus

SmokTeratosaurusSikannisuchusPostosuchus

DecuriasuchusPagosvenatorTurfanosuchusPoposaurusSacisaurusSilesaurusLotosaurusShuvosaurusEffigia

Protorothyris CM 8617Protorothyris MCZ 2149Vaughnictis

MFSN 19235Scandensia

ArchaeovenatorAnningia

OrovenatorPyoziaBroomiaMilleropsis BPI-720Erpetonyx

Ascendonanus MNC-TA0924

Rauisuchus

PVL 4597

Lewisuchus

TarjadiaParringtonia

GracilisuchusMCZ4116SaltopusScleromochlus

TeleocraterArizonasaurus

YarasuchusQianosuchusNundasuchusTicinosuchusAeotosauroidesAetosaurusStagonolepis

Wumengosaurus

ThaisaurusXinminosaurusUtatsusaurusShastasaurus alexandraeShastasaurus pacificus UCMP 9017GrippiaMacgowania

MikadocephalusYGMIRSPCV03108ParvinatatorChaohusaurusBesanosaurusQianichthyosaurus CMNH V1412Qianichthyosaurus IVPP V11839Phalarodon fraasiContectopalatus'Phalarodon' nordenskioeldiiMixosaurusCymbospondylus petrinusShonisaurus sikanniensisGuanlingsaurus GNG dq-50Guanlingsaurus YGMR SPC V03107

WimaniusPlatypterygiusGuizhouichthyosaurus tangae IVPP V 11853'Cymbospondylus' buchseriShonisaurus popularisIchthyosaurus communisOphthalmosaurusLeptonectes tenuirostrisHauffiopteryxBRLSI M1399

Suevoleviathan GPIT 328/4/5Suevoleviathan SMNS15390

Eurhinosaurus

Chasmatosaurus QR 880C.500Chasmatosaurus SAM PK 11208Chasmatosaurus TM 201

Elaphrosuchus RC59

DiandongosuchusTeraterpetonMesorhinosuchusPaleorhinusParasuchusChanaresuchusProterochampsa PVSJ 77Proterochampsa PVSJ606Pseudochampsa PVSJ 567GualosuchusCerritosaurusTropidosuchus PVL 4601Tropidosuchus PVL 4602Lagerpeton

Bergamodactylus MPUM 6009EudimorphodonPreondactyluisDimorphodon

Nyctiphruretus holotypeNycteroleter

Nyctiphruretus PIN 4659/1

BarasaurusKitchingnathusSantaisaurusSauropareionColettaPaliguanaSophineta

PsilotrachelosaurusPIMUZ AIII 0192SMF R 4710StereosternumBrazilosaurus holotypeMesosaurusSerpianosaurusConcavispinaXinpusaurus kohiXinpusaurus suniThalattosaurusWachtlerosaurusEndennasaurusClarazia

AskeptosaurusAnshunsaurusMiodentosaurus

Eusaurosphargis holotypeHelveticosaurus

Vancleavea

DiandongosaurusPalatodontaPappochelysPalacrodonMajiashanosaurusParaplacodus

Placodus gigasHenodusPlacochelysCyamodusPachypleurosaurus

CartorhynchusSclerocormus

Qianxisaurus

KeichousaurusDianmeisaurusDawazisaurusHanosaurusLariosaurusCorosaurusNothosaurusWangosaurus

SimosaurusAnningsauraPistosaurusYunguisaurus

AlbertonectesPlesiosaurus

TrinacromerumBrachaucheniusKronosaurus

Proterosuchus SAMPKK10603DoswelliaYoungina BPI 2871ElachistosuchusTchoiriaIkechosaurusChampsosaurusSimoedosaurusCteniogenysPhilydrosaurusLazarussuchusHyphalosaurusMonjurosuchus GMV216Monjurosuchus NMM3671

SMNS12591skullTerrestrisuchus

LitargosuchusErpetosuchus

HesperosuchusSMNS12352 partial skull

PedeticosaurusYonghesuchusDromicosuchusSichuanosuchusProtosuchusSphenosuchusSaltoposuchusDibothrosuchus

