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Executive Summary of the�� Stock Assessment Workshop

��th SAW� for theNew Jersey Delaware Bay Oyster Beds

Presenters

Eric Powell� Haskin Shell�sh Research LaboratoryJohn Kraeuter� Haskin Shell�sh Research LaboratoryDavid Bushek� Haskin Shell�sh Research Laboratory

Stock Assessment Review Committee

Russell Babb� New Jersey Department of Environmental ProtectionBarney Hollinger� Delaware Bay Section of the Shell Fisheries Council

Roger Mann� Virginia Institute of Marine ScienceSteve Fleetwood� Delaware Bay Section of the Shell Fisheries Council

Rich Wong� Delaware Department of Natural ResourcesMike Celestino� New Jersey Department of Environmental Protection

Joe Dobarro� Rutgers UniversityThomas Landry� Department of Fisheries and Oceans� Canada

Steve Jordan� U�S� Environmental Protection Agency

Editors�

Eric Powell� Haskin Shell�sh Research LaboratoryJohn Kraeuter� Haskin Shell�sh Research Laboratory

Kathryn Ashton�Alcox� Haskin Shell�sh Research Laboratory

Distribution List

Barney Hollinger� Chair� Delaware Bay Section of the Shell Fisheries CouncilJim Joseph� New Jersey Department of Environmental ProtectionSelected faculty and sta�� Haskin Shell�sh Research Laboratory

Oyster Industry Science Steering CommitteeStock Assessment Review Committee

February ��

Haskin Shell�sh Research Laboratory �� Rutgers� The State University of New JerseyHaskin Shell�sh Research Laboratory �� Rutgers� The State University of New JerseyHaskin Shell�sh Research Laboratory �� Rutgers� The State University of New Jersey

Status of the Stock

Figure � summarizes the condition of the oyster stock throughout the NewJersey waters of Delaware Bay at the end of �� The stock presents a mixtureof positive and negative indicators that approximately balance� Abundance is low�but abundance increased in three of four bay regions� Abundance continued tobe below target levels in all bay regions Figures � and �� but above thresholdlevels on the medium mortality beds and Shell Rock and near� but below� thresholdlevels on the high mortality beds� The shell planting program promises to increaseabundance on these downbay beds in �� Abundance has increased each year onthe high mortality beds since reaching a post �� low in �� Figures � and ��and abundance has moved in a positive direction for several years on Shell Rock�The stock continues to be disproportionately consolidated on the medium mortalitybeds� a process that began in the early ��s with persistent recruitment failure andthe in�uence of Dermo disease downbay�

Spawning stock biomass rose slightly in �� Increases were noted in all bayregions except upbay on the low mortality beds� SSB has increased steadily on thehigh mortality and medium mortality beds over the last three years and has risenfor the last two years on Shell Rock Figures � and �� SSB was above the biomasstarget in three of � bay regions and near the threshold for the low mortality beds�

Recruitment remains low bay wide Figure �� and was particularly low on thelow mortality beds� high mortality beds� and Shell Rock in �� A near averagerecruitment event occurred on the medium mortality beds� However� in no case didrecruitment reach the value of �� spat per adult� The ratio of spat to oysters hasbeen lower than �� over six of the last � years� Shell planting increased recruitmenton Shell Rock and the high mortality beds and this increase brought the spat per adult ratio above �� on Shell Rock and up to �� on the high mortality beds� Bothratios are normally associated with increasing abundance in the following year�Evidence exists that low spat abundance is associated with low adult abundance�and suggests that the explanation involves the contribution of live oyster shell tothe cultch resource preferred for settlement� This implies that high recruitmentmay be less likely under current conditions of low abundance�

The oyster population as a whole continues to be depauperate in the smallersize classes Figure �� with the proportion of animals �� being below the

��th percentile in all bay regions and no higher than the ��th percentile in three�If this trend continues� a decline in abundance and SSB is highly likely to occurover the coming few years and this will restrict �shery yield� In contrast� surplusproduction projected for �� is expected to permit an increase in market sizeabundance bay wide and in all bay regions� given average mortality rates� barringa higher than average rate of natural mortality� and not counting removals by the

�

�shery� This continues the trend of positive surplus production in most bay regionsobserved over the last few years�

Dermo disease rose to moderate levels in �� and natural mortality rates weresomewhat above average Figure �� A rising trend in Dermo disease weightedprevalence may presage epizootic rates of natural mortality in ��

Fishery exploitation levels since �� have been low �� of abundance per

year�� Recent improvements in collection of �shery dependent data indicate thatexploitation in terms of biomass has been �� over most of that time� Lowexploitation rates indicate that the �shery does not have a signi�cant e�ect onthe stock and that �shing mortality is not responsible for the current conditions oflow abundance�

The beds continued to su�er a net loss of shell in �� However� the shell planting program reduced this net loss to the lowest value during the �� time period when data are su�cient to estimate the trend Figure ��

Overall� the conditions on the low mortality beds are distinctly disadvanta geous� whereas the remaining bay regions appear to have improved since ��However� the fact that all bay regions fall below their abundance targets indicatesthat actions to enhance abundance are needed in all bay regions� A reduction in�shing e�ort will not signi�cantly address this need because exploitation rates arealready low� however� substantial increases in exploitation rate should be avoidedas the importance of adults as sites for larval settlement and the continued need tominimize shell loss reinforces the importance of maintaining biomass at or abovetarget levels� Abundance has been enhanced on the high mortality beds and ShellRock by downbay transplant and this program should be continued� The preferredmechanism to address low abundance overall is to enhance recruitment throughshell planting�

�� Management Goals

Cultch Management Goals

Continued shell planting is essential to maintain habitat quality as well asprovide substrate to enhance recruitment� Shell plants should target areas wheremarketable oysters grow but where cultch loss exceeds the addition of shell throughnatural mortality� The Ship John region is such a case� Due to the enhancedsurvival of juveniles in this region� replants from downbay plants should be movedto selected areas of Ship John in �� The same region� and Cohansey as well�should be considered for direct shell plants� In addition� Nantuxent Point� Hawk�sNest� Hog Shoal� Vexton� and Strawberry should be considered as planting localesfor direct shell plants� Downbay plants and replants should be expanded to the

�

extent funds permit to enhance recruitment� Shell Rock and Bennies Sand shouldnot be receive shell plants in ��

Bay Region Considerations�Shell Rock Downbay

Shell Rock and the high mortality beds have provided most of the �shedanimals since �� because market quality is consistently high� These beds havebeen successfully managed using a constant abundance reference point since ��with a precautionary component to guard against epizootic losses� Harvest levelsderived from exploitation based reference points that vary considerably from therange suggested by surplus production projections should be considered non

precautionary� This year� exploitation based reference points using the ��th to

�th percentiles fall within the range established by surplus production calculations

using the ��th to ��th percentile of natural mortality rate and the average of the�� and �� updated growth rates� Thus the two approaches provide relativelyequivalent allowable harvest levels�

Due to the uniqueness of medium mortality and high production� and given itsimportance to the �shery� Shell Rock must be managed independently of the high mortality beds and under more conservative guidelines� For surplus production

projections� the ��th percentile assumption should be retained�

Bay Region Considerations�Low�mortality Beds

This bay region is below threshold abundance levels and at threshold SSBlevels� The SARC recommends that this bay region be closed in ��

Bay Region Considerations�Medium�mortality Beds

These beds have contributed the bulk of the stock supporting the �sheryover the entire �� year history of the survey� excepting the ��s high abundanceperiod� SAW � recommended that management should emphasize increased directmarketing on these beds to reduce the exploitation rate downbay while stockrebuilding continues� Biomass continues to be high on these beds and continuedfocus on directing e�ort to these beds is important� The SARC recommendsthat the upper bed group Middle and Upper Middle� be used for intermediatetransplant and encourages that intermediate transplant to Shell Rock or BenniesSand be a component of the �� program� The SARC recommends that Sea Breeze�Cohansey� and Ship John be managed as direct market beds in �� as in ��

The high levels of surplus production anticipated for �� should not permitdramatically expanded exploitation of these beds� The SARC strongly recommendscontinued use of the exploitation reference points on these beds� The SARC notesthat the high surplus production expected in �� originates from the growth of

�

the last large cohort� recruited in the early ��s� into market size and that thesebeds do not have substantial numbers of smaller animals supporting continuedstock expansion in the future� Given the likely average age of animals growinginto market size � to years�� management of these beds should consider that a�� recruitment event will not bene�t the �shery for minimally � years� Thus�the surplus production anticipated in �� should be viewed as minimally a � year allotment� The �ve year time period has implications for the evaluation ofexploitation reference points in the context of minimizing yearly variations in theallocation� A yearly harvest level above �� to �� bushels risks a reductionin allocation from this bed region in future years that may be long term in duration�even with a substantive recruitment event in �� The SARC strongly recommendsthat exploitation rate percentiles permitting harvest above this level be viewed asinhibitory to the long term health of the �shery�

Allocation Projections � Constant�abundance Reference Point

The surplus production option assumes direct marketing� Consequently� esti mates are provided only for the high mortality beds� Shell Rock� and the lower groupof medium mortality beds Cohansey� Ship John� Sea Breeze�� The SARC notesthat surplus production estimates for these beds covered a wide range because ofuncertainty in growth rates and strongly recommends use of the exploitation basedreference points rather than the constant abundance reference point for all bed re gions except Shell Rock� Accordingly� the SARC recommends that exploitation ratereference points be used for allocation decisions for the high mortality beds in ��with a harvest value chosen that does not stray substantively from the range delin eated in the following table� so that rebuilding of abundance can be achieved during

times of shell planting� The SARC recommends that the ��th percentile constant abundance projection be used for allocation decisions for Shell Rock� A range ofoptions exists this year for the medium mortality beds� The SARC recommendsthat exploitation reference points be used for these beds with allocation estimatesin accordance with a � year distribution of �� surplus production as shown in thefollowing table�

Natural AllocationBay Region Mortality Percentile market equivalent bushels�

High mortality ��th ��th �� q

Shell Rock ��th ��

Lower medium mortality ��th �� to ��

Upper medium mortality NAxLow mortality NAx

�

xNA� not applicable to this reference point�qThe range given provides guidance for the implementation of an allocation plan using theexploitation reference points in the next table��The range given provides guidance for the implementation of an allocation plan using theexploitation reference points in the next table� The range is based on a ��year distribution of�� surplus production�

Allocation Projections � Exploitation Reference Point � Direct Marketing

Projections are provided for the high mortality beds� Shell Rock� and thelower group of medium mortality beds Cohansey� Ship John� Sea Breeze�� TheSARC recommends that the high mortality and medium mortality beds be managedusing this reference point in �� cognizant of the restrictions imposed by surplusproduction on the high mortality beds and the likely reduction in SSB over thenext �ve years on the medium mortality beds� The SARC notes than an upper

and lower bound for this reference point normally should be taken as the ��th and

�th percentiles of the �� time series� however� the SARC recognizes thatlower or higher percentiles may be investigated after consideration of surplusproduction projections and the position of abundance and SSB relative to thresholdand target levels� This year� these bay regions are above their SSB targets� but belowtheir abundance targets and� below the abundance threshold for the high mortalitybed group� thus stock rebuilding should be included in the management goal�

Numberof

Exploitation Animals Direct marketBay Region Percentile Rate Removed Bushels

High mortality ��th ��

��th ��

�th ��

��th ��


��th ��

�th ��

Lower medium mortality ��th ��

��th ��

�th ��

Upper medium mortality NAxLow mortality NAx

xNA� not applicable to this reference point�

�The inclusion of this percentile recommendation was not uniformly supported by the SARC�The SARC notes the following� �� This value falls within the range of values estimated usingthe constant�abundance approach� �� The high�mortality beds are above the biomass target�however a higher than anticipated mortality event is anticipated to occur on these beds in �� The high�mortality beds are below the threshold in abundance requiring the rebuilding ofabundance� however shell planting has targeted this area and initial estimates show a promisingincrease in abundance that might bring abundance above the threshold in �� The SARC wasabout evenly divided on the issue of whether the concern about rebuilding abundance shouldoutweigh the high�biomass level present considering that the volatility of the resource in this bayregion caused by the wide range of mortalites from Dermo disease may limit the long�term successof stock rebuilding programs� The SARC has chosen to include this option with the caveat that itis unlikely to be a precautionary option and with the further caveat that its inclusion should notbe considered precedent setting for application to management advise by future SARCs��The SARC notes that this value approximates the value obtained using the constant�abundancemethod with a ��th percentile natural mortality rate and average growth� It is therefore a pre�cautionary option given the prediction that natural mortality rates will exceed the ��th percentilein ��The SARC notes that this value is consistent with the ��th percentile surplus productionrecommendation�

Allocation Projections � Exploitation Reference Point � Intermediate Transplant

The estimates assume that intermediate transplant will be conducted on theupper medium mortality beds Middle� Upper Middle� and that direct marketingwill be conducted on beds downbay of these two beds� The estimated number ofbushels to be moved is derived from the mean of the number of oysters per bushelobtained from the �� survey� If cullers are used� the number of bushels can bereduced by an estimated factor of �� The proportion of animals available formarket is estimated based on the fraction of animals �� Note that allocationvalues are estimates and can only be speci�ed after the transplant is complete�Moreover� if cullers are used� the marketable bushel estimate may be low�

Number Deck loadof Deck load Marketable

Exploitation Animals Population Transplant BushelBay Region Percentile Rate Removed Oysters�Bu Bushels EquivalentsHigh mortality NAxShell Rock NAxLower medium mortality NAx

Upper medium mortality ��th ��

��th ��

�th ��

Low mortality CLOSED�

xNA� not applicable to this reference point��The SARC recommends closure of the low�mortality beds in ��

�

Caveats Apropos to Risk for �� Fishery Yield

Management options for �� provided in the preceding tables permit arelatively wide range of possible �� shery allocations� Mortality in �� wasabove that in �� but not yet at epizootic levels� Dermo levels are increasing�however� and Dermo induced mortality is likely to increase unless environmentalconditions inhibit further development� The corollary is the natural mortality ratesare expected to be above average in �� Increasing Dermo disease and continueduncertainty in recruitment suggests that moderation in setting �� allocation goalsmay be desirable� Speci�c observations include the following�

�� The SARC recognizes the need to manage Shell Rock and the high mortalitybeds conservatively� Present day abundance on the high mortality beds is be low threshold levels Figure �� requiring that rebuilding be included as a man agement goal� These beds have responded positively to abundance enhance ment programs by shell planting to increase recruitment� and transplantingto increase adult abundance� Retention of these mechanisms within yearlymanagement plans is essential while abundance is low�

�� The SARC notes that the low mortality beds will be re surveyed in �� andthat the updated abundance and biomass estimates thus derived should beused to re evaluate the closure recommendation�

�� Proper management of the medium mortality beds this year is critical� TheSARC notes that the surplus production expected in �� should be harvestedover a � year period to stabilize the �shery� Population demographics suggestthat harvesting in excess of this level will reduce SSB rapidly and result inlower allocations from this bed region and a possible closure in future years�Abundance remains below target levels Figure �� Consequently� managementof these beds should include rebuilding of abundance as a goal�

�� Evidence exists that low spat abundance is associated with low adult abun dance� and suggests that the explanation involves the contribution of live oys ter shell to the cultch resource preferred for settlement� This implies that highrecruitment may be less likely under current conditions of low abundance� Cou pled with the observation that all bay regions are below the abundance target�this trend emphasizes the essential need to include abundance enhancement inany management plan for ��

�� The SARC strongly encourages continuation of the intermediate transplantprogram and notes that any transplant option requires transplant to occurbefore the allocation derived therefrom can be set�

� The SARC strongly encourages retention of the area management program in

�

which the high mortality beds� Shell Rock� the medium mortality beds� and thelow mortality beds are managed as separate units with separately determinedallocations�

�

Figure Summary status of the stock for �� Green indicates variables judgedto be above average relative to the �� time period or having an improvingtrend relative to the previous year� Orange indicates variables judged to be belowaverage relative to the �� time period or having a degrading trend relative tothe previous year� Light green indicates near average conditions� generally de�ned

as conditions falling within the ��th to �th percentiles of the �� time period�but sometimes determined by scienti�c judgment� Fraction of stock refers to thedispersion of the stock across the salinity gradient in the four bay regions� Allpercentiles are relative to the �� time series� Parentheses are values thatinclude the �� shell plants�

Low Medium High

Mortality Beds Mortality Beds Shell Rock Mortality Beds

Fraction of Stock 0.14 0.64 0.07 0.15

Total Abundance

2006 Percentile 0.08 0.42 0.40 0.25

2005-2006 Trend Decreasing Increasing Increasing Increasing

Spawning Stock Biomass

2006 Percentile 0.27 0.67 0.67 0.67


Recruitment

2006 Percentile 0.18 0.42 0.18 (0.45) 0.08 (0.55)


Spat per Adult

2006 Ratio 0.16 0.34 0.32 (0.64) 0.42 (1.00)

2006 Percentile 0.36 0.47 0.25 (0.50) 0.18 (0.60)

2006 Juveniles (fract.<2.5") 0.63 0.46 0.53 0.55

2006 Percentile 0.06 0.13 0.25 0.38

Dermo Infection Status

2005-2006 trend increasing increasing increasing increasing

2006 Mortality Rate 0.06 0.16 0.18 0.21

2006 Percentile 0.08 0.58 0.47 0.31

Abundance Position vs

Target Below Below Below Below

Threshold Below Above Above Below

SSB Position vs

Target Below Above Above Above

Threshold Near Above Above Above

2007 Surplus Production

50th percentile mortality Positive Positive Positive Positive


��

Figure � Time series of oyster abundance� by bay region� for the Dermo era� �� High mortality� Beadons� Nantuxent Point� Strawberry� Hog Shoal� Vexton�Hawk�s Nest� New Beds� Egg Island� Ledge� Bennies� Bennies Sand� medium mortality less Shell Rock� Ship John� Cohansey� Sea Breeze� Middle� Upper Middle�low mortality� Arnolds� Upper Arnolds� Round Island�

1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109

3.5x109

4.0x109

4.5x109

Num

ber

of O

yste

rs

High Mortality Beds

Shell Rock

Medium Mortality Beds

Low Mortality Beds

��

Figure � Position of the oyster stock in �� with respect to biomass andabundance targets and thresholds� The target is taken as the median of abundanceor biomass during the �� time period� The threshold is taken as half thesevalues�

0.0x100

6.0x107

1.2x108

1.8x108

2.4x108

3.0x108

3.6x108

4.2x108

0.0x100

2.0x108

4.0x108

6.0x108

8.0x108

1.0x109

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance

Target Threshold 2003 2004 2005 2006

Low Mortality Beds

0.0x100

2.0x108

4.0x108

6.0x108

8.0x108

1.0x109

0.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109

3.5x109

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance


0.0x100

4.0x107

8.0x107

1.2x108

1.6x108

2.0x108

0.0x100

4.0x107

8.0x107

1.2x108

1.6x108

2.0x108

2.4x108

2.8x108

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance

Shell Rock

0.0x100

9.0x107

1.8x108

2.7x108

3.6x108

4.5x108

5.4x108

0.0x100

9.0x107

1.8x108

2.7x108

3.6x108

4.5x108

5.4x108

6.3x108

7.2x108

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance

High Mortality Beds

��

Figure � Time series of spawning stock biomass by bay region� Bed distributionsby region are given in Figure ��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

2.0x108

4.0x108

6.0x108

8.0x108

1.0x109

1.2x109

1.4x109

1.6x109

1.8x109

Spa

wni

ng S

tock

Bio

mas

s (g

)

High Mortality Beds

Shell Rock


Low Mortality Beds

��

Figure � Number of spat recruiting per year for the �� time series�

1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

1.0x109

2.0x109

3.0x109

4.0x109

5.0x109

6.0x109

Num

ber

of S

pat

��

Figure � The abundance of small� submarket and market size animals since ��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109

3.5x109

4.0x109

Num

ber

of O

yste

rs

<2.5 inches

2.5-3 inches

>3 inches

��

Figure � Time series of box count mortality on New Jersey Delaware Bay oysterbeds prorated by bay section� The height of each shaded area is proportional to thetotal number of deaths contributed by that bay region� The cumulative sum of thefour bay regions measures the bay wide mortality rate for that year�

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

0.45

0.5

Box

Cou

nt F

ract

ion

Year

High Mortality Beds

Shell Rock


Low Mortality Beds

�

Figure Estimated net change in sur�cial shell content in bushels by bay regionfor the New Jersey oyster beds for the time period �� Positive values onShell Rock in �� and �� re�ect the addition of shell through shell planting too�set the shell loss�

1999 2000 2001 2002 2003 2004 2005 2006-350000

-300000

-250000

-200000

-150000

-100000

-50000

0

50000

100000

Net

Cha

nge

in B

ushe

ls

Low Mortality Beds


Shell Rock

High Mortality Beds

��

Report of the�� Stock Assessment Workshop

��th SAW� for theNew Jersey Delaware Bay Oyster Beds

Presenters

Eric Powell� Haskin Shell�sh Research LaboratoryJohn Kraeuter� Haskin Shell�sh Research LaboratoryDavid Bushek� Haskin Shell�sh Research Laboratory

Stock Assessment Review Committee

Russell Babb� New Jersey Department of Environmental ProtectionBarney Hollinger� Delaware Bay Section of the Shell Fisheries Council

Roger Mann� Virginia Institute of Marine ScienceSteve Fleetwood� Delaware Bay Section of the Shell Fisheries Council

Rich Wong� Delaware Department of Natural ResourcesMike Celestino� New Jersey Department of Environmental Protection

Joe Dobarro� Rutgers UniversityThomas Landry� Department of Fisheries and Oceans� Canada

Steve Jordan� U�S� Environmental Protection Agency

Editors�

Eric Powell� Haskin Shell�sh Research LaboratoryJohn Kraeuter� Haskin Shell�sh Research Laboratory

Kathryn Ashton�Alcox� Haskin Shell�sh Research Laboratory

Distribution List

Barney Hollinger� Chair� Delaware Bay Section of the Shell Fisheries CouncilJim Joseph� New Jersey Department of Environmental ProtectionSelected faculty and sta�� Haskin Shell�sh Research Laboratory

Oyster Industry Science Steering CommitteeStock Assessment Review Committee

February ��

New Jersey Agricultural Experiment Station �� Rutgers UniversityNew Jersey Agricultural Experiment Station �� Rutgers UniversityNew Jersey Agricultural Experiment Station �� Rutgers University

Introduction

The natural oyster beds of the New Jersey portion of Delaware Bay �Figure ��have been surveyed yearly� in the fall andor winter� since �� Since � � thisperiod has been concentrated into about one week in the latter part of Octoberto early November� and has been conducted using a strati�ed random samplingmethod� Each bed is divided into �� latitude � �� longitude grids� each havingan area of approximately �� acres� These grids fall into one of three strata� thebed core �high quality�� the bed proper �medium quality�� or the bed margin �low

quality�� Each sample represents a composite of � one�third bushels from threeone�minute tows within each grid� The current survey instrument is a standard��m commercial oyster dredge on a typical large Delaware Bay dredge boat� theF�V Howard W� Sockwell�

Sample analysis includes measurement of the total volume of material obtainedin each measured dredge haul� the volume of live oysters� boxes� cultch� and debris�the number of spat� older oysters� and boxes per composite bushel� the size oflive oysters and boxes �� mm from the composite bushel� condition index� andthe intensity of Dermo and MSX infections� Until �� the principal data usedin management were based on the proportion of live oysters� excluding spat� ina composite ��quart bushel�� although spat set also entered the decision�makingprocess� Beginning in � � dredge tow lengths were measured and recorded every� seconds by GPS navigation during the survey and� in �� and ��separate dredge calibration studies were undertaken to determine dredge e�ciency�These data integrated into the regular sampling permit quantitative estimationof the number of oysters per square meter beginning in � � In �� at the

behest of the �th SAW� the entire survey time series from �� to the present�day was retrospectively quantitated� Also in �� a dock�side monitoring programbegan� This program obtains additional �shery�dependent information on the sizeand number of oysters marketed� permitting� beginning in �� the determinationof exploitation based on spawning stock biomass as well as abundance� In ��su�cient information was available from the dock�side monitoring program toreconstruct the �� exploitation rates�

Status of Stock and Fishery

Historical Overview

From �� to �� the bay�wide mean number of ��mm oysters per bushelwas about �� Table �� The highest numbers of oysters were on the beds upbayof Shell Rock and the lowest numbers were on the two most downbay beds� Egg

� A ��qt bushel is the New Jersey Standard Bushel�

�

Island and Ledge �Table �� Since � when Dermo became prevalent in the bay�the bay�wide overall mean of �� oystersbu� about half the long�term average�has varied little� and the changes� with the exception of the extremes ��

�� and �� have not been statistically signi�cant from any other year �Figure�� Throughout this report� except where noted� present�day conditions will becompared to these two periods of time� the �� period encompassing theentire survey time series and the � �� portion encompassing the period of timeduring which Dermo has been a primary source of mortality in the bay�� Statusof stock evaluations and management advice will refer exclusively to the � ��time period� because the advent of Dermo disease as an important determinant ofpopulation dynamics occurred in � and this disease has substantively controlled

natural mortality ratesz in all succeeding years� Two exceptions exist to thedependency on the � �� time series� All size�dependent indices begin in ��when size frequencies were �rst measured in survey samples� Evaluation of �sheryexploitation by abundance is focused on the �� time period during whichthe �shery has been conducted under a direct�marketing system�

The �� bay�wide mean number of spatbu was �� Table �� with thegreatest set of �� spatbu� occurring in �� Since � � the bay�wide averagehas been � spatbu� slightly less than half the long�term mean �Figure �� The

long�term �� average box�count mortality is approximately �� Table ��The appearance of Dermo in the bay has increased the average mortality since � to �� and in some years the mortality has exceeded �� Thus� both abundanceand recruitment have averaged signi�cantly lower since the onset of Dermo� whilenatural mortality rate has averaged higher� Since the direct landing of market�sizeoysters from the beds was instituted in �� the greatest landing occurred in � �� bu�� The average yearly landing since �� has been �� bu�

Survey Design

The survey has been conducted as a random survey of the twenty primaryoyster beds �Figure �� since �� with embedded strata de�ned by di�erences inabundance in the random design for much of that time� Each bed is divided into�� latitude � �� longitude grids� approximating �� acres in area� Each of thesegrids is assigned to a speci�ed stratum and a subset of grids� randomly selected� is

� Because the survey footprint changed in �� and �� as described in a subsequent sectionthe values provided in the time series plots have changed in most cases over the entiretime series in comparison to the values reported by SAW�� and SAW�� Values reportedherein are considered to be improvements in accuracy and should be used in lieu of SAW��or SAW�� values�

z Throughout the term �mortality rate applies to the fraction dying per year� Values givenare not true rates� rather they are equivalent to e�mt in the equation Nt � N�e

�mt with m inunits of yr�� and t � � yr�

�

chosen each year for survey� Prior to �� these strata were based on an historicalevaluation of relative abundance� the high�quality areas were considered areas ofthe bed with a high abundance of oysters �� or more of the time� medium�qualityareas were considered areas where oysters were abundant �� of the time� andlow�quality areas were considered areas where oysters were abundant less than ��of the time� Through �� most beds were sampled yearly� the remaining mostlyminor beds were sampled every other year� Beginning in �� sampling intensitywas revised on a number of beds to better re�ect their utilization by the �shery� and�to provide more accurate estimates of oyster abundance� fewer beds were sampledin alternate years�

Beginning in �� two important changes occurred� First� all beds weresampled each year with the exception of Egg Island and Ledge that continue toalternate due to their consistent low abundance� Second� the area from Middle toBennies Sand was re�surveyed resulting in a change in stratal de�nition and surveydesign�� In the new system� the strata for re�surveyed beds were based on orderinggrids within beds by abundance� Grids were de�ned by cumulatively accountingfor the �rst �� of the stock as low quality� the next � � of the stock as mediumquality� and the �nal �� of the stock as high quality� In �� the area downbay ofBennies Sand� except Egg Island� Ledge� and New Beds was similarly re�surveyedand this re�survey resulted in an equivalent change in stratal de�nition and surveydesign�

The spring �� re�survey of the area from Bennies Sand to Vexton exclusiveof Egg Island� Ledge� and New Beds� included all navigable ��acre grids� Thesesampled grids consisted of all previously designated grids and a number of gridsnot in the pre�� footprint� Each of the new grids were assigned to the nearestbed while maintaining simple linear boundaries between adjoining beds wheneverpossible� and given a unique grid number� In total� over �� grids were sampledover a two�week period�

Preliminary evaluation of the density of oysters among grids revealed that alarge number of grids could be deleted from the fall survey if the survey was focusedon the grids on each bed that support � of the stock on that bed� These gridswere assigned to a �low�quality� stratum� This designation is consistent with thede�nition of a low�quality grid adopted in SAW� after the re�survey of the BenniesSand to Middle reach� The remaining grids were input into a Monte Carlo modelin which grids were subsampled repeatedly� without replacement� under a givenset of rules� and the mean abundance estimated from the subsample compared tothe mean abundance obtained from the average of all grids� Analysis of manysimulations suggested that a random survey based on two strata would su�ce�

� Details of this revision can be found in HSRL� �� Report of the �� Stock AssessmentWorkshop ��th SAW� for the New Jersey Delaware Bay Oyster Beds� �� pp�

�

remembering that a third low�quality stratum had already been split out at thecost of �� of the stock� These two strata are de�ned by assigning grids orderedby increasing abundance that cumulatively account for the �rst � � of the stockto a �medium�quality� stratum and grids that cumulatively account for the upper�� of the stock to a �high�quality� stratum� These designations are also equivalentto those adopted in SAW� for the Bennies Sand to Middle reach� The new high�quality stratum generally includes most grids originally assigned to the high�qualitystratum used prior to �� and a few of the old medium�quality grids� The medium�quality stratum generally includes some of the old medium�quality and low�qualitygrids plus a number of new grids� Figure � shows the revised bed footprint de�nedby the high�quality and medium�quality strata for these beds�

Dredge e�ciency for grids surveyed in the �� spring re�survey and theprevious fall �� survey varied by a factor of �� Comparison to fall surveyabundance estimates� after this correction� revealed that the �� assessment ofthe re�surveyed beds underestimated abundance by about �� Evaluation of timeseries data identi�ed some old �� samples obtained from newly�designatedhigh�quality and medium�quality grids that did not fall within the previous bedfootprint� This suggests that not all of the �new� areas are �new� oyster bed� thatis� some portion of the �� underestimate is due to an historical rede�nition of thebed footprint used for survey design�

The October �� survey was constructed by randomly choosing a designatednumber of grids from each stratum on each bed� Sampling was conducted fromOctober �� to November �� using the oyster dredge boat F�V Howard W� Sock�well with Greg Peachey as Captain� The sampling intensity is shown in Table �and the speci�c grids sampled are shown in Figure �� Total sampling e�ort in�� was �� grids� a value about the same as �� These included �� transplantgrids selectively sampled because they were sites of �� and �� shell plants or�� intermediate transplants� The �� shell�plant grids were Bennies Sand ��and Shell Rock �� and �� The �� shell�plant grids were� Bennies Sand �� and �� Hawk�s Nest �� Nantuxent Point �� and Shell Rock �� and �� Theintermediate transplant grids were Shell Rock � and ��

In �� a few additional dredge e�ciency measurements were made for gridsinvolved in the �� shell�planting program� Dredge e�ciency experiments wereconducted on Hawk�s Nest and Nantuxent Point� Values were representative of

previous experiments� Live oyster q� averaged �� versus the �� value obtained in�� and used in this assessment� Boxes averaged �� versus �� Cultch averaged�� versus �� These additional measurements suggest that dredge e�ciency has

� The catchability coe�cient q as used herein is de�ned as the inverse of dredge e�ciency e�q � �

e�

�

changed little since �� Table ��

Oyster Abundance

Analytical Approach

The data that follow are presented in three ways� �� Data are presented interms of numbers per ��qt bushel� This is the datum used historically since theinception of the formal stock survey in �� Bay�region averages are obtainedby the averaging of survey samples per bed� summed over the beds in each bay�region group� �� Since � � swept areas have been directly measured� permittingestimation of oyster density� Bay�region point�estimates are obtained by averaging

the per�m� samples per stratum� expanding these averages for each bed accordingto the stratal area for that bed� and then summing over the beds in any bay�regiongroup� Throughout this report� these quantitative point estimates of abundancesum the high�quality �bed core�� medium�quality �bed proper�� and transplantstrata only� Low�quality areas are excluded� For much of the bay� exclusion of thelow�quality grids underestimates abundance by approximately �� Judging fromthe targeted spring re�survey of the Middle to Vexton reach� the underestimate ofabundance elsewhere in the bay is likely to be considerably larger� �� In �� the�� survey time series was retrospectively quantitated� Data including thisretrospective analysis will be termed �time�series estimates� throughout this report�These estimates were obtained by using bed�speci�c cultch density determinedempirically from � �� This quanti�cation assumes that cultch density isrelatively stable over time� Comparison of retrospective estimates for � ��obtained using the �stable cultch� assumption� with direct measurements for � �� suggests that yearly time�series estimates prior to �� may be biased bya factor of �� Cultch varies with input rate from natural mortality and thetemporal dynamics of this variation are unknown for the �� time frame�however� recent improvements in the understanding of shell dynamics on DelawareBay oyster beds show that shell is the most stable component of the survey sampleand support the belief that a �� error is unlikely to be exceeded� Accordingly� thequantitative time�series estimates are considered the best estimates for the �� time period�

All quantitative and post�� time�series estimates were corrected for dredgee�ciency using the dredge e�ciency measurements made in �� and �� Thesize�class�speci�c dredge e�ciencies were applied whenever size�class data wereanalyzed� The di�erential in dredge e�ciency between the upper and lower bedswas retained in all cases �Table ��

Throughout this report� �oyster� refers to all animals �� mm� Animals ��mm are referred to as �spat�� Adult oysters are animals �� mm� Calculations

�

of spawning stock biomass �SSB� are based on this size class and used bed� andyear�speci�c regressions between dry weight �g� and shell length �mm� to convertsize to biomass� Market�size animals are animals �� mm� Submarket size classesare variously de�ned depending on growth rates and analytical goals as indicated�Shell planting permitted an estimate of the accuracy of the ��mm size boundaryfor spat on the beds downbay of Ship John and Cohansey� �� shell plants revealedthat about �� of the spat exceeded �� mm in size and reached sizes as large as�� mm� Thus recruitment on the downbay beds may be underestimated by thisamount� Spat growth rates are unknown upbay of Shell Rock� however� the errorupbay is unlikely to exceed the �� value�

Abundance Trends

Since � � the natural oyster beds have experienced a two�fold �uctuationin the number of oysters per bushel� but� with the exception of the two highestand lowest values� no statistical di�erences exist �Figure �� High variances areto be expected because oysters are being sampled along a salinity gradient thatre�ects spat set� predation� disease� and growth� The bay�wide average number of oystersbu in �� was statistically the same as for most of the � �� period�

but �� of the long�term average of �� oystersbu �Table ��

Quantitative estimates using the time�series analysis indicate that oysterabundance summed across all strata and bay regions increased in �� after fouryears of decline to �� from the �� estimate of �� Figure ��

In �� abundance was at the ��th percentile of the �� time series and the

�th percentile for the post�� era�

Beds in the low�mortality and medium�mortality segments of the bay �seeFigure � for bed groupings� continue to support relatively high oyster abundance

�Table �� but many beds declined somewhat this year� Only Round Island� Middle�Ship John� and Shell Rock have �� oystersbu� In �� oyster abundance onbeds in the medium�mortality segment of the bay did not deviate statistically fromthe remaining years of the � �� time series �Figure �� but all beds except ShellRock experienced a slight decline on a per�bushel basis� Average oyster abundancein the high�mortality region was the same as last year and in the mid�range ofabundances of the � �� time series �Table �� Figure ��

Quantitative estimates con�rm that most �� of the oysters were on themedium�mortality beds �Ship John� Cohansey� Sea Breeze� Middle� Upper Middle��Figure �� This is a substantive increase in proportion for these beds over ��but in keeping with the distribution of oysters in most years post�� Abundance

on these beds ranked at the ��rd percentile of the ��yr time series and the

�

��nd percentile post�� Abundances also rose on the high�mortality beds andon Shell Rock in �� The high�mortality beds and Shell Rock ranked at the

� th and ��th percentiles� respectively� for the ��year time series and at the ��th and

��th percentiles post�� Abundance in �� on the high�mortality beds rosefrom �� by a factor of �� Figure �� This is the second consecutive yearabundance has increased on these beds� Abundance rose substantially as well onShell Rock �by �� and for the second consecutive year� The large increase on ShellRock was partly due to the �� shell�planting program� In contrast� abundancedeclined by �� on the low�mortality beds to one of the lowest levels ever recorded

��rd percentile� and the lowest level in the post�� era�

The dramatic decline in abundance on the low�mortality beds in �� isunsupported by mortality data reported subsequently and� so� may be� in part�a survey artifact� This section of the bay has� as yet� not been re�surveyed and theoriginal stratum footprint is unlikely to permit a stable assessment of abundanceon these beds�

SAW� projected an eventual harvest of �� bushels from the shell�plantingprogram of �� The yearly mortality rate for yearlings from the �� shell plantsin �� was �� on Bennies Sand and �� on Shell Rock� These rates are muchbelow the rate used for harvest projections by SAW� � Assuming marketable size

is reached in year � and that the mortality rate will average at the ��th percentileof the � �� time series in years � and � permits a revision of the projectedharvest from the �� program as �� bushels on Bennies Sand and �� onShell Rock� totaling �� bushels� The yearlings from these shell plants representan important source of the abundance increase observed in these bay regions in��

Spawning Stock Biomass SSB

Spawning stock biomass rose slightly bay�wide in �� Figure �� SSB

was at the ��rd percentile of the �� time series� SSB declined markedlyon the low�mortality beds� rose on the medium�mortality beds� and remainedessentially unchanged on Shell Rock and on the high�mortality beds� in keepingwith the management goals established by SAW� that sought to stabilize market�size abundance on Shell Rock and beds downbay� For the low�mortality beds� themedium�mortality beds� Shell Rock� and the high�mortality beds� the percentiles

were the ��th� ��rd� ��rd� and ��rd� respectively�

SSB is highest on the medium�mortality beds in most years �Figure �� In ��these beds contributed �� to bay�wide SSB� The high�mortality beds contributedan additional �� SSB was more concentrated on the medium�mortality beds in�� than in �� because of the decline in SSB on the low�mortality beds�

Oyster Size Frequency

Size�frequency trends were calculated for the �rst time in �� based on thecontribution of each sample to total bay�wide abundance� Perusal of the �� time series �Figure �� shows that the fraction of the population �� washigh in the early ��s� then declined somewhat� and rose again in the late ��sto early ��s� In �� bay�wide� �� of the animals were below �� and ��of the animals were � �� in size �Tables � and �� Thus� marketable animals

�� accounted for half of all animals� This size�frequency distribution is notrepresentative of the long�term trend �Figure �� In fact� the proportion of smallanimals in �� is the lowest in the �� time series� Over the timeseries� values of �� are more typical� The recent decrease in this percentageis primarily due to low recruitment rather than unusually low adult mortality� Thatis� the number of smaller oysters has declined as animals have grown to �� insize or have died� and these small oysters have not been replaced by new recruits�

The fraction of the population�� began a dramatic decline in �� downbayand �� above Shell Rock� and this size�frequency structure depauperate injuveniles has been retained since then� Small oysters accounted for �� of theanimals on the low�mortality beds in �� a fraction much below long�term trends�due to persistent very low recruitment �Figure �� More than half of all animals�� on the medium�mortality beds were � �� in size� Population percentage for� �� animals for Shell Rock was �� and for the high�mortality beds� �� Theselatter two values are above average for the �� time series� Bay�wide� of theanimals �� were �� in size �Figure �� This unbalanced size�frequencydistribution occurred in all bay areas except the low�mortality region and was mostextreme on the medium�mortality beds �Figure �� The moderate increase injuvenile proportions on Shell Rock and the high�mortality beds is substantially theresult of the shell�planting program� Nevertheless� the population remains overlyweighted towards older adult animals� Such populations are sensitive to processesincreasing adult mortality� namely disease epizootics and over�shing�

Oyster Condition and Growth

Condition bay�wide was calculated for the �rst time in �� based on thecontribution of each sample to total bay�wide abundance� Condition index declinedfrom an unusually high value in �� to a value representative of the �� timeseries �Figure �� and the decrease was similar in all areas of the bay� Condition rosedownbay� averaging on the high�mortality beds more than double the low�mortality�bed value �Figure ��

Growth rates were updated from �� for each of the bay regions� Growthof submarket individuals to market size was evaluated by three separate methods�

size�at�age� repeated measures of known individuals throughout the growth season�and annual shifts of population size�frequency peaks from one fall sampling to thenext� These measures were supplemented using information on the size changes ofcohorts of animals on clam�shell plants and in hatchery�produced cohorts held intrays in the lower bay� These latter individuals also provide a means to check thesize�at�age estimates�

All methods indicated that the di�erential in size at age for oysters acrossthe salinity gradient is established by di�erential growth in the �rst two to threeyears of life� and� after that initial growth� the year�to�year growth increment isremarkably similar throughout the bay� Data were obtained for representative bedsfor the low �Arnolds�� medium �Cohansey� Middle� and high�mortality �New Beds�

Bennies Sand� bay sections �Table �� New data were obtained for Shell Rock� butheavy exploitation of this resource and recent additions by intermediate transplanthave modi�ed the size�frequency distribution enough to reduce con�dence in thosedeterminations� Thus Shell Rock growth rates were not updated in �� Theminimum�sized animal reaching �� in one growth year was found to be� high�mortality beds and Shell Rock� �� medium�mortality beds� �� low�mortalitybeds� �� Table �� These compare to values used in �� high�mortality beds�

�� Shell Rock� �� medium�mortality beds� �� low�mortality beds� ��

While the minimum submarket sizes yielding animals of market size with oneyear�s growth are only separated by a few millimeters� the age of the animalsreaching this size is appreciably di�erent in the various parts of the bay� Timeto market size was estimated from growth rings as� high�mortality beds� �� yr�medium�mortality beds� �� yr� low�mortality beds� �� yr �Table ��

Surplus Production

Surplus production is de�ned in this assessment as the number of animals avail�able for harvest under the expectation of no net change in market�size abundanceover the year� given a speci�ed natural mortality rate and growth rate� If �shingmortality rate is set to zero� surplus production as calculated herein is equivalentto the di�erential between the number of animals expected to recruit to marketsize in a year less the number of market�size animals expected to die naturally� Inthe absence of �shing� a positive surplus production indicates that the market�sizepopulation is expected to expand in abundance� If negative� the market�size pop�ulation is expected to contract even in the absence of �shing� The model used forthe calculation assumes an uneven distribution of mortality rate during the yearas observed� however this assumption is only noteworthy if market�size animals areremoved from the population by means other than natural mortality� A detaileddescription is found in Klinck et al� ��

� Klinck J�M� E�N� Powell J�N� Kraeuter S�E� Ford and K�A� Ashton�Alcox� �� A �sheries

��

Surplus production was estimated using the ��th and ��th percentiles of naturalmortality rate� Growth rates were updated for this assessment� As a probabilisticapplication of growth rate cannot yet be done� surplus production estimates weremade using the �� growth rates from SAW� � the updated �� growth rates�Table �� and the average of the two� �� updated growth rates producedsubstantively higher surplus production estimates in comparison to the growth ratesused in �� by SAW� upbay of Shell Rock� despite the relatively small incrementin the estimate� �� mm� On the high�mortality beds� the change in growth ratesreduced surplus production from the estimate using the growth rates from SAW� �Table ��

Surplus production estimates were generally positive and in keeping withprevious estimates on the high�mortality beds and Shell Rock �Table �� Surplusproduction estimates were very high on the medium�mortality and low�mortalitybeds using the �� updated growth rates� as a consequence of a large cohort ofindividuals of about �� in size on these beds� and low natural mortality rates�This is likely the �� or �� cohort� Negative surplus production estimates occur

only twice� both using the ��th percentile of natural mortality� These are bothassociated with low growth and high natural mortality� Overall� surplus productionestimates suggest that SSB should increase in all bay regions in �� barringunusually high mortality rates� unusually low growth rates� or overharvesting�

Recruitment

Recruitment was low for the seventh year in a row� bay�wide� and very lowfor the fourth year in a row �Figures �� and �� Seven consecutive years of lowrecruitment is unprecedented in the �� time series� The bay�wide �� spatcount �mean � ��bu� was far below the long�term mean of �� spatbu �Table

�� and well below the post�� long�term mean of � spatbu �Figure �� recruitment �spatbu� was signi�cantly below six other years in the � �� timeseries� all in the ��s �Figure �� No bed achieved a spat set of �� spatbu andspat set was �� spatbu or higher on only three beds� Middle� Ship John� and Shell

Rock �Table �� spat settlement ranked at the ��rd percentile for the ��

�� time series and at the ��nd percentile post�� The higher ranking post�� indicates a long�term decline in recruitment rate relative to the earlier portion ofthe time series �Figure ��

The number of spat recruiting per oyster was similar to �� however� it

continued to be very low� �� Figure �� The �� value is at the ��rd percentileof the �� time series� Shell planting raised this ratio to �� a factor of ��

model for managing the oyster �shery during times of disease� J� Shell�sh Res� ��

��

above the �� ratio for the entire bay region� and a value near the ��th percentile�The ratio has been above �� only one year since � ��

The ratio varies from bay region to bay region with high recruitment events�de�ned as exceeding � spat per oyster� occurring simultaneously on all bay regionsinfrequently� In particular� the low�mortality and high�mortality beds often showdistinctive patterns �Table �� Viewed in this way� recruitment has been unusuallylow on the medium�mortality beds since �� but much more representative of thenormal condition on beds downbay of this region� For �� the spat�to�adult ratioswere �� and �� for the low�mortality beds� the medium�mortalitybeds� Shell Rock� and the high�mortality beds� respectively� Recruitment has beenconsistently higher downbay than upbay� per adult� for many years� The respective

percentiles for the �� time period are� ��th� ��th� ��th� and ��th� and for

the post�� era� ��th� ��th� ��th� and ��th�

Recruitment enhancement programswere successful in �� Shell�planting wascarried out in June�July� �� Ocean quahog andor surf�clam shell was plantedon Hawk�s Nest� �� bu� Nantuxent Point� �� bu� Bennies Sand� �� bu�and Shell Rock� �� bu �Figure � � Table �� This totals �� bushels� Ofthese� �� were replants from downbay shell plants �Figure � �� These latter werereplanted on Bennies Sand �Table �� Shell planting in �� enhanced recruitment

by a factor of �� bay�wide� providing �� of total bay�wide recruitment� On ShellRock� shell plants accounted for �� of total recruitment� On the high�mortalitybeds� shell plants accounted for � � of total recruitment� Spat�per�adult ratios�after including the shell plants� rose to �� on Shell Rock and �� on the high�

mortality beds �Table �� with percentile positions� respectively� of ��th and ��th forthe post�� period� Values � � are desirable because they are always associatedwith stock expansion� An increase in abundance can be anticipated on Shell Rockand beds downbay in �� barring an unusual mortality event or overharvesting�

Projections of marketable bushels from the �� shell plants were obtained byassuming a ��year time to market size and natural mortality at the juvenile rate inyear � and at the adult rate in years � and �� The mortality rates used were the

��th percentiles of the � �� time series� for Shell Rock� �� for years �� and �� respectively� for Bennies Sand� �� shellplants are expected to provide �� bushels for market in �� Table ��

Shell was planted directly on the oyster beds and downbay o� Cape Shore�Figure � �� These latter plants were replanted upbay� As in �� even directplants signi�cantly out�performed native shell� with an average of �� spat perbushel� Native shell on Bennies Sand attracted �� spat per bushel and on ShellRock� � spat per bushel in comparison� thus� the increase was about a factor of

��

�� Downbay plants average �� spat per bushel �Table �� Thus� in contrastto �� downbay plants returned more than double the spat per bushel of directplants �Table ��

Shell planted in �� continued to attract spat in �� however the rate ofattraction ��bu� was little better than native shell in the same grids ��bu�� Nev�ertheless� the net addition of shell to these beds sustained an increased recruitmentrate for a second year� Year � recruitment will contribute minimally an additional�� marketable bushels in ��

A monitoring program for settlement potential was initiated in �� The ��program showed the anticipated trend of greater spat availability downbay �Figure�� but a lower setting potential overall than in �� Figure �� The monitoringprogram suggested that two recruitment waves occurred in �� one early� in July�and downbay and another later� in August� and upbay �Figure �� High settlementpotential in the Cape Shore region conforms to the observed higher settlement rateson shell planted in this area and subsequently replanted upbay �Figure � � Table ��The higher settlement potential on Middle was realized by the higher recruitmenton this bed� The higher settlement potential on Nantuxent Point and New Beds�however� was not�

Shell Budget Projections

A shell budget was constructed using bed�speci�c half�life estimates for catch��Half lives ranged generally between � and �� years �Table �� The analyses aresubject to substantial yearly variations when analyzed retrospectively because notall beds were sampled each year in the �rst two�thirds of the time series and becausethe addition of shell beginning in �� increases the di�culty of analysis as industrydredging redistributes the shell beyond its original grid placement and the half�livesfor surf clam and ocean quahog shell may diverge substantively from that for oystershell� Outlier half�life values occurred on beds poorly sampled in the �rst two�thirdsof the survey or beds heavily impacted by shell planting in �� Three bedshave negative half�life estimates� Round Island� Upper Middle� and Sea Breeze�All three were surveyed in alternate years from �� thus the time series isinadequate�

New Jersey oyster beds have been losing on the order of �� to ��bushels of shell annually since � �Figure �� is the �rst year an estimatecan be made as � is the �rst year that full survey data are available� The shellbudget shows a substantial reduction in shell loss in �� and �� as a result of the

� Powell E�N� J�N� Kraeuter and K�A� Ashton�Alcox� �� How long does oyster shell laston an oyster reef� Estuar� Coast� Shelf Sci� ��

��

shell�planting program that has reduced by at least two�thirds the yearly de�cit�

By region� the low�mortality beds have been losing about �� bushelsannually �Figure �� This low level of shell loss is due to low taphonomic loss rates�The medium�mortality beds are losing �� to �� bushels annually due tohigher loss rates and larger total area� Shell Rock has shown a net gain since ��due to shell planting� The high�mortality beds are losing �� to �� bushelsannually due mostly to the larger area of coverage� A reduction in the rate of declinein �� is due to the substantial shell planting that occurred downbay of Shell Rock�

By bed� Ship John and Bennies have the largest negative numbers� Benniesdue to its large size and Ship John due to its high loss rate �Table �� Otherbeds exceeding a loss of � million kg per year include Arnolds� New Beds� HogShoal� Vexton� and Egg Island� Four of these beds are high�mortality beds withlow abundance and thus low rates of natural shell addition� Five beds had positivebalances in �� four of these were the beds on which shell planting occurred� The�fth is Beadons�

Mortality and Disease

MSX disease� caused by Haplosporidium nelsoni� and Dermo disease� causedby Perkinsus marinus� remain the two primary disease concerns in Delaware Bay�Following a major bay�wide MSX epizootic in the mid�� s� most of the oysterpopulation appears to have become resistant to MSX� Monitoring via standardhistological methods showed that MSX continued to be insigni�cant during ��

In general� Dermo disease� and mortality increase downbay as salinity increases�A regression between Fall Dermo disease and box�count mortality explains approx�imately �� of the variation in mortality among beds since �� Figure �� They�intercept for this regression is just below �� indicating that background �non�disease� box�count mortality is about �� The regression by bay region revealsthat backgroundmortality is about �� for all bay regions except the high�mortalityregion� where it rises to � � �Figure ��

In �� the prevalence and infection intensity of Dermo increased from ��and approached or exceeded long�term means on most beds �Figures �� and�� In �� and �� summer water temperatures were cooler than normal

� The percent of oysters in the sample with detectable infections is termed prevalence� Infectionintensity is scored along the Mackin scale from zero �� pathogen not detected� to �ve ��heavily infected� and then averaged among all oysters in the sample to calculate a weightedprevalence� A full analysis of the �� disease monitoring program is available as an HSRLreport� Bushek D� �� Delaware Bay Oyster Seedbed Monitoring Program �� StatusReport�

��

and watershed runo� was higher than normal� The increased �ow of freshwaterprobably decreased disease transmission while cooler temperatures likely reducedproliferation rates within infected oysters� Higher salinities and temperatures during�� encouraged proliferation of Dermo and weighted prevalence rose �Figure ��In �� temperature and salinity showed normal seasonal �uctuations� but bothtended to be higher than average and Dermo weighted prevalences rose again relativeto �� Figure ��

Since the onset of Dermo disease in �� three epizootics� most of them multi�year� have occurred �Figure �� with peaks in �� and �� Each ofthese epizootics was characterized by multiyear increases and decreases in diseaseintensity with a tendency for disease prevalence to follow a � year cycle� SinceFall �� prevalence greater than �� or weighted prevalence approaching ��has corresponded to mortalities exceeding �� In �� Dermo levels were intheir second year of increase following a �� low and near these prevalence andweighted prevalence thresholds �Figure �� Mortality in �� was above thatin �� but not yet at epizootic levels� Dermo levels are increasing� however�and Dermo�induced mortality is likely to increase unless environmental conditionsinhibit further development� The corollary is the natural mortality rates areexpected to be above average in ��

Quantitative box�count mortality rates were obtained by calculating the num�

ber of boxes per m� and summing over strata and beds within bay regions� Box�count mortality was �� bay�wide in �� This is a slight increase from ��but still below epizootic mortality levels �Figure �� This is the fourth year in arow with mortality below �� This trend is partly due to the concentration ofanimals on the medium�mortality beds that occurred in the early ��s as a con�sequence of the last epizootic and persisted subsequently as a result of recruitmentfailure� Despite a relatively low mortality rate in �� box�count mortality was at

the ��st percentile of the ��yr time series and at the ��th percentile post��

The high� mortality and medium�mortality beds accounted for the bulk of totalmortality in �� Figure �� Although mortality rates are lower on the medium�mortality beds �Figure � �� the total number of adults is much higher� so thatcumulative deaths nearly equaled those in the high�mortality region� The high�mortality beds su�er a much higher mortality rate �Figure � � so that total deathsare high despite lower abundance on these beds �Figure �� Box�count mortality

rose on the high�mortality market beds to �� in �� falling at the ��th percentile

of the ��year time series� but only the ��rd percentile of the post�� time series�Mortality on Shell Rock was somewhat lower at � �� with percentile positions

of ��th and ��th� respectively� The percentile rank indicates that mortality wasunusually high on Shell Rock in �� in comparison to other bay areas� Shell Rock

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accounts for part of the higher bay�wide mortality rate in ��

Box�count mortality on the medium�mortality beds was �� a value distinctlyhigher than observed in �� and in keeping with the increasing proportion of largeanimals in the population� The �� level of mortality was at the ��st percentile for

the ��year time series and the ��th percentile for the post�� time series� The�� rate� due primarily to the high proportion of older adult animals� accountsfor the remainder of the explanation for the higher bay�wide mortality rate in ��Figure ��

Box�count mortality fell to �� on the low�mortality beds in comparison to ��

The �� level of mortality is at the ��th percentile for the ��year time series and

at the ��th percentile for the post�� time series� Nothing in the mortality dataexplains the large decline in total abundance in �� on these beds �Figure ��Possibly� this variation is� in part� due to a survey artifact�

Population Dynamics Trends

Broodstock�recruitment� abundance�mortality� and mortality�recruitment re�lationships were updated� The broodstock�recruitment diagram �Figure �� sug�gests that present�day abundance limits recruitment in some way� Oyster larvaetend to set preferentially on live oysters and boxes� so that one cannot excludethe possibility that broodstock abundance modulates settlement success by beinga principal source of clean shell� The shell�planting program strongly suggests that

the bay is not larvae limited� The distribution of points in the four quadrants�

of the broodstock�recruitment diagram �yx � recruitmentbroodstock abundance�is� lowlow � �� highlow � �� lowhigh � � and highhigh � �� This is sig�ni�cantly di�erent from the expectation that one�quarter of the years should fallinto each quadrant� P � �� P � �� P � �� P � �� respectively�High recruitment events are much more likely with high abundances� However� lowrecruitment events are as likely at any abundance� The ��year average recruitmentrate expressed as the number of spat per ��mm oyster per year is �� Since� � the same long�term average has been somewhat lower� �� The long�termlikelihood of a one�year population replacement event� � spat per ��mm oyster�is �� of �� and a recruitment rate half that high occurred in �� of �� years� Since� � the same two probabilities� � of � and of � � are not signi�cantly di�erent�so that the expectation of a respectable recruitment event remains approximately��

� To assign data points to each of the four quadrants each x�y datum pair is assessed as to itsposition versus the median of the x and y values� An assignment to the �low�low quadrantfor example would occur if the datum pair was below the median of the y values and belowthe median of the x values�

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Epizootics �bay�wide mortality events greater than �� of the stock� haveoccurred in about half of the years since � �Figure �� Non�epizootic yearstend to average around �� mortality �Figures �� and �� The bay�wide average

for �� was �� a non�epizootic mortality rate� Geographic contraction of thestock� an ongoing process since �� ceased in �� Figure �� Over the previousfew years� the stock became increasingly concentrated in the central part of the baywhere mortality rates tend to be moderate� In �� of the stock was onthe medium�mortality beds above Shell Rock �Ship John� Cohansey� Sea Breeze�Middle� Upper Middle�� on the low�mortality beds �Arnolds� Upper Arnolds�Round Island�� on Shell Rock� and �� on the high�mortality beds� In ��a somewhat higher fraction� �� was present on the medium�mortality beds�In �� of the stock was on the low�mortality beds� �� on Shell Rock�and �� on the high�mortality beds� Most of the increased proportion on themedium�mortality beds was due to declining abundance on the low�mortality beds�

This stock contraction should reduce total mortality rate and therefore decreasethe chance of epizootics at low abundance �Figure �� The relationship betweenbroodstock and mortality continues to clarify as low abundance values accumulate�The distribution of points in the four quadrants �yx � mortality ratebroodstockabundance� is� lowlow � �� highlow � �� lowhigh � �� highhigh � �Figure

