stanford university jasper ridge biological …trees.stanford.edu/pdf/porter_rev.pdf · cirsium...

31
RESEARCH REPORT NO. 2 STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL EXPERIMENTAL AREA p'e pEl rtme nt.of Bio10eical ScLences THE VASCULAR PLANTS OF THE JASPER RIDGE BIOLCGICAL EXPERIMENTAL AREA OF STANFORD UNIVERSITY Duncan M. Porter INTRODUCTION Anril 1. 1962 (Revised slightly, March 26, 1971, by J. H. Thomas.) Jasper Ridge and Searsville Lake have long been an important collecting area for the botanists and students of Stanford University. Botanical collections were first made in this area about 1860 by H. N. Bolander and have continued to the present. Many theses and scientific papers have been written that bear partially or directly on the area. Those that have been used in the present study are those b,y Cooper (1922), Chaney (1925), Springer (1935), Moeur (1947), and Thomas (1958, 1961). The nomenclature used is the same as that in the latest flora of the area (Thomas, 1961). The standard floras of California (Abrams & Ferris, 1923-1960; Jepson, 1909-1943, 1925; MUnz, 1959) have been consulted. THE VEGETATION OF THE JASPER RIDGE BIOLCGICAL EXPERIMENTAL AREA The vegetation ,of Jasper Ridge may be divided into five distinct associations. These are oak-madrone forest, oak-woodland, chaparral, grasslands, and the aquatic flora of Searsville Lake. The areas of transition between the first four associa- tions may be either gradual or abrupt, so there mayor may not be an intermingling of their species complexes. In the following lists, only those species either confined to, or indicative of, an association will be listed for that association. The more densely wooded forest, occurring mainly upon the steep north-facing slopes of the ridge, may be termed the oak-madrone forest. The dominants are Quercus ap.rifo1ia and Arbutus Inp.nziesii. Indicative species are: trees: ~lercus a~rifo11a Quercus kelloe~ii Umbellularia ca1ifornica shrubs: Rosa californica Rosa evmnocarpa Rubus ursinus Holodiscus discolor herbs: Adiantum pedatU1l1 var. aleuttcum PitvroRramma trianp.ularts Pnlvstichum munttum Aesculus californica Arbutus menziesii Qsmaronia cerasiformis Rhamnus crocea ssp. crocea Dirce occidentalis AouileRia formosa Var. truncate Thelict.rum polvcaroum DAntaria californica

Upload: others

Post on 01-Aug-2020

8 views

Category:

Documents


0 download

TRANSCRIPT

Page 1: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

RESEARCH REPORT NO. 2

STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL EXPERIMENTAL AREA

p'e pEl rtme nt.of Bio10eical ScLences

THE VASCULAR PLANTS OF THEJASPER RIDGE BIOLCGICAL EXPERIMENTAL AREA

OF STANFORD UNIVERSITY

Duncan M. Porter

INTRODUCTION

Anril 1. 1962

(Revised slightly,

March 26, 1971, byJ. H. Thomas.)

Jasper Ridge and Searsville Lake have long been an important collecting area

for the botanists and students of Stanford University. Botanical collections

were first made in this area about 1860 by H. N. Bolander and have continuedto the present.

Many theses and scientific papers have been written that bear partially or

directly on the area. Those that have been used in the present study are those

b,y Cooper (1922), Chaney (1925), Springer (1935), Moeur (1947), and Thomas (1958,1961). The nomenclature used is the same as that in the latest flora of the area

(Thomas, 1961). The standard floras of California (Abrams & Ferris, 1923-1960;Jepson, 1909-1943, 1925; MUnz, 1959) have been consulted.

THE VEGETATION OF THE JASPER RIDGE BIOLCGICAL EXPERIMENTAL AREA

The vegetation ,of Jasper Ridge may be divided into five distinct associations.

These are oak-madrone forest, oak-woodland, chaparral, grasslands, and the aquaticflora of Searsville Lake. The areas of transition between the first four associa­

tions may be either gradual or abrupt, so there mayor may not be an intermingling

of their species complexes. In the following lists, only those species either

confined to, or indicative of, an association will be listed for that association.

The more densely wooded forest, occurring mainly upon the steep north-facingslopes of the ridge, may be termed the oak-madrone forest. The dominants are

Quercus ap.rifo1ia and Arbutus Inp.nziesii. Indicative species are:

trees: ~lercus a~rifo11a

Quercus kelloe~ii

Umbellularia ca1ifornica

shrubs: Rosa californica

Rosa evmnocarpaRubus ursinus

Holodiscus discolor

herbs: Adiantum pedatU1l1var. aleuttcum

PitvroRramma trianp.ularts

Pnlvstichum munttum

Aesculus californica

Arbutus menziesii

Qsmaronia cerasiformisRhamnus crocea ssp. croceaDirce occidentalis

AouileRia formosaVar. truncate

Thelict.rum polvcaroumDAntaria californica

Page 2: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Ptaridium aouilinumvar. oubescens

Desch~mpsia elon~ata

Fritillarta lanceolataDtsnorum hookeri

Smilecina ~acemose

var. amplexicaulis

Smilaeine stellate

var. sp-ssifoUa

Tr 1111um chlorooetalmn

Corallorhiza striata

HeMnarta unalaRcenstsvar. eleta

Silane c~rnica

var. 9alifornieaFra~aria ealifornica"

Anthriscus scancHclna

Senicula crassicaulis

Trientalis latifolta

Nemoohtla hl"lteronhvlla

PQolistoma 811ritum

Cvno~lossum Rranoe

Satl1reja d01U!lasU

Scroohularia cali forntca

Lontcera hispidula \Hierectum albiflorum

Adenocaulon bicolor

The streambank vegetation along San Francisquito Creek may be included as a sub­

unit of the oak-madrone forest association, as the two types here intermingle, with

many of the species common along the creek being found along its minor tributaries

in the oak-madrone forest. Some of these species are also found along Corte deMadera Creek and the moist ravines above Searsville Lake. These are:

trees: Seouoia sempl"Jrvirens

§Alli laevt!:!at,e

Salix lasiendra

Salix lasioleoi8

Alnus rhombifolia

shrubs: Corvlus californica

Whinolea modestaGrossularia divaricate

Grossularia leptosma

Phvsoearnus canitatus

Rubus parviflorusvar. velut.inus

herbs: Urtiea holosericea

Nastl1rtium offie inaleReuehera micrantha

Conium maculatum

Heracleum maximum

UmbelDllaria ealifornica

Acer macroohvlluJn

~ ME'-undoyare californica

Cornus californica

Cornus g,lahrata

Prunus emarRinataPrunus subcordata

Amelanchie~ oellide

Ellnon~us occtdentelisLonicera involucrata

Artemis is dOllE?lasiana

Mentha Pllle~ium

Mlmulus p.uttatnsBac~hArls dOllP-lasii

Helentum puberulumSenecio miklmioides

The more openly yooded forest, occurring upon the flatter, gently sloping areasof the ridge may be termed the oak-woodland. The characteristic species is Qnercusdouglasi!. Species indicative of this association are:

trees: Quercus douelastt

herbs: Calochortus Albus

Brodiaea lutp,a

Montia perfoliata

Delnhinium patens

Lathvrus vestitus

ssp. puberulusVieia americana

var. !!!.!.o.m:

Viela ext~ua

Clarkia unRuiculataCaucalls microcaroa

Castilleja affinis

~edicularis densiflora

Aster chtlensls

Madia exi!:!ua

Erionhvllum lanatumvar. arachnoideum

The chaparral association is found on the gentle south-facing slopes and flatter

-2-

Page 3: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

8~eas on the crest of the ridge. The dominants are Adenostoma fasclculatum, CeR"othuscuneatus var. d~bius, and Arotostapbylos crustacea var. crustacea. Indicative

species are:

shrubs: Quercus dumosa

9uercus dwata

gperou~ vislizeniivsr. frutescens

Clematis lastantha

RiMs malvaceum

~denostoma fasciculatum

geroocsrous betuloides

Prunus i1101fo118

Photlnta arbutifolia

Ceanothus ouneatus

vsr. dubius

herbs: DeJ.phinium.'~a.1ifornicumL(')tusscop~rius

Hesperollnon micr8nt~~

Hellanthemum scoparium

var. vulaare

Convolvulus occidentalis

Erlastrum abramsii

Navarretta heterodoxa

Scutella~is tuberosa

9aanothus soredietus

Ceanothus thvrsiflorus

~laoothamnus arcuatus

Garrva elliotica

Arctostaohvlos crustacea

var. orustacea

~riodictvon californicumLeoechinia cslvci.na

Diplacus eurantiacus

Artemisia oalifornlca

O~obanche bulbosa

Galium nuttell!i

Sambucus mexicana

1riodanus biflor~

Aster radulinus

Lavis hieractoides

Eriophvllum confertiflorum

Much of the Jasper Ridge, especially along its crest, consists of grasslands.