DakosaurusTeleosaurusPelagosaurusDyoplax

MetriorhynchusSebecusIsisfordiaCrocodylus niloticusAraripesuchusSimosuchus

MariliasuchusArmadillosuchus

CaipirasuchusBaurusuchus

Herrerasaurus

TchingisaurusChometokadmonGekko smithiiLialisGekko geckoEublepharisNorelliusArdeosaurusEichstaettisaurus schroederiEichstaettisaurus gouldiJucaraseps

AphanizocnemusAdriosaurusPontosaurusPrimitivus

TetrapodophisDinilysiaPachyrhachisBoaCrotalusLoxocemus

Yabeinosaurus IVPP V13284

XenopeltisAniliusCylindrophisUropeltisAnomochilusLeptotyphlops

Bahndwivici FMNH PR2260

AigialosaurusTethysaurusTylosaurusPlotosaurus

Not Ascendonanus MNC TA1045

Saniwa ensidens FMNHEstesiaVaranus komodensisVaranus griseus

ShinisaurusHelodermaLanthanotusAnniellaGobiderma

OphisaurusKuroyuriella

MyrmecodaptriaElgaria

Cryptolacerta

Chalcides ocellatusChalcides guentheriGymnophthalmusVanzosauraSirenoscincusSlavoia MGR-I-112SineoamphisbaenaCrythiosaurusSpathorhynchus

DibamusTamaulipasaurus

BipesAmphisbaena

BarberenasuchusEodromaeus

Sakurasaurus

LimnoscelisOrobatesTseajaiaTetraceratopsMilleretta RC14Milleretta RC70

BashkyroleterMacroleterEmeroleterLanthanosuchus

UrumqiaBruktererpetonThuringothyris MNG 7729Thuringothyris MNG 10183

Reiszorhinus

CephalerpetonOpisthodontosaurus

Concordia KUVP 8702aConcordia KUVP 96/95Romeria primus

Saurorictus

Romeria texanaProtocaptorhinusParacaptorhinusEocaptorhinusCaptorhinusLabidosaurus

Calanguban

Acanthodactylus

EolacertaLacerta

TupinambisDorsetisaurus

Slavoia holotype ZPAL MgR-I/8Slavoia ZPAL MgR-III/77MacrocephalosaurusTianyusaurus

Diadectes CM25471Silvadectes absitus MNG 8853PhonodusBolosaurusBelebeyOradectes sanmiguelensisDiadectes UC706Diadectes sideropelicusProcolophonKapesScolaparia NSM 99 GF82.1LeptopleuronPentaedrusaurusHypsognathus

Youngina BPI 375Prolacerta AMNH 9502IxalerpetonCzatkowiellaMalerisaurusProlacerta BPI 1/471OzimekProtorosaurus NHMW 1943I4*BoreopriceaJaxtasuchusProtorosaurus NMK S 180Protorosaurus WMsN P47361Pamelaria

ZupaysaurusMegapnosaurus

StephanospondylusProvelosaurusDeltavjatia KPM232ScutosaurusBradysaurus NHMWAnthodonPareiasuchusPumiliopareia

JZC Bu1803 - amberLiushusaurusPurbicella

EuposaurusIguana

TriocerosPhrynosoma

BasiliscusAnolis

Pristidactylus

Chlamydosaurus

JKZ Bu267 -amber

Tawa

Staurikosaurus

SegisaurusZuolong

MarasuchusProcompsognathus

Dracoraptor

CoelophysisDilophosaurus

Sinocalliopteryx

Buriolestes CAPPA/UFSM 0035

VelociraptorBalaurHaplocheirusShuvuuiaHalszkaraptorSinornithoidesSauronithoides

Jinfengopteryx Anchiornis LPM B00169 Anchiornis PKUP V1068Anchiornis IVPP V14378

AurornisCaihongBuitreraptorSinovenator

Eosinopteryx

Xiaotingia

ElliotsmithiaApsisaurusAerosaurusVaranopsVaranodon

Varanosaurus FMNH PR 1760Varanosaurus BSPHM 1901 XV20ArchaeothyrisOphiacodon

HaptodusPantelosaurus

SecodontosaurusDimetrodonSphenacodonIanthodonEdaphosaurusIanthasaurusGlaucosaurus