Most of this association is on flat or gently sloping land, but in many placesit extends up and over the steeper slopes where the original chaparral has been re­

moved to provide for more grazing area. Here and there, especially on the steeper

slopes, individuals of Quercus sQrifolia, Q. douelasi\, and Q. kelloei~ may beencountered. Typical grassland herbs are:

Brf-za minor

Bromus c~rinatus

Bromus mollis

Bromus racemosusBromus rubens

testuoa mvuros

Melioa imOArfect~

Poa scabrella

Elymus Qlaucus

Hordeum hvstrix

Lolium ~lttflorum

Lolium tremulentum

Airs earvoohvlla

Avena barbat.a

Avena. fatua

Holeus lanatusGastridium ventricosum

Calohortus luteus

Brodiae~ ele~ans

Brodiaea hycinthinsBr.odiaea.laxe

Brodiae~ pulchella

SlsyrinchiWTJ bell1lm

~r10gonum nudum

Ertoeoovm vimineum

-3-·

THleea erects

Alchemilla occidentalis

Lotus ID+cranthus

Lotus purshianus

Lotus suhoinnatus

Luoinus densiflorus

Lupinus bicolor

var. mierophvllus

Lupinus nanusTrifolium barbi~erum

Trifolium Qifidum

.. var. dec.1plens .Trifolium eiliolatum

Trifoli~ fucatum

Trifolium gracilentum

Trifolium microcephalum

:frlfolium variegatum

Erodium botrys

Erodium circutarium

~rodium ohtuspUcatum

Qeranium dissectumGeranium molle

Eremocarpus seti~erus

Clarkia purourea

ssp gyadrivulnera

Page 4: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Ca1apdrinla ei1i~tavar. lllilnziesii

Month st>athulataCerastium viseosum

Silene Ralliea

Delphinium hesoerium

Delphinium var!e~atumvar. ealifornicus

Eschseholzia ea1iforniea

~tystemon californieus

Athvsanus pnsillus

Lepidium nitidumCentuneu1us minimusGilla e1ivorum

Llnanthus androsaceous

Linanthus bico10r

Linanthus dichotomus

Linanthus linif10rus

Llnanthus parviflorusNavarretia viseidula

Pholox eracilis

Nemop~i1a Menzies!!var. menziesii

Nemophila menzies!ivar. atomAria

Amsinekia intermedia

CX'v:ptanthamicromeres

Pla~iobothrys nothofulvus

P1aeiobothrys tenel1usMonarde11a vil10sa

var. vil10sa

Pop.o€vne seroy1loidesSalvia co1umbarae

.cordv1anthus pilosusLinaria texana

Orthocarpus attenuatus

Orthocarous densiflorus

Orthoearous erianthusvar. eriant,hus

Orthocarous ourourascens

var. purol~ascens

Orthocarous pusi11us

Planta~o erecta

Clarkia pnrpures

ssp. viminea

Oenothera Qvata

ErvnRium maculetumLomatium dasv~arpumLomat.hun utricnlatum

Senicula bipinnatifida

ssp. -pstu1umDodecatheon hendersonii

ssp. cruciatnm

Micl'ocale quadraMularis

Plect.ritisei1iosa

ssp. !nsi.p,nis

Plectritis macroc~A€oseris erandifld a

Hvpochaeris E!labraMlcroseris doup-lasii

ssp. doudasii

Chaetopappa a1sinoide~

Eri€eron fo1iosusGrindelia cemoorumGrindelia hirsutu1a

Lessin~ia germanorumval'. tenuioes

Lessinpia holo1euc~var. ho101euca

Evax soorsif10ra

Gnaohalium QuroureumMlcroous californiclls

Psi10carphlls tene11us

Fe1ianthel1a californica

Wyeth ia ammstifo1ia

Achyrachaene mollis.Hemizon i8 cOl'vmbosaHemizonia l11zulaefolia

ssp. l'udis

LaRonhv11a ramosissima

Lavie platye10ssa

ssp. camoestris

Madia BracHis

Baeria chrvsostoma

ssp. era~ilis

Many of the species listed above are found in the serpentine soil of the meadow

on the crest of Jasper Ridge, but those species in the following list are confined

solely .to this serpentine.

Festuca lJacifica

Sitanion jubatllln

Agrostis microphy11~

Stipa pu1chra

Junclls phaeoceoha1usCalochortus venustus

Allium amolectensAllium breweri

,Il.lliumlacunosllln

Brodiaea terl'estris

~r1~ dou~lasii

Astra~a1us gambel1ianus

Crvotantha f1accida

Mimulu~ doue1asii

Qrthocarous lithosn~rmoides

Calvcadenia mu1tig1andu10sa

ssp. robusta

Many weedy species have invaded Jasper Ridge that are confined to disturbed areas.

-4-

Page 5: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

These are in the main found in the well trampled land to the north and west of the

ridge abutting upon Searsville Lake, but many of them occur along the roadsides on

the ridge proper. Introduced weeds of disturbed areas are:

Bromus commutatus

Bromus rieidusEoa annua

Hordeum lenorinum

Cspsells ~-~~

Rsohanus sativus

Sisvmbrium officinalis

Melilotus indicus

Medica~o R91~o~ha

val'. ~llgAris f. vul~ar!s

Medica!:!osativa

Victa sativa

Malva parviflora

Convolvulus arvensi~

Planta~o lanceola~

D!osaeus fullonum

RUmex acetosellaRumex CriS01,lS

Brasslcs ~amnestris

Brassles nip-ra

Picris ft£h1oidesSonehus esper

§onchus olereceusTeraxacum officinale

Anthemts cotula

Senecio vulgarisCarduus tenuiflorus

Centaurea ealcitrepa

Centaurea melitensisCentaurea solstitialis

Cirsium vul~are

SilybuJIlmarianum

.The aquatic and simiaquatic plants in and around Searsville Lake comprise thelast association. These are:

Azolla filiculoides

Tvpha lat ifolia

SparQanum eurvcaroum

Potamo~eton folios~s

var. mecellus

Potamo~eton pusillusvar. minor

Alisma plantago-aQuat\ca

A~rostis exaratavar. exarata

Polvpogon monspeliensis

PaspaJll{lld isUchU!!!Scirpus scutusIris pseudoacornus

The following annotated list consists of:82 Families

273 Genera

458 Species, Subspecies, and Varieties

-5-

Polvgonum cocci.neum

Poly~onum laoathtfolium

Polvp-onum punctatumAtriolex patula

var. hasta:t,a

Lvthrum hvssopifolia

Epilobium adenocaulonval'. occidentale

Oenanth~ sarrnentosa

SaHdMo occi.dentalisGnanhalium oalustreXanthium strumarium

Artemisia biennis

Page 6: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

ANNOTATED LIST OF THE VASCULAR PLANTS

Class F111c1nae

1. Salviniaceae

*Azolla filiculoides Lam.Searsville Lake.

2. Polypodiaceae

Adiantum jordon! Muell.

Reported by Cooper (1922) from the oak-madrone forest.

~diantum oedatum L. var. aleuticum Rupr.

Dense oak-madrone forest on north-facing slope above San Francisquito Creek.*Pellaea mucro nata (Eaton) Eaton

x Sandstone outcroppings on north-facing slope above San Francisquito Creek.PitvroQramma trianQularis (Kaulf.) Maxon

Common in oak-madrone forest, occasional in chaparral.Polvstichum munitum (Kaulf.) Presl.

Dense oak-madrone forest on north-facing slope above San Francisquito Creek.

Pteridium aauilinum (L.) Kuhn var. pubescens UndeN.Oak-madrone forest.

Class Gymnospermae

1. Pinaceae

Xpseudotsuga menziesii (Mirb.) Franco

Scattered on the north-facing slope above San Francisquito Creek.

2. Taxodiaceae

~ .Seauoia semoervirens (Lamb.) Endl.

Common along San Francisquito Creek.

Class Angiospermae

Subclass Monocotyledoneae

1. Typhaceae

*Tyoha latifolia L.Searsville Lake.

2. Sparganiaceae

*SoarQanium eurvcarpum Engelm.Searsville Lake.

3. Potamogetonaceae

*PotamoQeton folios~ Raf. var. macelDls Fern.Searsville Lake.

*PotamoReton pusillus L. var. minor (Biv.) Fern. & Schub.

4. Alismataceae

-6-

Page 7: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*Alisma plantaeo-aquatica L.Searsville Lake.

5. Gramlneae

Festuceae

~ .Briza m~nor L •

.,r Grasslands and openly wooded· slopes.Bromus carinatus H. & A.

Grasslands and openly wooded slopes.Bromus comml1tatu~ Schrad.

Disturbed areas in grasslands.Bromus laevioes Shear

Reported by Springer (1935) from a "shady slope".*Bromus mollis L.

The most common grass in grasslands and on openly wooded slopes.Bromus racemosus L.

Grasslands and openly wooded slopes.Bromus riRidus Roth.

Disturbed areas in grasslands.Bromus ruoons L.

Grasslands.

Bromus tectorum L.

Reported by-Springer (1935) from "open field near picnic grounds".Festuca elmeri Scribn. & Marr.

Reported by Moeur (1947) from "wooded hillsides".

Festuca mvuros L.Grasslands and disturbed areas.

*Festuca occidentalis Hook.

"Moist areas in redwoods and redwood-Douglas fir forests •••• "

(Thomas, 1961). Collected by Rattan (~• .n.) at "Searsv1l1e."Festuca octo flora Walt.

Reported by Moeur (1947) from "openly wooded hillside".

*Festuca PAcifica Piper

Serpentine soil in meadow.*Melica imnerfecta Trin.

Oak madrone forest.

Melica torrevana Scribn.

~ Reported by Springer (1935) from "grassy field near picnic grounds".'Poa annua L.

Disturbed areas.

*~ scabrella (Thurb.) Benth. ex Vasey

Grasslands and openly wooded slopes.

Hordeae

,cElymus glaucus Buckl.

Grasslands and openly wooded slopes.

Hordeum brachvantherum Nevski

Reported by both Springer (1935) and Moeur (1947) as limited to the "serpentine ofthe meadow". Listed as H. nodosum L.

*Hordeum hystrix Roth.Grasslands.

Hordeum leporimlm LinkDisturbed areas.

-7-

Page 8: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

~Lo1ium mu1tif1orum Lam.

Grasslands and openly wooded slopes.Lo1ium tremu1entum L.

X Grasslands and openly wooded slopes.Sitanion iu~tum Smith

Serpentine soil in the meadow.

Aveneae

yAira sarvophv11~ L.

~ Grasslands and openly wooded slopes.Avena barbata Brot.

Grasslands and openly wooded slopes.Avena fatua L. var. fatue

Grasslands and openly wooded slopes.*Deschampsia danthonioides (Trin.) Munro ex Benth.

"Low moist areas where water has stood during the winter and early Sprlng •••• "

(Thomas, 1961). Collected by Mason (May 13, 1921) at "Jasper Ridge. n

*Deschamnsia elongata (Hook.) Munro ex Benth.Oak-madrone forest.

*Ho1cus lanatus L.

Grasslands.

Koe1eria mecrantha (Ledeb.) Spreng.

Reported by Moeur (1947) from "open fields and on openly wooded slopes". Listed8S !. cristata (L.) Pers.

'.Agrostideae

*A~rostis exarats Trin. var. exarataSearsville Lake.

*Aerostis microphv11a Steud.

Serpentine soil in the meadow.

Agrostis semiverticillata (Forsk.) Christ.

y Reported by Moeur (1947) from "wooded hillside". Listed as A. verticillata VilI.Gastridium ventricosum (Gouan) Schinz & TheIl.

1< Grasslands.Polvpo~on interruptus RBK

Reported by Moeur (1947) from "near the lake shore". Listed as f. lutosus (Poir)Hitchc.

*Polynogon monspeliensis (L.) Desf.

Searsville Lake. Abrams 1639 was collected at "Jasper Ridge."

*Stjpa pulchra Hitchc.

Serpentine soil in the meadow.

Phal~rideae

*Hierochloe occidenta1is Buckl.

"A common grass in redwoods and redwood-Douglas fir forests •••• " (Thomas, 1961).Rattan (March 1867) 'Wes collected at "Searsvll1e.1f

Panlceae

*Paspalum distichum L.Searsvi1le Lake.

6. Cyperaceae

Carex barbarae Dewey

Reported by Moeur (1947) from "creek 'tank".

-8-

Page 9: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Cvperus eraqrostis Lam.

Probably Moeur's ~ virens Michx. from "San Francisquito Creek".*Scirpus acutus Muhl. ex Bigel.

Searsvi1le Lake

7. Juncaceae

~Juncus occidentalis (Cov.) Wieg.

Reported by Moeur (1947) from "San Francisquito Creek". Listed as J. tenuis

var. conqesta Engelm.

Juncus patens Meyer

San Francisquito Creek ann wet grasslands.

Juncus phaeocephalus Engelm.Serpentine soil in the meadow.

~Luzula multiflora (Retz.) Lejeune

Oak-madrone forest and openly wooded slopes.

8. Melanthaceae

1<Zvqadenus fremontii (Torr.) Torr. ex Wats.· var. fremontii

Wooded hillsides and chaparral.

9. Liliaceae

*Calochortus albus Dougl. ex Benth.Oak woodland.

*Calochortus luteus Dougl. ex Lindl.

Grasslands and openly wooded slopes.

*Calochortus venUStU5 Dougl. ex Benth.

Serpentine soil in the meadow.

*Chloroqalum pomeridianum (DC.) KunthGrasslands and edges of chaparral.

)r Frit illar ia lanceolata Pursh

Oak-madrone forest.

Lilium pardalinum Kell.

Reported by Moeur (1947) from "deep shade of the redwood area".

10. Convallariaceae

*Disporum hookeri (Torr.) Nichols.

Oak-madrone forest on north-facing slope above San Francisquito Creek.

*Scoliopus biqelovii Torr.

"Moist slopes and along creek banks in redwood forests •••••• (Thomas, 1961).

Collected by Rattan (March 1867) at "Searsville."

*Smilacina racemosa (L.) Desf. var. amplexicaulis (Nutt.) Wats.

X Oak-madrone forest.Smi1acina ste11ata (L.) Desf. var. sessi1ifo1ia .(Baker) Henders.

X Oak-madrone forest.lrillium chloropetalum (Torr.) Howell

Oak-madrone forest, especially common along streambanks.

11. Amaryllidaceae

*Allium amplectens Torr.

Serpentine soil in the meadow.

-9-

Page 10: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*Al1ium breweri Wats.

Serpentine soil of the meadow and adjacent slopes.

*Al1ium dich1amydeum Greene

Sandstone outcroppings on north-facing slope above San Francisquito Creek.*Allium lacunosum Wats.

Serpentine soil in the meadow.Allium serratum Wats.

Serpentine soil of the mendow and adjacent slopes.*Brodiaea e1eQans Hoover

Grasslands.

~Brodiaea hyacinthina (Lindl.) Baker

x Grasslands and openly wooded slopes.rBrodiaea laxa (Benth.) V·fats.

Grasslands and openly wooded slopes.

Brodiaea lutea (Lindl.) Mort.Oak woodland.

*Bro~iaea pulchella (Salisb.) Greene

Grasslands and openly wooded slopes.*Brodiaea terrestris Kel1.