CutleriaStenocybusIVPP V18117KenyasaurusGalechirusSuminiaVenjukoviaEodicynodonDicynodon

HeleosaurusMycterosaurus FMNH UC169NikkasaurusMesenosaurus PIN 158/1Niaftasuchus

Beipiaosaurus

RahonavisJianchangosaurusLimusaurusCaudipteryxKhaan

JuravenatorSinosauropteryx

Compsognathus holotypeAorunStruthiomimusGallimimus

OrnitholestesScriurumimusMicroraptor IVPP V 13352Microraptor M V1002Sinornithosaurus holotypeSinornithosaurus NGMC 91

FukuivenatorMasiakasaurusTianyuraptorHuaxiagnathusZhenyuanlongAlioramus altai

GorgosaurusRaptorex

Tyrannosaurus

Reptile Family Tree - Peters 2018 1326 taxa, 231 characters

EoraptorPampadromaeus

LeyesaurusSaturnaliaPantydracoEfraasiaMassospondylus adultMassospondylus embryoYizhousaurus

ChilesaurusDaemonosaurusJeholosaurus

JesairosaurusHypuronector

VallesaurusAvicranium

Megalancosaurus

TridentinosaurusCoelurosauravusMecistotrachelosLanthanolaniaIcarosaurusKuehneosaurusXianglong

Owenetta rubidgeiOwenetta kitchingorumCandelaria

FeeserpetonAustralothyrisEocaseaDelorhynchusMicroleterAcleistorhinusEunotosaurusEorhynchochelys

OedaleopsEothyrisColobomycter vaughniColobomycter pholeterDatheosaurusEnnatosaurusCaseaCotylorhynchus

Placodus inexpectatus

HipposaurusBiarmosuchusTitanophoneusProcynosuchusThrinaxodonProzostrodonProbainognathusChiniquodonCastorocaudaHaldanodon

Liaoconodon

Pachygenelus

RiograndiaChaliminia

OligokyphusKayentatheriumTritylodonSpinolestesJeholodens

GobiconodonRepenomamus robustusRepenomamus giganticus

Yanoconodon

SinoconodonBrasilodonHaramiyavia

Therioherpeton

Megazostrodon

Hadrocodium

Dromiciops

Asioryctes holotype

Monodelphis domestica

Marmosa

Chironectes

Docofossor

Volaticotherium

Alcidedorbignya

Maotherium

Maelestes

AnagaleLeptictisRhynchocyonHarpagolestes macrocephalusSinonyx

AndrewsarchusHemicentetesTenrecLeptictidiumIndohyus

Onychodectes

EctoconusPantolambda

BarylambdaGlyptodonParamylodonBradypusPeltephilusFruitafossor

Orycteropus

Myrmecophaga

HolmesinaChlamyphorus

DasypusTamanduaCyclopesMeniscotherium

OrthaspidotheriumTitanoides

AstrapotheriumPhenacodusThomashuxleya

GobiatheriumArsinoitheriumCoryphodonUintatherium

MesonyxHarpagolestes uintensisOcepeia

HomalodotheriumMiocochiliusProtypotherium

Litolophus

EmbolotheriumEotitanops

MetamynodonAceratherium

Hyracotherium

HyracodonUintaceras

Hyrachyus

Mesohippus

Pappoceras

Equus

JuxiaParaceratherium

Ceratotherium

Anisodon grande

PakicetusMaiacetus

Mammalodon

CoronodonAetiocetus

Zygorhiza

Mystacodon

Physeter

MonodonDelphinapterusTursiopsOrcinus

Hippopotamus

AnthracobuneCambaytheriumCornwallius

Janjucetus

PaleoparadoxiaNeoparadoxia

Desmostylus

Behemotops RBCM sp.