Serpentine soil in the meadow.

12 •. Iridaceae

Iris macrosiphon Torr.

Reported by Springer (1935) from "edge of the road through the chaparral abovethe lake."

*Iris pseudacorus L.

~ Searsville Lake.?isyrinchium bellum Wats.

Grasslands and openly wooded slopes.

13. Orchidaceae

*Corallorhiza striata Lindl.Oak-madrone forest.

~abenaria unalascensis (Spreng.) Wats. var. ~ (Jeps.) CorrellOak-madrone forest.

Subclass Dicotyledoneae

14. Salicaceae

Populus trichocarpa T. & G. ex Hook.

Reported from San Francisquito Creek by Chaney (1925).Salix hindsiana Benth.

~ported by Moeur (1947) from "edge of road above lake."

*~ laeviQata Bebb.Corte de Madera Creek, San Francisquito Creek, and marshy area at southern end ofSearsville Lake.

*Salix lasiandra Benth.

Streambanks and marshy area at southern end of Searsville Lake.

*Salix 1asio1epis Benth.

Streambanks and marshy area at southern end of Searsvi11e Lake.

15. Juglandaceae

-10-

Page 11: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Juqlans hinds ii Jeps.

Reported from the edge of San Francisquito Creek by both Chaney (1925) (as J.californica wats.) and Moeur (1947). Specimens of this species are unknownfrom the Santa Cruz Mountains.

16. Betulaceae

~Alnus rhombifolia Nutt.

San Francisquito Creek.

17. Corylaceae

*Corylus californica (A. DC.) Rose

Shady streambanks.

18. Fagaceae

*Quercus aqrifolia Nee

The principal oak of the oak-madrone forest, to a lesser extent in the oak­

woodland and on openly-wooded slopes.

*Quercus douqlasii H. & A.

The prinicpal oak of the oak-woodland and the openly-wooded slopes, occasionallyin the oak-madrone forest.

*Quercus dumosa Nutt.

Chaparral.

*Quercus durata Jeps.

Chaparral.

*Quercus kelloqqii Newb.Oak-madrone forest and oak-woodland.

Quercus lobata Nee

Openly wooded slopes on the lower edges of Jasper Ridge, occasional.

*Quercus X morehus Kell.

Collected by Fisher (March 19, 1898) on "Jasper Ridge."

*Quercus wislizenii A. DC. yare frutescens Engelm.

Chaparral.

19. Urticaceae

)f Urtica holosericea Nutt.

Streambanks.

20. Loranthaceae

*Phoradendron villosum Nutt.

Parasitic on oaks (Quercus aqrifolia, Q. douqlasii, Q. lobata), and occasionallyon Adenostoma fasciculatum.

21. Polygonaceae

*Chorixanthe diffusa Benth.

Dry rocky areas in chaparral and grasslands.

~ Erioqonum nudum Dougl. ex Benth.Grasslands.

*Erioqonum vimineum Dougl. ~x Benth.Grasslands.

Polyqonum aviculare L.

Reported by Maeur (1947) from "edge of road above the lake."

-11-

Page 12: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*PolYQonum coccineum Muhl. ex Wi11d.Searsville Lake.

*Polyqonum lapathifolium L.Searsv1.Ue Lake.

*PolYQonum punctatum Ell.Searsville Lake.

*Rumex acetose1la L.

Disturbed areas.

Rumex conQlomeratus Murr.

Reported by Moeur (1947) from "along roadsides and near the lake."

Rumex crispus L.Disturbed areas around Searsvi1le Lake.

Rumex obtusifolius L.

Reported by Springer (1935) from "roadsides, near the lake, and near buildings,"

and by Moeur (1947) from "edges of the lake and along roadsides."

22. Chenopodiaceae

*Atriplex patula L. var. hastata (L.) GraySearsville Lake.

23. Portulacaceae

*Calandrinia ciliata (R. & P.) DC. var. menziesii (Hook.) Macbr.Grasslands.

~Montia perfoliata (Donn ex Willd.) Howell

Oak-woodland and moist openly wooded slopes.

*Montia spathulata (Dougl.) Howell

Moist grassl?nds and openly wooded slopes.

24. Caryophyllaceae

Arenaria douQlasii Fenzl ex T. & G.

~ Serpentine soil in the meadow.Cerastium viscosum L.

~ Grasslands and disturbed areas.Silene californica Durand

Oak-madrone forest.

§ilene Qallica L.Grasslands.

*Sperqularia marina (L.) Griseb. var. marina

~ Baker (1959) was collected at "Searsville."Stel1aria media (L.) Cyril1

Grasslands, openly wooded slopes, oak-woodland, and oak-madrone forest.

25. Ranunculaceae

Actaea arquta Nutt .ex T. & G.Occasional in dense oak-manrone forest on north-facing slope above San FrancisquitoCreek.

~Quileqia forrnosa Fisch. var. truneata (F. & M.) JonesOak-madrone woodland.

*Clematis lasiantha Nutt.

Chaparral.

Clematisliqust ieifolia Nutt.

Reported by Springer (1935) from "bank of San Francisquito Creek."

-12-

Page 13: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*Delphinium californicum T. & G.

Chaparral.

Delphinium hesperium GrayGrasslands.

*Delphinium patens Benth.Oak-woodland.

~lphinium varieqatum T. & G.

Grasslands and openly wooded hillsides.*Ranunculus californicus Benth. var. californicus

Grasslands and openly wooded hillsides.

Ranunculus hebecarpus H. & A.

Reported from the oak-madrone forest by Cooper (1922).Ranunculus muricatus L.

Reported by Springer (1935) from "moist shady places near the banks of San Fran­

c isquito Creek".

*Thalictrum polycarpus (Torr.) Vlats.

Oak-madrone forest, occasional in the chaparral.

26. Berberidaceae

*Berberis pinnata Lag.

Sandstone outcroppings on north-facing slope above San Franeisquito Creek.

27. Lauraceae

i*Umbellularia californiea (H. & A.) Nutt.

Streambanks and oak-madrone forest.

28. Papaveraeeae

~ Eschscholzia ealifornica Cham.

Grasslands.

*Platvstemon californieus Benth.Grasslands.

29. Crue iferae

Athysanus pusillus (Hook.) GreeneGrasslands.

Brassica campestris L.

Disturbed areas, occasionally in grasslands.

Brassica kaber (DC.) Wheeler

Reported by Moeur (1947) from "open fields and disturbed areas."

Brassica niQra (L.) KochDisturbed areas.

*Cardamine oliqosperma Nutt.

Oak-woodland and moist openly wooded slopes.

Capsella bursa-pastoris (L.) Medic

Disturbed areas and grasslands.*Dentaria californica Nutt. var. californica

Oak-madrone forest.

)CLepidium nitidum Nutt.Grasslands.

*Nasturtium officinale R. Br.

Streambanks.

-13-

Page 14: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Raphanus sativus L.Disturbed areas.

~Thvsanocarpus curvipes Hook. var. curvipes

Openly wooded hillsides.

lhvsanocarpus curvipes Hook. var. eleqans (F. & M.) Robins.

Reported by Springer (1935) from "openly wooded hillside."

Jropidocarpum qracile Hook.

Reported by Springer (l935) from "open hillsides, especially near the edges of

areas of chaparral."

30. Crassulaceae

~edum spathulifolium Hook.

Reported by Moeur (1947) from "shaded moss-covered rocke in the heavily woodedforest."

Jillaea erecta H. & A.Grasslands.

31. Saxifragaceae

Heuchera micrantha Dougl. ex Lind1.

San Francisquito Creek.

Lithophraqma affinis Gray

Oak-madrone forest, oak-woodland, and openly wooded slopes.

Lithophraqma heterophvlla (H. & A.) T. & G.

Reported from the oak-madrone forest by Cooper (1922) (as Tellima heterophylla).

*Saxifraqa californica Greene

Oak-madrone forest, oak-woodland, and openly wooded slopes.

)CTellima qrandiflora (Pursh) Dougl. ex Lind1.Reported by Moeur (1947) from "deep shade of tha redwood zone."