EschrichtiusIsanacetusBalaenoptera

TokarahiaYamatocetus

CetiotheriumSitsqwayk

CapereaEubalaena

ArganacerasSphodrosaurusSclerosaurus

Odontochelys

TrionyxPerochelys

Ocepechelon

Bunostegos

Elginia mirabilisElginia wuyongae

NiolamiaMeiolaniaProterochersisProganochelys

KayentachelysBaena

Kallokibotion_UBB ToK-2Arvinachelys

GlyptopsMongolochelysChubutemysCheloniaSantanachelysDermochelys

Macrochelys temminckiiTerrapene carolinaFoxemys

Carettochelys

PelomedusaAraripemysChelus

SaichangurvelMagnuviator

PleuraspidotheriumPeriptychus

Nambaroo

?Hapalodectes IVPP V2385

70

Anthracodromeus

Reptilia

UtegeniaBalanerpetonCaerorhachisAmphibamusPlatyrhinops

DendrerpetonCochleosaurus

TersomiusAchelomaPlatyhistrix

StegopsDissorophus

BroiliellusCacops

EcolsoniaFedexiaTambachia

GeorgenthaliaMicromelerpetonDoleserpetonApateonGerobatrachusAndriasCeltedensRana Utaherpeton

TuditanusKirktonecta

SauropleuraPtyonius

BatropetesRhynchonkosAsaphesteraMicraroterEuryodus primusPantylus

ScincosaurusBatrachiderpetonKeraterpetonDiceratosaurusDiplocaulusDiploceraspis Acherontiscus

MicrobrachisAdelospondylusAdelogyrinusChinlestegophisBrachydectesLysorophusEocaeciliaDermophis

CheirolepisEusthenopteronGogonasusOsteolepis

PholidogasterColosteus

PanderichthysTiktaalik

GreererpetonKoilopsSpathicephalusTrimerorhachis

DvinosaurusBatrachosuchusGerrothoraxNeldasaurusTrematosaurusMetoposaurusRhineceps

WantzosaurusTrematosaurus long rostrum

OssinodusAcanthostegaMGUH-VP-8160

DeltaherpetonVentastega

Crassigyrinus

PederpesWhatcheeria

BaphetesMegalocephalusSclerocephalus smns90055IberospondylusEryopsNigerpetonSaharastegaIchthyostegaProterogyrinusEoherpetonNeopteroplaxAnthracosaurus

Eucritta

DiscosauriscusSeymouria

MakowskiaSeymouria.tadpole

Guaibasaurus

Miohippus

Pectodens

AncodusLambdotherium

SusCainotherium

AnthracotheriumArchaeotherium

Danjiangia

Ectocion ralstonensis

NotostylopsProcavia

DusisirenProrastomus

PaleomastodonGomphotheriumElephas

Moeritherium

Amynodon

Note: This tree does not support DNA topologies over long phylogenetic distances.Certain dental traits are convergent and do not define clades.

HeptodonLophiodonTapirusProtapirus

Parotosuchus

OestocephalusLethiscus

RileymillerusPhlegethontia CGH129Phlegethontia AMNH6966

Acroplous

Eusaurosphargis juvenile

Fostercooperia

Junggarsuchus

CarnufexPseudhesperosuchusTrialestes

Asioryctes long rostrum

AriekanerpetonKotlassia

PseudocrypturusApteryx

RhynchotusEudromia

StruthioPatagopteryx

CasuariusRhea

CariamaPhoenicopterusSylviornis

SagittariusFalcoPandion

ButeoTorgos

ApusParargornis

TytoAegotheles

Llallawavis

Yanornis

ChangzuiornisJiangchangornis

LongicrusavisJinguofortis

Juehuaornis

RhamphocephalusHongshanornisDingavisCryputurusMegapodius

ApsaravisIchthyornisGansusHesperornis

CoccyzusMonias

Ardea

Ardeotis

Eozygodactylus

Butorides

AnhimaPteroclesFulica

Crex

AepyornisPalaelodus

PasserLonchuraTetrao

GallusPavo

EogranivoraMeleagris

Crax

OpisthocomusMacrocephalon

Dinornis

Ara macaoMelopsittacusNestor

Gastornis

AramusRallus

GaviaAechmophorus

Thalasseus

Charadrius

BalearicaJacanaHimantopus

EurypygaGrus

ChroicocephalusEocypselusArchilochus

MegaceryleJabiru

CinclusTroglodytes

MelanerpesSitta

UriaAptenodytes

Geococcyx

PsophiaMusophaga

MenuraSemiopteraParadisaea

CorvusTurdus

Quiscalus

Ciconia

Hirundo

Cyanocitta

Rhynochetos

PalaechthonCynocephalus

ChriacusOnychonycteris

RousettusMyotisChoeronycteris

ZhangheotheriumMetachcheiromysManis

Archaeopteryx siemensii Thermopolis

Archaeopteryx albersdoerferi Daiting

Archaeopteryx siemensii BerlinLiaoning embryo birdArchaeopteryx LondonChongmingiaCathayornisSulcavisPengornisChiappeavis

Yanornis STM9-52

IberomesornisProtopteryx

SerikornisArchaeopteryx recurva (Eichstätt)JianianhualongLiaoningvenator

Sapeornis IVPP V13276Sapeornis DNHM-F3078

Confuciusornis GMV-2132

Archaeopteryx #11

Confuciusornis MBAv1168Changchengornis

Zhongornis

ArchaeornithuraVegavis

Ostromia - Haarlem

Oedicnemus

Falcarius

DeltatheriumBishops

Caluromys

Ectocion cedrus UM 8615

MicromeryxGiraffa

Camelus

98

Miocaperea

Chonecetus

Archaeopteryx 12 private

Archaeopteryx bavarica (Munich)Jeholornis primaJeholornis curvipesHong Kong specimen

Mei

ScansoriopteryxYi

EpidexipteryxOmnivoropteryx

Archaeovolans (=Yanornis)

Draco volansMoloch

Lyriocephalus

MegachirellaBRSUG 29950-12

Pleurosaurus

MarmorettaGephyrosaurusAnkylosphenodonPaleopleurosaurusPlanocephalosaurusHeleosuchusKallimodonSphenodonBrachyrhinodonClevosaurusSphenotitanLeptosaurusVadasaurusSapheosaurusNoteosuchus

TrilophosaurusAzendohsaurusShringasaurus

EohyosaurusMesosuchusPriosphenodonRhynchosaurus articepsBentonyxHyperodapedon

Colobops

SaurosternonPalaegama

PolychrusJZC Bu154 in amber

Archaeopteryx (Wellnhoferia) grandis

Buriolestes ULBRA-PVT280

PteroglossusBuceros

Septencoarcias

Pelecanus

Catharacta

Balaeniceps

Fregata

PelagornisPuffinus

MacronectesMorus

Fraterculus

RynchopsPinguinus

Psilopogon

Cyrilavis

Phaethon

Upupa

Scopus

Phalacrocorax

ThreskiornisPlatalea

AnasAnserMergusPresbyornisPumiliornis

ColumbaCaloenas

PezophapsRaphus

AptornisDidunculus

CoragypsVultur

Goura

Lepidosaurmorpha

Archosauromorpha

JuramaiaChaoyangodensBrasilitheriumKuehneotherium

CifelliodonTachyglossusAkidolestesOrnithorhynchusUkhaatherium

Morganucodon

EomaiaAgilodocodon

AcristatheriumCronopio

DidelphisAmbolestes

Arctocyonnot AmphicyonThylacinus

Thylacosmilus

OxyaenaVincelestes

HyaenodonErnanodonDidelphodonMasrasectorBorhyaena

PeramelesMacrotisAnebodon

Notoryctes

InteratheriumMesotherium

Procoptodon

Dendrolagus

Proargyrolagus

Macropus

EurygeniumToxodon

PyrotheriumVombatusPhascolarctos

PaedotheriumGroeberiaVintanaBalbarooPhalangerPetaurusThylacoleo

Sinodelphys

TheosodonMacrauchenia

Chalicotherium? goldfussi

PaludidracoCymatosaurus

Lactodens

?Hapalodectes IVPP V5235NotharctusIndriRooneyiaDarwiniusTarsiusAegyptopithecusProconsulPanGorillaArdipithecusHomo sapiens

Ptilocercus

Tupaia

MacroscelidesChrysochlorisNecrolestes

ApatemysTrogosusRhyncholestesScutisorexLimnogaleMicrogale cowaniPotamogaleMicropotamogaleZalambdalestesSolenodonHenkelotheriumGomphosOryctolagus