32. Hydrangeaceae

*Whipplea modesta Torr.

San Francisquito Creek.

33. Grossulariaceae

*9rossularia californica (H. & A.) COv. & Britt.

Oak-madrone forest, oak-woodland, openly wooded hillsides, and occasionally at

edges of the chaparral.

*Grossularia divaricata (Dougl.) COv. & Britt.Streambanks.

*Grossularia leptosma Cov.Streambanks.

~Ribes q1utinosum Benth.

Reported by Moeur (1947) from "thicket at the edge of the lake."*Ribes malvaceum Smith

Chaparral.

34. Rosaceae

*Adenostoma fascicul~tum H. & A.

The index plant of the chaparral.

~Alchemilla pccidentalis Nutt. ex T. & G.Grasslands.

·-14-

Page 15: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*Cercocarnus betuloide~ Nutt. ex T. & G.

Chaparral.*Fra~aria californica C. & S.

Oak-madrone forest on north-facing slope above San Francisquito Creek.XHolodiscus discolor (Pursh) Maxim.

Oak-madrone forest and streambanks.

~Phvsocarpus capitatus (Pursh) KuntzeStreambanks.

~Potentilla Rlandulosa Lindl.

Oak-yoodland and openly wooded hillsides.Rosa californica C. & S.

--O;k-madrone forest.

*Rosa Rymnocarpa Nutt. ex T. & G.Oak-madrone forest.

Rosa spithamea Wats.

Reported by Springer (1935) from "openly wooded slope at the top of the hill nearthe lake."

Rubus oarv1florus Nutt. var. velutinus (H. & A.) GreeneStreambanks.

*Rubus ursinus C. & S.

Oak-madrone forest, especially along streambanks.

35. Amygdalaceae

~Osmaronia cerasiformis (T. & G. ex H. & A.) GreeneOak-madrone forest.

*Prunus ~marRinAta(Dougl.) Walp.Streambanks.

*Prunus 1licifolia (Nutt. ex H. & A.) Walp.Chaparral.

*Prunus subcordata Benth.Streambanks.

36. Malaceae

*Amelanchier pallida GreeneStreambanks.

*Photinia arbutifolia (Ait.) Lindl.

Chaparral, occasionally in oak-yoodland or oak-madrone forest.

37. Fa baceae

*Astragalus Rambellianus ~heldon

Serpentine soil in the meadow.

*Cvtisus maderensis ~~sf.

~ Thomas ~ yas collected~in oak-woodland."Glvcyrrhiza 1epidota Pursh yare ~lutinosa (Nutt.) Wats.

Reported by Springer (1935) from "field that was dry, but close to the marshy

part of the lake."

Lathvrus vestitus Nutt. ex T. & G. ssp. puberulus (White ex Greene) Hitchc.Oak-woodland.

Lupinus bicolor Lindl. var. microphvllus (Wets.) SmithGrasslands.

Lupinus densiflorus Benth.Grasslands.

Lupinus formosus Greene var. formosus

Reported by Moeur (1947) from "open fields."

-15-

Page 16: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

~upinus ~ Dougl. ex Benth.Grasslands.

Lotus humistratus Greene

Reporte~ by Moeur (1947) from "grassy hillsides."*Lotus micranthus Benth.

Grasslands and openly wooded slopes.

*Lotus purshianus (Benth.) Clem. & Clem.

Grasslands and openly wooded slopes.

*~otus scoparius (Nutt.) OttleyChaparral.

*Lotus striqosus (Nutt.) Greene

Collected by Davis (April 10, 1903) on "North slope, Searsville Ridge."

*Lotus subpinnatus Lag.Grasslands.

Melilotus albus Desr. ex Lam.

- Reported by Moeur (1947) from "along creek banks."

Melilotus indicus (L.) All.Disturbed areas.

Medicaqo arabica (L.) Huds.Disturbed areas.

YMedicaQo pOlymorpha L. var. vulQaris (Benth.) ShiMers f. vulqaris

Disturbed areas and grasslands.MedicaQo sativa L.

y Roadsides and disturbed areas.Psoralea physodes Dougl. ex Hook.

)( Oak-woodland and oak-madrone forest.I!ifolium albopurpureum T. & G.

Reported by Moeur (1947) from "moist, grassy fields."*Trifolium barbiQerum Torr.

Grasslands and openly wooded slopes.

*Irifolium bifidum Gray var. decipiens GreeneGrasslands and openly wooded slopes.

Irifolium ciliolatum Benth.Grasslands.

*Trifolium fucatum Lindl.

Thomas (1958, p. 460) states, "On Jasper Ridge this plant is very common and

almost a weed in low moist areas, especially where cows have ~vergrazed and have

trampled the soil dur ing the rainy season." Grasslands.

Trifolium qracilentum T. & G.

Grasslands and openly wooded slopes.

'Trifolium microcephalum PurshGrasslands.

*Trifolium tridentatum Lindl.

Grasslands, openly wooded slopes and oak-woodland.

*Trifolium varieqatum Nutt.Grasslands.

Trifolium wormsk;oldii Lehm.

Reported by Moeur (1947) from "along stream banks."

*Vicia americana Muhl. ex Willd. var. miD£r Hook.Oak-woodland.

Vicia anqustifolia L.

Reported by Moeur (1947) from "fields and along roadsides."

Vicia exiqua Nutt.Oak-woodland.

~..?.ili sativa L.Disturbed areas.

Vicia villosa Roth.

Reported by Moaur (1947) as "mostly along roadsides."

-16-

Page 17: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

38. Geraniaceae

)(Erodium botrys (Cav.) Berto1Grasslands.

Erodium circutarium (L.) L'Her.Grasslands.

Erodium moschatum (L.) L'Her. ex Ait.

Reported by Moeur (1947) from "fields and pasture lands."

*Erodium obtusip1icatum (Maire, Wei11er, & Wilczek) HowellGrasslands.

~Geranium dissectum L.

Grasslands.

*Geranium molle L.

Grasslands and openly wooded slopes.

39. Oxa lidaceae

*Oxalis oreqana Nutt.

P~a.ulson(Apr. 1915) was collected at "Jasper Ridge near Searsville Lake."

40. Linaceae

*Hesperolinon micranthum (Gray) Small

Chaparral.

41. POlygalaceae

Po1yqala californica Nutt.

Reported by Springer (1935) from "under the shade of some of the chaparral shrubs

growing on a lightly northward slope."

42. Euphorbiaceae

*Eremocarpus setioerus (Hook.) Benth.

Disturbed areas in grasslands.

)( 43. AnacardiaceaeRhus diversiloba T. & G.

Common.

44. Celastraceae

*Euonvmus occidentalis Nutt. ex Torr.Streambanks.

45. Aceraceae

*~ macrophyllum purshStreambanks.

*~ nequndo L. var. californicum (T. & G.) SargeStreambanks.

46. Hippocastanaceae

Aesculus californica (Spach) Nutt.

Oak-madrone forest, especially along streambanks.

-17-

Page 18: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

ssp. rubicunda

This is probably the species which Sprin

arnoena (Lehm.) Lilja.

47. Rhamnaceae

*Ceanothus cuneatus (Hook.) Nutt. var. dubius Howell

Chaparral.*Ceanothus sorediatus H. & A.

Chaparral, occasionally in oak-madrone forest.

Ceanothus th~siflorus Esch.

Chaparral •.

Rhamnus californica Esch. ssp. californicaOak-woodland and oak-madrone forest.

*Rhamnus crocea Nutt. ssp. croceaOak-madrone forest.

48. Malvaceae

)CMalacothamnus arcuatus (Greene) Greene

Chaparral.

Malva parviflora L.Disturbed areas.

Sidalcea ma1vaeflora (DC.) Gray ex Benth.Grasslands.

49. Cistaceae

*Helianthemum scoparium Nutt. var. vulqare Jeps.

Chaparral.

50. Vio1aceae

*Viola ocellata T. & G.

Dudlev (Apr. 1, 1893) was collected from "Hill above Searsville."

51. Datiscaceae

*Datisca 910merata (Presl) Bai11.

San Francisquito Creek.

52. Thyme1aeaceae

*Dirca occidentalis GrayOak-madrone forest.

53. Lythraceae

*L~thrum hvsso~ifolia L.Searsville Lake.

54. Onagraceae

Boisduvalia densiflora (Lindl.) Wats.

Reported by Moeur (1947) from "moist ~round.around the lake."

Clarkia purpur~a (Curtis) Nels. & Macbr. ssp. quadrivulnera (Dougl.) Lewis & Lewis

Grasslands and openly wooded hillsides.

*Clarkia purpurea (Curtis) Nels. & Macbr. ssp. viminea (Dougl.) Lewis & Lewis

Grasslands and openly wooded hillsides.

Clarkia rubicunda (Lindl.) Lewis and Lewis

Grasslands and openly wooded hillsides.

(1935) and Moeur (1947) list as Godetia

-18-

Page 19: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Clarkia unquicu1ata Lindl.

Oak-woodland.

*Epi10bium adenocau10n Hausskn. var. occidentale Tre1.Searsville Lake.

gpilobium panicu1atum Nutt.

Reported by both Springer (1935) and Moeur (1947) to occur along roadsides through

the chaparral.

~pilobium franciscanum Barbey

Reported by both Springer (1935) and Moeur (1947) to occur in the moist area nearSearsville Lake.

*Oenothera ovata Nutt.

Grasslands.

55. Umbe 11iferae

y Anthriscus 8candicina (Weber) Mansf.

~ Oak-madrone forest.Caucalis microcarpa H. & A.

)( Oak-woodland.Conium maculatum L.

Streambanks.

*Ervnqium aristu1atum Jeps.

Moist grasslands.*Foeniculum vulqare Mill.

Disturbed areas.

y "erae leum maximum Bartr.

Streambanks.

Lomatium dasvcarpum (T. & G.) C. & R.Grasslands.

~ !-~matium macroearpum (H. & A.) C. & R.

Reported by Moeur (1947) from "open hillsides."

Lomatium utriculatum (Nutt.) C. & R.

Grasslands and openly wooded slopes.*Oenanthe sarmentosa Presl •.

Searsville Lake.

X Osmorhiza chilensis H. & A.

Reportedby Moeur (1947) from "shady woods near San Francisquito Creek."Perideridia qairdneri (H. & A.) Mathias

Reported by Moeur (1947) from "both open and openly wooded hillsides" (as ~

qairdneri Gray).

Eerideridia kelloqqii (Gray) Mathias

Reported by Springer (1935) from "dry open hillsides" (as ~ kelloqqii Gray).

Sanicula bipinnatifida Dougl. ex Hook.Grasslands.

*Sanicula crassicaulis Poepp. ex DC.Oak-madrone forest.

Sanicula laciniata H. & A.

Reported by Springer (1935) from "edges of a road cut through the chaparral."

Tauschia kelloqqii (Gray) Macbr.

Reported by Springer (1935) from "an openly wooded slope~" as Velaea kelloQ9ii

(Gray) C. & R.

Torilis nodosa (L.) Gaertn.

Reported by Moeur (1947) from "openly wooded hillsides."

56. Cornaeeae

)Ccorn~s ~alifornica C. A. MeySan Franeisquito Creek.

-19-

Page 20: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

eornus qlabrata Benth.Streambanks.

57. Garryaceae

*Garrva elliptica Dougl •

. Chaparra 1.

58. Ericaceae

*Arbutus menziesii Pursh

Dominant with Quercus agrifolia in the oak-madrone forest.

*Arctostaphvlos canescens Eastw.

fobrams 4990, collected on "Jasper Ridge, It bears the notation, "A. canescensEastwood? Poor specimen. A. E. " ,

*Arctostaphvlos crustacea Eastw. var. crustacea

Chaparral.

59. Primulaceae

*Anaqallis arvensis L.

Grasslands, oak-woodland, and disturbed areas in the chaparral.*Centriculus minimus L.

Moist grassland$.

*Dodecatheon clevelandii Greene ssp. patulum (Greene) ThompsonGrasslands and openly wooded slopes.

*Dodecatheon hendersonii Gray ssp. cruciatum (Greene) Thompson

Grasslands and openly wooded slopes.*Trientalis latifolia Hook.

Oak-madrone forest.

60. Gentianaceae

~ ~entaurium Qavvi (Jeps.) AbramsReported by Springer (1935) from "openly wooded slopes."

*Microcala Quadranqularis (Lam.) Griseb.Grasslands.

61. Apocynaceae

Yinca major L.

Reported by Moeur (1947) from "bank of San Francisquito Creek."

62. Convolvulaceae

Convolvulus arvensis L.

Disturbed areas.

Convolvulus occidentalis Gray

Chaparral.

')< 9m.Y9lvulus subacaulis (H. & A.) GreeneReported by Moeur (1947) from "meadows and pasture lands."

63. POlemoniaceae

~Collomia heterophylla Hook.

Reported by Moeur (1947) from "shaded areas of the wooded slopes."

*Eriastrum abr?msii (Elmer) Mason

Chaparral.

-20-

Page 21: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*Gi11a achilleaefolia Benth.

Grasslands, oak-woodland, and oak-madrone forest.

*Gi11a clivorum (Jeps.) GrantGrasslands.

Linanthus ambiquus (Rattan) Greene

Reported by Moeur (1947) from "open hillsides."Lin~nthus androsaceus (Benth.) Greene

Grasslands and openly wood~d slopes.

*Linanthus bicolor (Nutt.) Greene

Grasslands and openly wooded slopes.*Linanthus dichotomus Benth.

Grasslands.

*Linanthus liniflJ2J'_l1.S(B.enth.) GreeneGrasslands.

*Linanthus parviflorus (Benth.) Greene

Grasslands and openly wooded slopes.

*Navarretia heterodoxa (Greene) Greene

Chaparral.*Navarretia viscidula Benth.

X Grasslands.Pholox qracilis (HOOk.) Greene

Grasslands and openly wooded slopes.

64. Hydrophyllaceae

*Eriodictyon californicum (H. & A.) Torr.Chaparral.

*Nemophila heterophylla F. & M.Oak-madrone forest.

*Nemoqhila menziesii H. & A. var. menziesiiGrasslands.

Nemophila menziesii H. & A. var. atomaria (F. & M.) ChandlerGrasslands.

Phacelia californica Cham.