BrachyerixErinaceusEchinopsThomomysHeterocephalusCeratogaulusCoendouMaiopatagiumNeoreomysChinchillaCaviaHydrochoerus

ShenshouVilevolodonRattusMusParamysRatufa

PaulchoffatiaIgnaciusPlesiadapisDaubentoniaCarpolestesMegaconusRugosodonXianshouKryptobaatarPtilodusYubaatarTaeniolabis

Mirischia

ColiusPharomachrus

BrachiosaurusCamarasaurus

DiplodocusRapetosaurus

Pteropus

VulpavusProtictisNandiniaProcyonAilurusProhesperocyonHerpestesTalpaMiacisHyopsodusEnaliarctosDesmatophocaPithanotariaZalophusProtelesCanisAmphicyon majorHesperocyonPantheraCrocutaAmphicyon galushiMustelaAiluropodaPuijilaNeotheriumUrsusGuloArctodusStylinodonAmphicynodonPsittacotheriumPalaeosinopaPhocaOdobenus

DiplovertebronRomeriscusBystrowiellaSolenodonsaurusLaosuchusChroniosaurus PIN3585/124ChroniosaurusChroniosuchus

HayaPisanosaurusScelidosaurusEmausaurusMinmiAgilisaurusStegocerasKulindadromaeusTianyulongHeterodontosaurusLesothosaurusScutellosaurusLiaoningosaurusStegosaurusChangchunsaurusDryosaurusEdmontosaurusLaquintasauraAquilopsHexinlusaurusYinlongPsittacosaurus mongoliensisLiaoceratopsAuroraceratopsLeptoceratopsStenopelix

Triceratops adultTriceratops juvenileChasmosaurus adultChasmosaurusjuvenile

72

86

9796

88

71100

8799

69

9498

82

70

9598

6264

64

79

7496

62

100 100

100<50

Compsognathus CNJ79

ProceratosaurusScipionyx

Deinocheirus

XiongguanlongSuchomimusSpinosaurusDilong

GuanlongMegaraptor

Yutyrannus

SinraptorMajungasaurus

AllosaurusSinosaurusMonolophosaurusAcrocanthosaurusEustreptospondylusCeratosaurus

Tanycolagreus

83

92

10067

9676

78

<50

62

9270

97

7180

6267

77

Anchiornis BMNHC PH804

Barlochersaurus

Llanocetus

EohupehsuchusParahupehsuchusHupehsuchus

88

100

74

87

82

89

95

95

87

75

79

98

87

100

97

87

100100 92

72

75

85

72

94

83

85100

100

100

98100

72

84

89

100

94

9860

100

91

85

100

73

6694

<50

99

80

<50 959960

63

82

99

7791 78

87

84

93

94

85

94

82

100

9194

99

9797

99

96

100

95

84

8093

84

60

95

9772

92

90

99

85

98100

89

97

98

99

9390

97

98

72

9991

98

57

65

54

57

80

62

81

88

95

89

89

100100

100

98

89

5592

81

70

90

7577

96

9564

51

100

<5091

93

7856

80

5585

7882

73

6460

95

62

98

62

8079

9897

66

84

8176

8955

7162

6094

99

90

7195

89

91

87

81

9396

91100

83

5792

9088

9495

95

76

7588

84

100

7998

62

57

80

65

89

72

75

78

55

100

95 90

67

93

99

86

92

76

6169

9696

99

83

100

7794

98

87<50

79

8368

100100

100

97

8698

77

9997

9998

83

83

100

92

57

64

79

9059

81<50

<5085

9281

100100

99

96

99

86

93

98

97

88

9497

100

9997

9464100

99

87

96

92

66

7377

99

97

81

100

91

100

97

88

98

100

100

90

90

98

96

97

9991

82

84

84

<50

87

51

93

10095

9688

7167

6569

6791

73

73

79100

100

95

94

100

79

10097

98100

95

86

62

60

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68

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6887

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57

65

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66

80

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78

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80

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76

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82

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67

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76

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82

87

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62

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52

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8199

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84

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Basal Therapsida59 taxa75 skull only characters

82

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Bootstrap scores are based on testing overlapping subsetsdue to limitations in computing power.

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PriacodonTrioracodon

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