Reported by Moeur (1947) from "rocky hillsides."Phacelia imbricata Greene

- Reported by Springer (1935) from "dry rocky hillsides" (as~. californica var.imbricata Jepson).

~~~l~~~~~?auritum (Lindl.) Lilja ex Lindb.Oak-madrone forest.

65. Boraginaceae

Allocar~ chorisiana (Cham.) Greene

Reported by Moeur (1947) from "open fields."~Amsinckia inter~Edia F. & M.

Grasslands.

*Cynoqlossum qrande Dougl. ex Lehm.Oak-madrone forest.

*Cryptantha flaccida (Dougl.) Greene

Serpentine soil in the meadow.

Cryptantha leiocarpa (F. & M.) Greene

Reported by Springer (1935) from lithe more open places in the chaparral."*gryptantha micromeres (Gray) Greene

Dry, rocky grasslands.

Cryptantha torreyana (Gray) Greene var. pumila (Heller) Johnston

Reported by Moeur (1947) from "dry hillside."

-21-

Page 22: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

}(Heliotrop1um curassavicurn L. var. oculatum (Heller) Johnston ex Tidestrom

Reported by Springer (1935) from "along the side of the road very close to thelake. "

*Pectocarve pus1lla (A. DC.) Gray

"Chaparral, open woods, and on serpentine •••• " (Thomas, 1961, p. 290).

Pla~iobothrvs nothofulvus (Gray) GrayGrasslands.

*Plagiobothrvs tenellus (Nutt.) GrayGrasslands.

66. Verbenaceae

Verbena lasiostachvs Link.Disturbed areas.

67. Labiatae

*Lepechiniq calycine (Benth.) Epling

Chaparral.)r Melissa officinalis L.

Reported by Moeur (1947) from "creek banks."

Mentha {lrvensis L. var. lanatEj Piper

Reported by Springer (1935) as "Frequent in moist places - near the overflow from

the watering trough, and at the edge of the lake."*Mentha pule~ium L.

Searsville Lake and San Francisquito Creek.*Monardella villose Benth. var. villose

Grasslands and openly wooded slopes.

*Po~o~yne serovlloides (Torr.) GrayGrasslands.

Prunella vulgaris L. ssp. lanceole~~ (Barton) HultenReported by Moeur (1947) from "banks of San Francisco Creek" (as ~z:unelle vulgaris L

*Salvia columbarlli Benth.Grasslands.

Satureja dou~lasii (Benth.) Briq.Oak-madrone forest.

*Scutellaria tuberosa Benth.

Chaparral.Stechvs bullate Benth.

~ Openly wooded slopes, oak-woodland, and oak-madrone forest.Stachls pvcnantha Benth.

Reported by Springer (1935) from "edge of the lake."

68. Solanaceae

*Petunia parviflora Juss.Searsville Lake.

Solanum nodiflorum Jacq.

Reported by Moeur (1947) from "the dry bed of San Francisquito Creek." (as 2,.

)( ni~rum L.).polanum umbelliferum Esch.

Oak-madrone forest, less common in the chaparral.

69. Scroph~ariaceae

Antirrhinum vexlllo-calyculatlJn1 Kell.

Reported by Springer (1935) from "edge of the shady area at the margin of the

lake." (as A. vagans Gray).*Castilleia affinis H. & A.

Oak-woodland.

-22-

Page 23: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Castilleia foliolosa H. & A.

Reported by Springer (1935) from "dry hillsides"and "the edge of the chaparral."

Collinsia heterophvlla Buist ex Grah.Oak-madrone forest and oak-woodland.

Collinsia sparsiflora F. & M. var. collina (Jeps.) Newsom

Reported by Springer (1935) from "steep, moist, open hillside above San Franc is­

quito Creek" (as ~. §£arsiflora F. & M.)

Cordvlanthus pilosus Gray

Grasslands and openly wooded slopes.

*Diplacus aurantiacus (Curtis) Jeps.

Chaparral.*Linaria texana Scheele

Grasslands.

*Mimulus douqlasii (Benth.) Gray

Serpentine soil in the meadow.

*Mimulus quttatus DC.Streambanks.

*Orthocarpus attenuatus GrayGrasslands.

*Qrthocarpus densiflorus Benth.Grasslands.

*Orthocaxpus erianthus Benth. var. erianthusGrasslands.

Orthocarpus faucibarbatus Gray var. albidus (Keck) Howell

Reported by both Springer (1935) and Moeur (1947) from the serpentine soil of the

meadow (as Q. faucibarbatus Gray).

*Orthocarpus lithospermoides Benth.

Serpentine soil of the meadow.

Orthocarpus purpurascens Benth.var. purpurascensGrasslands.

~Orthocarpus pusillus Benth.

Openly wooded hillsides.*Pedicularis densiflora Benth. ex Hook.

Oak-woodland.

*Scrophularia califo~ c. & S.

Oak-madrone forest, occasional in chaparral.

70. Orobanchaceae

*Orobanche bulbosa Beck

Chaparral.Orobanche fasciculata Nutt.

Reported by Springer (1935) from "dry open hillside."

71. Plantaginaceae

*Plantaqo erecta MorrisGrasslands.

*Plantaqo lanceolata L.Disturbed areas.

*Plantaqo major L.Reported by Moeur (1947) from "near the lake,"

72. Rubiaceae

Galium aQarine L.Grasslands, oak-woodland, and oak-madrone forest.

-23-

Page 24: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

Galium californicum H. & A.

Reported~by Cooper (1922) from chaparral and oak-madrone forest.

*Galium murale (L.) All.

~ Grasslands, chap~rral, and oak-woodland.Galium nuttallii Gray

Chaparral.

Galium trifidum L. var. subbiflorum Wieg.

Reported by both Springer (1935) and Moeur (1947) from "shady woods."Galium trif10rum Michx.

Reported by Moeur (1947) from '~oody thickets and openly forested slopes."*Sherardia arvensis L.

Thomas 4996 was collected at "Jasper Ridge. Small draw on south side of ridge."

73. Caprifoliaceae

)<Lonicera hispidu1a Dougl. ex Lindl.Oak-madrone forest.

~onicera involucrata (Richards) Banks ex S~rengStreambanks.

~Sambucus mexicana Presl ex DC.

Chaparral.

XSymphoricarpos albus (L.) Blake var. laevigatus (Fern.) BlakeChaparral, oak-woodland, and oak-madrone forest.

74. Dipsacaceae

~Dipsacus fullonum L.

Disturbed areas.

75. Valerianaceae

Plectritis ciliosa (Greene) Jeps. ssp insiqnis (Suksd.) MoreyGrasslands.

Plectritis macrocera T. & G.

Grasslands and openly wooded slopes.

76. Cucurbitaceae

x .M~ fabaceus (Naud.) Greene

Chaparral, oak-woodland, and oak-madrone forest.

77. Campanulaceae

*Triodanus biflora (R. & P.) Greene

Chaparral.

78. Compositae

Cichorieae

YAqOseris qrandiflora (Nutt.) Greene

~ Grasslands and openly wooded slopes.Hieracium albiflorum Hook.

~ Oak-madrone forest.Hypocha~ qlabra L.

Grasslands and openly wooded slopes.

-24-

Page 25: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*Lactuca saliqna L.

Thomas 8074 was found "Growing in serpentine rock crevice."

*Micxoseris douqlasii (DC.) Sch.-Bip. ssp. douqlasii

Grasslands and openly wooded slopes.

Microseris heterocarpa (Nutt.) Chambers

Reported by Moeur (1947) as "in fields and on open hillsides" (as Vropappus

~ lindleyi var. leucocarpus Jepson.)Picris echioidesL.

Disturbed areas.

~F2finesquia californica Nutt.

Reported by Springer (1935) from "openly wooded slopes."

Sonchus asper (L.) Hill

y Disturbed areas in moist grasslands and openly wooded slopes.f'Sonchus oleraceus L.

Disturbed areas.

Stephanomeria vir9ata Benth.Disturbed areas.

Taraxacum officinale Weber

Disturbed areas.

Astereae

Aster chilensis Nees

Oak-woodland and openly wooded slopes.

*A2.ill.radulinus Gray

Chaparral, occasionally on openly wooded slopes.

*Baccharis douqlasii DC.Streambanks.

*Baccharis pilularis DC. var. consanquinea (DC.) Kuntze

Chaparral and openly wooded slopes.

*Chaetopappa alsinoides (Greene) Keck

Grasslands and openly wooded slopes.~rigeron foliosus Nutt.

Grasslands.

g~iqeron philadelphicus L.

Reported by Springer (1935) from "open hillside near the bank of a drying stream."

*Grindelia camporum GreeneGrasslands.

Grindelia hirsutula H. & A.

Grasslands and openly wooded slopes.

*Lessinqia qermanorum Cham. var. tenuipes HowellGrasslands.

Lessinqia holo~ Greene vax. hololeucaGrasslands.

*SolidaQo ca1ifornica Nutt.

Openly wooded slopes.

rSOlidaQo occidentalis (nutt.) T. & G.Searsv ilIe Lake.

lnuleae

*Adenocaulon bicolor HookOak-madrone forest.

Anaphalis marqaritacea (L.) Gray

Reported by both Springer (1935) and Moeur (1947) from both open and openlywooded hillsides.

-25-

Page 26: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

*~ sparsiflora (Gray) Jeps.Grasslands.

*Gnaphalium californicum DC.

"Fairly cornmon in poor soils, in chaparral, and on dry ridges •••• " (Thomsa, 1961,p. 355).

Gnaphalium chilense Spreng.

)C Reported by Moeur (1947) from "dry, open hillside."Gnaphalium luteo-album L.

Reported by Moeur (1947) from "along the side of the road above the lake."

*Gnaphalium palustre Nutt.Searsville Lake.

Gnaphalium purpureum L.Grasslands.

~Micropus californicus F. & M.

~ Grasslands and openly wooded slopes.Psilocarphus tenellus Nutt.

Grasslands and openly wooded slopes.

Heliantheae

*Helianthella californica Gray

Grasslands and openly wooded slopes.

*Wyethia anQustifolia (DC.) Nutt.Grasslands.

Madieae

*Achyrachaena mollis SchauerGrasslands.

*Calycadenia multiqlandulosa DC. ssp. robusta Keck

Serpentine soil in the meadow.

*Hemizoni~ corymbosa (DC.) T. & G.Grasslands.

*Laqophylla ramosissima Nutt.Grasslands.

*Laxia hieraciodes (DC.) H. & A.Chaparral.

~LaYia platyqlossa (F. & M.) Gray ssp. campestris KeckGrasslands.

*Madia exiqua (Smith) GrayOak-woodland.

*Madia qracilis (Smith) Keck

Grasslands and openly wooded slopes.

Madia madioides (Nutt.) Greene

Reported by Moeur (1947) from "heavily wooded slopes.lIMadia sativa Molina

Disturbed areas.

Ambrosieae

>'Xanth ium strumar ium L.Searsville Lake.

Helenieae

Baeria chrysostoma F. & M. ssp. gracilis (DC.) FerrisGrasslands.

-26-

Page 27: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

)CEriophyllum confertiflorum (ne.) Gray

Chaparral.

*Eriophyllum lanatum (pursh) Forbes var. arachnoideum (Fisch. & Ave-Lall.) Jeps.Oak-woodland.

XHelenium puberulum DC.Streambanks.

Monolopia qracilens Gray

Reported by Springer (1935) from "near the edge of the chaparral above the lake,"

and by Moeur (1947) from "openly wooded hill-slopes."

Antl,1emideae

Achillea millefolium L. var. californica (Pollard) Jeps.Oak-woodland and oak-madrone forest.

YAnthemis cotula L.

Grasslands and disturbed areas.

*Artemisiabiennis Willd.

Searsville Lake.

*Artemisia californica Less.

Chaparral, often forming thick stands in which it is the dominant shrub.

Artemisia douqlasiana Bess.Streambanks.

Matricaria matricariodes (Less.) PorterDisturbed areas.

Senec ioneae

*Arnica discoidea Benth.

Mason (May 18, 1921) was collected "Near redwood area, Jasper Ridge."*Senecio aronicoides DC.

Chaparral and openly wooded slopes.

~enecio breweri DavyReported by Moeur (1947) from "openly wooded areas."

*Senecio mikanioides Otto

Streambanks.

~Senecio vulqari~ L.Disturbed areas.

Cynareae

*Carduus ~iflorus CurtisDisturbed areas.

Centaurea calcitrapa L.Disturbed areas.

~ Centaurea me Iitens is L.

Disturbed areas in grasslands and along the roads.Centaurea sol~titialis L.

Disturbed areas.

Cirs~ brevistvlum Cronq.

Reported by Springer from "a place kept moist by the overflow of a watering

trough." (as g. edule Nutt.).Cirsium occidentale (Nutt.) Jeps.

X Reported by Maeur (1947) from "dry, openly wooded slopes."Cirsium proteanum Howell

Reported by Springer (1935) from "dry, more openly wooded slopes." (as g.

occidental~ var. venustum Jepson).

-27-

Page 28: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

K~irsium vulQar~ (Savi) TenoreDisturbed areas.

Si)ybu~ m~riAnum (t.) Gaertn.Disturbed areas.

As you can see, this list of the vascular plants of Jasper Ridge is

not complete. Nor are there specimens in the Dudley Herbarium represen-:jLng

each of the t,.:lxalisted. Theya~e many reasons for t his, mainly historical.

For some years I have been slouly completing the list and getting specima,s.

A separate herbarium case housed the plants from Jasper Ridge. This is available

for use, but only after receiving permission to do so. If you come across

plants that you do not think are in the list,·please communicate this to me.

We can then get a good sample for a permanent herbarium specimen.

I ~nllbe glad to help with identification of specimens, but there is a

limi t to my time. If yon Hant help 1·r'.L th identification, l'lease be sure that

you have enough plant material so that :-t can be identified. Sterile grassblades - ~o thankyouo

John H. Thomas

Harch 26, 1971

-28-

Page 29: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

ADDITIONS TO THE FLORA OF JASPER RIDGE

2. Polypodiaceae

*Drvopteris ar~uta (Kaulf.) lJatt.

PJDOng redwoods

*Polvpodium californicum Kaulf.Rock crevices and on rocks among mosses in shade.

3. potamogetonaceae

*Potamo~eton illinoensis MorongEdge of Searsville Lake near bridge.

4. Alismataceae

*Echinodorus berteroi (Spreng.) Fassett

Dry lake bed at Searsville Dam.

5. Gramineae

*Festuca eastwoodae PiperOpen oak woodlands.

*.f2! howellii V. & S.Between Searsville Dam and redwood area.

*Scribneria bolanderi (Thurb.) Hack.Chaparral.

*Calama2rostis koelerioides VaseyOak woodland.

*Stipa lepida Hitchc.Grasslands.

*Phalaris tuberosa L. var. stenoptera (Hack.) Hitchc.

Along San Francisquito Creek below Searsville Dam.

7 • Juncaceae

*Juncus bufonius L.

Grassy meadows in oak woodland.

11. Amaryllldaceae

*Brodiaea con~esta Sm.Oak woodland.

13 • Orchidaceae

*Corallorhiza maculata Raf.Without specific locality.

19. Urticaceae

*Urtica urens L.

In shade at base of sandstone outcroppings below Searsville Dam.

Page 30: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

2.

22. Chenopodiaeeae

*~opodium !E!?!'.2.:~~ L. var. ambrosioidesBelow Searsville Dam.

22a. Amaranthaceae

*Amaranthus Albus L.

Dry lake bed near Searsville Dam.

24. Caryophyllaceae

*Sa~ina apetala Ard. var. barbata Fenzl ex Ledeb.Edge of road in oak woodland.

25. Ranunculaeeae

*Ranuneulus hebecarpus H. & A.

Moist, shaded slopes near Searsville Lake.

29. Cruciferae

*Brassiea aenieulata (Desf.) BallDisturbed, grassy slope above Searsville Lake.

37.• Fabaceae

*L!thyrus !atifolius L.Along San Francisquito Creek below Searsville Dam.

*Lupinus latifolius Agardh var. latifoliusOak woodlands.

*Trifolium amplectens T. & G.Open serpentine grasslands.

*Trifolium microdon H. & A.Oak woodlands.

54. Onagraeeae

*Boisduvalia stricta (Gray) Greene

Along road in oak woodland.

*EpUobium minutum Lind!. ex Hook.Chaparral.

54a. Haloragidaeeae

*Mvriophvllum brasilienee Camb.Mud at edge of Searsville Lake near bridge.

55. Umbel1iferae

*Scandix pectin-veneris L.Oak woodlands.

Page 31: STANFORD UNIVERSITY JASPER RIDGE BIOLOGICAL …trees.stanford.edu/PDF/porter_rev.pdf · Cirsium vul~are SilybuJIl marianum. The aquatic and simiaquatic plants in and around Searsville

3.

6la. Asclepiadaceae

*Asclepias fascicularis Decne.

Edge of grassland bordering chaparral at Westridge end of Ridge.

64. B;ydrophyllaceae

*Phacelia rattanii Gray

Edge of chaparral.

65. Boraginaceae

*Mvosotis latifolia Poir.Below Searsville Dam.

69. Scrophulariaceae

*Kickxia spuria (L.) Dumort.

Along San Francisquito Creek near Searsville Dam.

*Vero.nica persica Poir.Grassy slope near Searsville Dam.

78. Compositae

*Crepis vesicaria L. ssp. taraxacifolia (Thuill.) TheIl.Oak 't<loodland.

*Convza canadensis (L.) Cronq.Edge of road above Searsville Lake.

*G~aphalium beneolens Dav.

Edge of lake at Searsville Dam.

*Rigiopappus leptocladus GrayChaparral.

*~~ethia glabra GrayOak woodland.

Compiled by John H. Thomas" April 26. 1965