�� This distribution is signi�cantly di�erent from the expectation that one�quarterof the years should fall into each quadrant� P � �� P � �� P � ��P � �� Low�mortality events are much more likely with high abundance thanwith low abundance� However� epizootics are less likely to occur as well at lowestabundance levels� This is because stock contraction occurs at low abundance� Thus�epizootics are most likely to occur in a narrow window of abundance as the stockexpands from its place of refuge on the medium�mortality beds�

A relationship between box�count mortality and recruitment continues to bepresent �Figure �� The distribution of points in the four quadrants �yx �mortality raterecruitment� is� lowlow � �� highlow � �� lowhigh � ��highhigh � �� This is signi�cantly di�erent from the expectation that one�quarterof the years should fall into each quadrant� P � �� P � �� P � ��P � �� respectively� Cases of high recruitment�low mortality occur more oftenthan expected by chance� Cases of high mortality and low recruitment occur lessoften than expected by chance� Possibly� the increased number of boxes increasesrecruitment in these latter cases�

The important areas for the oyster industry are the beds in the medium�mortality and high�mortality region� Examination of the trends on the individualbeds indicates that these two regions have substantially di�erent processes control�ling oyster abundance �Figure �� The average number of oysters on the medium�

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mortality beds for the � to �� period was statistically greater than for thehigh�mortality beds �Figure �� The spat set was not statistically di�erent over

the same period �Figure �� Surplus production generally has been positive on thehigh�mortality beds and commonly negative on the medium�mortality beds upbayof Shell Rock� although modeling suggests a strongly positive surplus productionin �� as a large cohort matures �Table �� The number of spat recruiting peradult has been consistently higher on the high�mortality beds and growth rates areconsistently higher� Present information suggests that the high�mortality beds arecharacterized by multiple cohorts moving through the population of relatively equiv�alent size� whereas the medium�mortality beds are characterized by aperiodically�occurring large cohorts that dominate the population for an extended period of time�The di�erential explains the tendency for surplus�production based reference pointsto work well downbay and the tendency for exploitation�based reference points towork more consistently upbay�

Harvest

In �� bushels were landed� somewhat less than the �� direct�

market average of �� bushels� �Figure �� Figure �� shows the time�series ofoyster harvest in Delaware Bay� Since �� an intermediate transplant program hasmoved oysters among beds� In this �gure� the total stock manipulation� includingtransplant and direct�market� is identi�ed as the apparent harvest� those oysterstaken to market are identi�ed as the real harvest� Harvest has been relatively

stable during direct�marketing times and below all bay�season� years�

Beds were harvested almost continually from April � to November �� Harvest was from �� beds� Five beds accounted for nearly �� of the harvest�Cohansey �� Shell Rock �� Nantuxent Point �� Ship John

�� and Hawk�s Nest �� Table �� The recommended area�managementpolicy formulated at SAW� resulted in signi�cant catches upbay of Shell Rock� Thise�ort was concentrated on Ship John and Cohansey�

Seventy�two boats participated in the �shery and worked for a total of ��boat�days� These included �� single�dredge boats working for �� boat�days ��daysboat� and � dual�dredge boats working for �� boat�days � �� daysboat��

� Catch and e�ort data have been provided by the New Jersey Department of EnvironmentalProtection�

� Prior to �� oysters were taken from the natural beds by deck�loading them and movingthem downbay to leased grounds during a few weeks in the spring� This time period wastermed �bay season � During this time oysters were taken from beds for which survey bushelsamples contained an average oyster volume of �� This �� rule was the �rst referencepoint and was used for management decisions from the late ��s until �� hence theidenti�cation of bushel samples �� in Table ��

�

The catch per boat�day for dual�dredge boats increased for the fourth year in arow �Figure �� The catch�per�boat�day for single�dredge boats also increased

this year� rising to the highest value since �� Figure �� This stabilization orincrease in catch per boat�day may re�ect the high percentage of marketable ornearly marketable oysters on most of the exploited beds�

Total dredging impact was estimated� Five beds were covered by industrydredges more than once during �� Cohansey� Ship John� Shell Rock� NantuxentPoint� and Hawk�s Nest �Table �� Highest coverage was �� on Nantuxent Point�

No other bed exceeded ��

The number of oysters per ��qt marketed bushel averaged �� in �� a dropfrom �� in �� Table �� Of these� �� were � �� in size� Incidental captureaveraged �� per bushel� fewer than in �� These were mostly animals that couldnot be culled from chosen oysters� Size of harvested individuals was about that of�� and larger than observed in �� Table �� Most animals marketed were�� to �� in length� Catch approximated a knife�edge process with few oystersmarketed below �� Figure ��

In �� the intermediate transplant program moved animals exclusively toShell Rock� �� bushels were moved from Arnolds and �� bushels were movedfrom Middle� Cullers were used for the Middle transplant� so this transplant wasenriched in larger animals� Oysters per bushel in the Arnolds transplant averaged�� and in the Middle transplant� �� The net of all �shing and transplantactivities was that most oysters taken to market ultimately were debited from thehigh�mortality and medium�mortality beds �Figures �� and �� continuesa program begun in �� to reduce net removals from Shell Rock� This wasaccomplished in �� by transplant downbay from Middle that nearly balancedremovals� A similar result occurred in �� with transplants from Arnolds andMiddle� This was a goal of the �� management plan�

Apparent �shing mortality was �� of the stock� that is� �� of the stockwas manipulated whether through transplant or harvest� True �shing mortality was�� of the stock �Figure � �� that is� the direct�market harvest in �� removed

about �� of the stock by number� This equates to �� of the spawning stockbiomass �Figure �� Fishing mortality measured by abundance has been below

The method for estimation is described in� Banta S�E� E�N� Powell and K�A� Ashton�Alcox�� Evaluation of dredging e�ort by the Delaware Bay oyster �shery in New Jersey waters�N� Am� J� Fish� Manag� ��

� This intensity of dredging is unlikely to negatively impact these beds � Powell E�N� K�A�Ashton�Alcox S�E� Banta and A�J� Bonner� �� Impact of repeated dredging on a DelawareBay oyster reef� J� Shell�sh Res� ��

�

�� every year except one since �� shing mortality so measured was at

the ��th percentile of the ��yr time series excluding closure years� and at the

��th percentile of years post�� For the �rst time� in �� the �shing mortalityof the larger size classes could be calculated� Fishing removed �� of the animals�� in �� Figure �� This is a value representative of the time series�

By bay section� �shing and management activities removed �� and �� of the animals for the low�mortality beds� medium�mortality beds� ShellRock� and the high�mortality beds� Values for the �rst two include intermediatetransplant removals� The value for the third takes into account intermediatetransplant plantings that took place� in �� exclusively on Shell Rock� Apparentexploitation rates for Shell Rock and the high�mortality beds were �� and ��respectively� As a consequence� the majority of animals taken to market originatedabout evenly from the medium�mortality and high�mortality beds �Figure ��

Management Advice

Summary of Stock Status and Population Management Goals

Target and threshold values for SSB and abundance were recalculated basedon updated numbers for the period �� This update incorporates revisedinformation on abundance on the high�mortality beds obtained during the spring�� re�survey� Year �� was not included in determining target and thresholdvalues�

Figure �� summarizes the condition of the oyster stock throughout the NewJersey waters of Delaware Bay and by bay region� All percentiles are based on the� �� period� This period is chosen because the advent of Dermo as a majorin�uence on population dynamics began in � �� and evidence indicates asubstantive change in population dynamics as a consequence� In particular� averagemortality rates are up� the frequency of epizootics is up� the average abundance isdown� and the average recruitment rate is down with respect to the �� timeperiod� These changes commenced in the �rst part of the ��s when the �sherywas closed in most years� Harvest was signi�cant during the � �� period inonly a single year� ��

The stock presents a mixture of positive and negative indicators that approx�imately balance� Abundance is low� but abundance increased in three of four bayregions� Abundance continued to be below target levels in all bay regions �Figure�� but above threshold levels on the medium�mortality beds and Shell Rock andnear� but still below� threshold levels on the high�mortality beds� Abundance waswell below threshold levels on the low�mortality beds� however� The shell�plantingprogram promises to increase abundance on these downbay beds in �� Abun�

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dance has increased each year on the high�mortality beds since reaching a post�� low in �� Figure �� and abundance has moved in a positive direction for severalyears on Shell Rock� The stock continues to be disproportionately consolidated onthe medium�mortality beds� a process that began in the early ��s with persistentrecruitment failure and the in�uence of Dermo disease downbay�

Spawning stock biomass is relatively low bay�wide� but rose in �� Increaseswere noted in all bay regions except upbay on the low�mortality beds� SSB hasincreased steadily on the high�mortality and medium�mortality beds over the lastthree years and has risen for the last two years on Shell Rock �Figure �� SSB wasabove the biomass target in three of � bay regions and near the threshold for thelow�mortality beds�

Recruitment remains low bay�wide and particularly low on the low�mortalitybeds� high�mortality beds� and Shell Rock� A near�average recruitment eventoccurred on the medium�mortality beds� However� in no case did recruitment reachthe value of �� spat per adult� Shell planting increased recruitment on Shell Rockand the high�mortality beds and this increase brought the spat�per�adult ratio above�� on Shell Rock and up to �� on the high�mortality beds� Both ratios are normallyassociated with increasing abundance in the following year� Evidence exists thatlow spat abundance is associated with low adult abundance� and suggests thatthe explanation involves the contribution of live oyster shell to the cultch resourcepreferred for settlement� This implies that high recruitment may be less likely undercurrent conditions of low abundance�

The oyster population as a whole continues to be depauperate in the smaller

size classes� with the proportion of animals �� being below the ��th percentile

in all bay regions and no higher than the ��th percentile in three� In contrast� thisyear� surplus production is expected to permit an increase in market�size abundancebay�wide and in all bay regions� given average growth rates� barring a higher thanaverage rate of natural mortality� and not counting removals by the �shery� Thiscontinues the trend of positive surplus production in most bay regions observed overthe last few years�

Dermo disease rose to moderate levels in �� and natural mortality rates weresomewhat above average� A rising trend in Dermo disease weighted prevalence maypresage increased rates of natural mortality in ��

Fishery exploitation levels since � have been low �� of abundance per

year�� Recent improvements in collection of �shery�dependent data indicate thatexploitation in terms of biomass has been �� for most of that time� Lowexploitation rates indicate that the �shery does not have a signi�cant e�ect onthe stock and that �shing mortality is not responsible for the current conditions of

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low abundance�

Overall� the conditions on the low�mortality beds are distinctly disadvanta�geous� whereas the remaining bay regions appear to have improved since ��However� the fact that all bay regions fall below their abundance targets indicatesthat actions to enhance abundance are needed in all bay regions� A reduction in�shing e�ort will not address this need because exploitation rates are already low�however� substantial increases in exploitation rate should be avoided as the impor�tance of adults as sites for larval settlement and the continued need to minimizeshell loss reinforces the importance of maintaining biomass at or above target levels�Abundance has been enhanced on the high�mortality beds and Shell Rock by down�bay transplant and this program should be continued� The preferred mechanism toaddress low abundance is to enhance recruitment and this program began in ��focused on Shell Rock and the high�mortality beds� Additional emphasis on themedium�mortality beds is desirable�

Cultch Management Goals

Most beds not receiving shell plants in �� su�ered a loss of sur�cial shell�Continued shell planting is essential to maintain habitat quality as well as providesubstrate to enhance recruitment� Shell plants should target areas where marketableoysters grow but where cultch loss exceeds the addition of shell through naturalmortality� The Ship John region is such a case� Due to the enhanced survival ofjuveniles in this region� replants from downbay plants should be moved to selectedareas of Ship John in �� This will maximize the return from this more costlyendeavor� The same region� and Cohansey as well� have the lowest fraction of smalloysters in the size�frequency distribution of any bay region� Hence� these beds alsoshould be considered for direct shell plants in �� Downbay plants and replantsshould be expanded to the extent funds permit to enhance recruitment�

Shell Rock and Bennies Sand should not be planted in ��

The high�mortality beds have the fastest growth rates and best oysters for mar�keting� but increasing abundance in this region increases the risk of epizootics� Theshell�planting program should not exclusively target this area� however� NantuxentPoint� Hawk�s Nest� Hog Shoal� Vexton� and Strawberry should be considered asplanting locales for direct shell plants� Planting should not occur on Bennies andNew Beds as evidence indicates that oysters in this region su�er proportionatelyhigher Dermo mortality for a given disease level than the inshore beds �Figure ��

�� Management Goals

Fishery Reference Points

Evidence indicates that the oyster stock varies in its population dynamics

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within bay regions and� as a consequence� management goals must be establishedseparately for each region� Since � � a constant�abundance reference pointhas been used� Under this reference point� �shing allocation is determined bythe surplus production of the population in each region� The use of a natural

mortality rate above the ��th percentile incorporates into this reference point arebuilding plan that can be expected to increase market�size abundance� Use of the

��th percentile guards against a high level of over�shing in epizootic years� Thisreference point has been successfully applied to Shell Rock and the high�mortalitybeds� These beds have consistently attained a productivity level permitting theexpansion of market�size abundance in most years� The reference point is dependentupon good information on the past record of natural mortality and growth� Theformer is represented by an � �year time series and is therefore substantive� Thelatter is represented by relatively few measurements� but the dataset was updatedin �� Nevertheless� data on growth continue to be inadequate to estimateprobabilities� In contrast� application of the constant�abundance reference pointhas been di�cult upbay of Shell Rock where less consistent recruitment results inpopulation expansions and contractions driven by the dynamics of aperiodic largecohorts� During times when these cohorts reach market size� overharvesting is alikely outcome� if long�term stability of the �shery is a desired goal�

As a consequence� an alternate� exploitation�based reference point was pro�posed by SAW� with the recommendation that both reference points be consid�ered as management options� The exploitation reference point recognizes that the�shery has been successfully prosecuted at relatively low exploitation levels since�� SAW� promulgated an exploitation�based reference point based on the me�dian exploitation rate� de�ned in terms of the fraction of abundance removed� foreach bay region for the years �� This approach has been substantiallyrevised this year based on the �� time series and new software permittingmore accurate estimates of size�dependent exploitation rates has been implemented�As these abundance�based exploitation reference points are derived from a periodof conservative �shery management characterized by low exploitation rates� theabundance�based exploitation reference points are likely to provide conservativemanagement goals� The SARC notes that these newly�formulated reference points�based on �� data� should not be updated yearly� but retained until suchtime as a Term of Reference permits formal review based on new information�

The newly�formulated exploitation reference points are introduced with the fol�lowing cautions as to their use� Two sets of exploitation percentiles were calculated�one using the assumption that all size classes were removed proportionately and oneusing a knife�edge assumption that all size classes �� were removed proportion�ately� Insu�cient data are available for the low�mortality beds� The exploitationindices for the upper group of medium�mortality beds �Middle�Upper Middle� wereapplied also to the low�mortality beds�

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Exploitation rates can be calculated based on real removals and apparent re�movals� Real removals are de�ned as the net of the market catch� increased or deb�ited by the removals and additions by intermediate transplant� Apparent removalsare de�ned as the market catch plus removals by intermediate transplant� The twovalues are identical for beds upbay of Shell Rock because transplants to these bedshave never occurred� In some cases� negative real exploitation rates appear in thetime series for Shell Rock and the high�mortality beds because the number addedby intermediate transplant exceeds the number removed� The alternative� use ofthe apparent exploitation rates� overestimates the inherent productivity of thesebeds� however� and would permit potentially unsustainable harvest levels withoutcareful implementation of the intermediate transplant program� Use of the realexploitation rates represents a precautionary approach to managing these beds�

The SARC recommends that the real exploitation rate reference points be usedfor any analysis for direct marketing and that the reference points used should bebased on the �� values for the �� size class �Table �� The SARC

recommends that the ��th� ��th� and ��th percentiles normally be employed� Lowerpercentiles might be evaluated when abundance or SSB values are near thresholdlevels to enhance stock rebuilding� Higher percentiles might be employed in timesof high surplus production or when abundance or biomass are over target levels�however� in this case� the SARC also notes that the employment of percentile

harvests above the ��th percentile reduces the likelihood of a consistent harvestover a period of years that would otherwise be permitted by the oyster�s life span�particularly upbay of the high�mortality beds�

Intermediate transplant can be conducted by suction dredge or dry dredgewith or without a culling device� Exploitation rates for suction dredge or drydredge without a culling device should be estimated assuming all size classes areremoved proportionately� The concentration factor for culling devices is of the order

of �� a concentration factor insu�cient to use the exploitation rates for ��

animals� Thus� all intermediate transplant estimates should rely on the �all�animal�exploitation rate reference points �Table ��

Bay Region Considerations�Shell Rock Downbay

Shell Rock and the high�mortality beds have provided most of the �shed ani�mals since �� because market quality is consistently high� however a substantialfraction of these animals have originated from the medium�mortality beds throughthe intermediate transplant program �Figure �� The high�mortality beds in par�ticular are highly in�uenced by disease and therefore susceptible to rapid population

� Powell E�N� and K�A� Ashton�Alcox� �� A comparison between a suction dredge and atraditional oyster dredge in the transplantation of oysters in Delaware Bay� J� Shell�sh Res�

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declines� Juvenile mortality rates also are high� Nevertheless� these beds normallyhave been characterized by positive surplus production due to high growth ratesand adequate recruitment rates� These beds have been successfully managed usinga constant�abundance reference point since � with a precautionary componentto guard against epizootic losses� That is� the beds have been managed in what isinherently a rebuilding mode�

The SARC recognizes the need to manage these beds conservatively� Present�day abundance is near threshold levels �Figure �� requiring that rebuildingbe included as a management goal� These beds have responded positively toabundance enhancement programs by shell planting to increase recruitment� andtransplanting to increase adult abundance� Retention of these mechanisms withinyearly management plans is essential while abundance is low� The constant�abundance approach using a projection of surplus production has proven itself inthis area and contains adequate precaution� Harvest levels at variance to thosesuggested by surplus production projections� derived for example from exploitation�based reference points� should be considered carefully� However� the SARC alsonotes that surplus production estimates covered a wide range due to uncertaintyin growth rates� Thus� the SARC considers the use of exploitation reference pointsfor this bay region to be the best management option� The SARC notes that�

this year� exploitation�based reference points using the ��th to ��th percentiles fallwithin a range of the central �� of the values established by surplus production

calculations derived from a �� matrix de�ned by the ��th to ��th percentile ofnatural mortality rate and the �� updated� and their average growth rates�Table �� Thus the two approaches provide relatively equivalent allowable harvestlevels�

Due to the uniqueness of medium mortality and high production� and given itsimportance to the �shery� Shell Rock must be managed independently of the high�mortality beds and under more conservative guidelines� For surplus production

projections� the ��th percentile assumption should be retained�

Bay Region Considerations�Low�mortality Beds

These beds have rarely contributed much to the �shery and none in most yearssince �� This bay region is below threshold abundance levels and at thresholdSSB levels �Figure �� The SARC recommends that this bay region be closed in�� However� the SARC also notes that this bay region will be re�surveyed in�� and that the updated abundance and biomass estimates thus derived shouldbe used to re�evaluate this recommendation�

Bay Region Considerations�Medium�mortality Beds

These beds are susceptible to negative surplus production� Positive surplus

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production has occurred in less than half of the years since �� In contrast� highlevels of surplus production are expected to occur in �� Table �� These bedshave contributed the bulk of the stock supporting the �shery over the entire ��yearhistory of the survey� excepting the ��s high�abundance period �Figure �� Overthe �� direct�market period� these beds contributed a substantial fraction ofthe animals supporting the �shery� albeit indirectly through transplant to replaceanimals �shed from the beds farther downbay �Figure �� Abundance and SSBare highest here �Figures � and � and the animals are moderately protected from

disease �Figure �� These beds must be included in the �shery� otherwise thepressure on the beds downbay of Shell Rock will be too high�

However� the high levels of surplus production anticipated for �� should notpermit dramatically expanded exploitation of these beds� The SARC notes thatthe high surplus production rate originates from the growth of the last large cohort�recruited in the early ��s� into market size and that these beds do not havesubstantial numbers of smaller animals supporting continued stock expansion inthe future �Figure �� Thus� the surplus production anticipated for �� should beviewed as the basis for �shery yield over a number of years� Given the likely averageage of animals growing into market size of � to � years �Table �� management ofthese beds should consider that a �� recruitment event will not bene�t the �sheryfor minimally � years� Thus� the projected surplus production for �� should beviewed as minimally a ��year allotment� As an example� the surplus productionestimates for �� for Cohansey� Sea Breeze� and Ship John range from �� to

�� bushels for the ��th percentile to the ��th percentile of natural mortalityrate and average growth �Table �� A yearly harvest level above �� to ��bushels risks a reduction in allocation from this bed region in future years that maybe long�term in duration� even with a substantive recruitment event in �� The�ve�year time period has implications for the evaluation of exploitation referencepoints in the context of minimizing yearly variations in the allocation� The SARCstrongly recommends that exploitation rate percentiles permitting harvest outsideof the �� to �� bushel range be viewed as inhibitory to the long�term healthof the �shery�

SAW� recommended that management should emphasize increased directmarketing on these beds to reduce the exploitation rate downbay while stockrebuilding continues� Biomass continues to be high on these beds and continuedfocus on directing e�ort to these beds is important� The SARC assumes thatthe upper bed group �Middle and Upper Middle� will be used for intermediatetransplant exclusively and encourages intermediate transplant to Shell Rock orBennies Sand be a component of the �� program� The SARC recommends thatSea Breeze� Cohansey and Ship John be managed as direct�market beds in ��

��

Surplus Production Projections � Constant�abundance Reference Point

Surplus production projections were run under the proviso that the number ofmarket�size oysters at the end of the year would equal the number at the beginningof the year� In essence� this allocates to the �shery a number of oysters equivalentto the number expected to grow into market size during the year after accountingfor replacement of those lost to natural mortality� Projections were run usingnatural mortality rates for each bay region� with percentiles calculated from the� �� time series� An average growth rate was assumed� based on the averageof the �� and �� updated data� Submarket�size oysters were de�ned using thesmallest individual that could attain �� mm during the year� Natural mortalityrates were taken from box counts because unrecorded mortality is assumed to be

mostly juvenile� The �nd SAW recommended a precautionary approach of managing

at the ��th percentile of the box�count mortality rate� The �th SAW recommended

relaxation of this assumption to the ��th percentile for the high�mortality beds�

The SARC recommends that the ��th percentile continue to be used for Shell Rock�

but that the remainder of the direct�market beds be managed within the ��th�to�

��th percentile range� Projections assumed a continuous �shing season from April� to November �� as has been typical of the last few years� This approach permitssome harvest to be compensatory� as a certain proportion of the animals takenwould otherwise die from disease� Allocation estimates used an updated value of ��to convert market�size and submarket�size abundance to market�bushel equivalents�The updated values were obtained by direct measurement of selected bushels landed

throughout the �� seasons �Table ��y� Note that the surplus�productionoption assumes direct�marketing� Consequently� estimates are provided only forthe high�mortality beds� Shell Rock� and the lower group of medium�mortality beds�Cohansey� Ship John� Sea Breeze��

Natural AllocationBay Region Mortality Percentile �market�equivalent bushels�

High Mortality ��th��th ��q


Lower Medium Mortality ��th �� to ��

Upper Medium Mortality NAxLow Mortality NAx

y This is an average of the three years �� and the mean of the total oysters andchosen oysters� The rationale for taking the mean is that the number of attached smallanimals will vary widely between years depending on recruitment dynamics so the use of thetotal number risks underestimating the allocation� On the other hand the smaller numberdoes not account for all of the oyster removals and this undervalues the �shing mortalityrate�

��

xNA� not applicable to this reference point�qThe SARC recommends that exploitation rate reference points be used for these beds with aharvest value chosen that does not stray substantively from this range so that rebuilding ofabundance can be achieved during times of shell planting� The SARCnotes that surplus productionestimates for these beds covered a wide range� The range provided represents the central �� ofa �� matrix of estimates using the ��th �th and ��th percentiles of mortality and the ��updated �� and average �� growth rates��Allocation estimates based on a ��year distribution of �� surplus production� The SARCrecommends that exploitation rate reference points be used for these beds with a harvest valuechosen that does not stray substantively from this range if long�term harvest stability is desired�

Abundance�based Exploitation Reference Point � Direct Marketing

Projections are provided for the high�mortality beds� Shell Rock� and the lowergroup of medium�mortality beds �Cohansey� Ship John� Sea Breeze�� The SARCrecommends that this reference point be de�ned based on the exploitation recordfrom �� using the abundance of �� animals in each bay region as thebasis to estimate an exploitation index �Tables �� and �� An upper and lower

bound normally should be taken as the ��th and ��th percentiles of the ��time series using data on the total removals from each bay region �transplant or

harvest�� The SARC recognizes that lower or higher percentiles may be investigatedgiven consideration of surplus production and position of abundance and SSBrelative to threshold and target levels� This year� these bay regions are above theirSSB targets� but below their abundance targets and� below the abundance thresholdfor the high�mortality bed group� thus stock rebuilding should be included in themanagement goal� This projection uses the average numbers per marketed bushelof �� derived from the �� dock�side monitoring program�

Exploitation Number of Direct�marketBay Region Percentile Rate Animals Removed Bushels

High Mortality ��th ��

��th ��

��th ��

�th ��


��th ��

��th ��

Lower Medium Mortality ��th ��

��th ��

��th ��

Upper Medium Mortality NAxLow Mortality NAx

xNA� not applicable to this reference point�

��

�The inclusion of this percentile recommendation was not uniformly supported by the SARC�The SARC notes the following� �� This value falls within the range of values estimated usingthe constant�abundance approach� �� The high�mortality beds are above the biomass target�however a higher than anticipated mortality event is anticipated to occur on these beds in �� The high�mortality beds are below the threshold in abundance requiring the rebuilding ofabundance� however shell planting has targeted this area and initial estimates show a promisingincrease in abundance that might bring abundance above the threshold in �� The SARC wasabout evenly divided on the issue of whether the concern about rebuilding abundance shouldoutweigh the high�biomass level present considering that the volatility of the resource in this bayregion caused by the wide range of mortalites from Dermo disease may limit the long�term successof stock rebuilding programs� The SARC has chosen to include this option with the caveat that itis unlikely to be a precautionary option and with the further caveat that its inclusion should notbe considered precedent setting for application to management advise by future SARCs��The SARC notes that this value approximates the value obtained using the constant�abundancemethod with a �th percentile natural mortality rate and average growth� It is therefore a pre�cautionary option given the prediction that natural mortality rates will exceed the ��th percentilein ��The SARC notes that this value is consistent with the ��th percentile surplus productionrecommendation�

Abundance�based Exploitation Reference Point � Intermediate Transplant

The estimates assume that intermediate transplant will be conducted on theupper medium�mortality beds �Middle� Upper Middle� and that direct�marketingwill be conducted on beds downbay of these two beds� Numbers to be movedby intermediate transplant are based on the assumption that transplant involvesthe removal of all size classes approximately in proportion to their representa�tion in the population as would occur by suction dredge� deck loading by drydredge� or ine�cient culling� The estimated number of bushels to be movedis derived from the mean of the number of oysters per bushel obtained fromthe �� survey� If cullers are used� the number of bushels can be reducedby an estimated factor of �� The proportion of animals available for mar�ket is estimated based on the fraction of animals �� and these animals areconverted to bushels using the �� animalbu conversion� Note that transplantoptions will require transplant before the allocation can be set because alloca�tion estimates provided herein can only be con�rmed after the transplant is com�plete� Moreover� if cullers are used� the marketable�bushel estimate may be low�

MarketableExploitation Animals Deck�load Transplant Bushel

Bay Region Percentile Rate Removed OystersBu Bushels EquivalentsHigh Mortality NAxShell Rock NAxLower Medium Mortality NAx

Upper Medium Mortality ��th ��

��th ��

��th ��

Low Mortality CLOSED�

��

xNA� not applicable to this reference point��The SARC recommends closure of the low�mortality beds in ��

Science and Management Issues

Management Issues

Abundance is at or below the abundance threshold in most bay regions� Ashell�planting program aimed at enhancing abundance by enhancing recruitmentmust continue with the aim of planting not less than �� bushels annually�

The continuing decline in the low�mortality beds requires a plan of redressincluding evaluation of the reasons for the decline�

The dock�side monitoring programmust continue� This program is required forSSB estimates of landings� improved abundance�to�bushel conversions� estimationof the shell budget� and evaluation of exploitation rates� as well as any developmentof size� or age�based models incorporating mortality�

Science Recommendations

These science recommendations are not ordered as to priority� The SARC madespecial note� however� of the need to improve growth data� conduct the remainingre�survey program� develop new recruitment indices for animals recruiting into the�shery� and the development of a �clean cultch� index to compare to recruitment�

The Dermo monitoring program should continue� Collection of ancillary dataon mortality� size�frequency distribution� and growth rate should be continued�

A spat settlement monitoring program should be continued�

A special survey of the low�mortality bed region should occur in �� to provideimproved survey design and stock estimates�

A sampling program should be undertaken to evaluate the ��tows�per�gridsampling protocol�

A program should be developed to permit yearly re�evaluation of grid allocationto strata to take into account changes in oyster distribution on beds as a consequenceof natural population dynamics and population enhancement programs�

A growth monitoring program should be expanded with emphasis on determin�ing size at age�

A model should be formulated to provide an estimate of the amount of clean

��

shell and a clean�shell versus recruitment relationship investigated�

Recruitment�into�the �shery indices should be formulated�

A geostatistical model for estimating abundance and SSB should be investi�gated and compared to the standard estimated based on the sum of stratal averages�

Further dredge calibration information is urgently needed to determine if tow�based dredge e�ciencies are su�ciently accurate to be used in survey quanti�cationand to determine if a temporal change in dredge e�ciency is occurring or hasoccurred� This study should use experiments occurring simultaneously with thesurvey to directly test the tow�based regressions�

A size�dependent model should be expanded to include box�frequencies so thatsize�dependent mortality can be included in the assessment� Retrospective analysisshould be used to better estimate mortality rates with the goal of developing ademographic model for the population�

Conversions for improving the shell budget model should be obtained� Theseinclude the amount of cultch attached to live oysters and boxes and the conversionof cultch and shell�plant volume to weight�

An improved estimate of the numbers per bushel in catch projections might beobtained by investigating the relationship between the size�frequency of catch andpopulation size frequency at the bed level of resolution�

The use of the ��mm cut�o� for de�ning spat should be re�examined�

��

Table �� Long�term �� average numbers of oyster and spat per bushel�and percent mortality �total box count�� Low mortality � Round Island� Arnolds�and Upper Arnolds� Medium mortality � Upper Middle� Middle� Cohansey�Ship John� Sea Breeze� and Shell Rock� High�mortality Group � � BenniesSand� Bennies� New Beds� Nantuxent Point� Hog Shoal� Strawberry� Hawk�s Nest�Beadons� and Vexton� High�mortality Group � � Ledge and Egg Island�

Oyster Spat � MortalityBay Average �� Low�mortality Beds �� Medium�mortality Beds �� High�mortality Beds � Group � � � �� High�mortality Beds � Group � ��

��

Table �� sampling scheme for the October survey of the Delaware Bay oysterbeds� The numbers given are the number of samples devoted to that bed� Arrowsindicate beds with the new con�guration of strata based on the �� and �� Springre�surveys� For these beds� no low�quality grids were sampled� For the remainder�the pre�� three�stratum sampling scheme was used�

Sampled Bed High�quality Medium�quality Low�quality TransplantRound Island � � � �Upper Arnolds � � � �Arnolds � � � �Upper Middle � � � �� Cohansey � � � �� Ship John � � � �� Middle � � � �� Sea Breeze � � � �� Shell Rock � � � � Bennies Sand � � � �� Bennies � � �New Beds � � � �� Nantuxent Point � � � �� Hog Shoal � � � �� Strawberry � � � �� Vexton � � � �� Beadons � � � �� Hawk�s Nest � � � �Egg Island � � � �Ledge � � � �

��

Table � Dredge e�ciency estimates expressed as the reciprocal of the e�ciency e�

q � �e� The value q is the multiplier by which swept area estimates were converted

to per�meter�square values� The upper bay includes all beds upbay of Shell Rock�

Live BoxLive Sub� Live Live Box Sub� Box Box

Juvenile market Market Total Juvenile market Market Total Cultch�� Lower�bay ��

�� Lower�bay ��

�� Upbay �� Lower�bay ��

�� Upbay �� Lower�bay ��

� �� and �� values are taken from� Powell E�N� K�A� Ashton�Alcox J�A� Dobarro M�Cummings and S�E� Banta� �� The inherent e�ciency of oyster dredges in surveymode� J� Shell�sh Res� �� and Powell E�N� K�A� Ashton�Alcox J�N� Kraeuter�in press Re�evaluation of Eastern oyster dredge e�ciency in survey mode� Application instock assessment� N� Am� J� Fish� Manage�

��

Table �� Results of the �� random sampling program for the Delaware Baynatural oyster beds� Included for comparison are data for �� and �� Dataare displayed from the farthest upbay beds to those downbay� The column called�Bushelshaul� to the left of the column headed �Percent Oyster� indicates theaverage number of bushels brought up by the � dredge hauls from each grid� Foreach bed the percentage of oysters for each sample is presented� with rankings fromhighest to lowest� Percent oyster is based on volume of oyster in the sample dividedby the total volume of shell� oyster� and debris� Those samples that have over ��oyster are underlined� Oysters per bushel and spat per bushel are based on actualcounts adjusted to a ��quart bushel� �Size� columns indicate the number of oystersgreater than �� and their percentage based on measured size frequencies �Table

�� Condition index is a measure of the dry meat weight in an oyster relative to thehinge�to�lip �greatest� dimension� The �Percentage Mortality� value is based on thenumber of boxes counted in the samples� Prevalence is the percentage of oysterswith detectable infections by Dermo� Weighted Prevalence is the average infectionintensity �scored from � to �� of all sampled oysters� Grids selected for non�randomsampling� because of recent transplants or shell plants� are listed separately at theend of the table�

��

Table �� page ��Su

mm

ary

of th

e 20

06 R

ando

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ampl

ing

of th

e Se

ed B

eds

Tab

le 4

. D

elaw

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Bay

See

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ed d

ata

for

2006

.D

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oD

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Per

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Per

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Mor

talit

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nt P

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Pre

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No.

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t/Hei

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Hau

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0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

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Table �� page ��Su

mm

ary

of th

e 20

06 R

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ampl

ing

of th

e Se

ed B

eds

Tab

le 4

. D

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are

Bay

See

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ed d

ata

for

2006

.D

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Per

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1320

207

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6530

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7010

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28.8

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2.5

27.5

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k4.

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3.3

56.5

47.7

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Shel

l Roc

k1.

552.9

47.1

35.7

213

173

175

5247

8219

1111

100

9460

3.28

2.09

1.25

9586

104

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59.5

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347

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9327

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4.0

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Shel

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5.6

46.6

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346

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3.9

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��

Table �� page �

Sum

mar

y of

the

200

6 R

ando

m S

ampl

ing

of t

he S

eed

Bed

sT

able

4.

Del

awar

e B

ay S

eed

Bed

dat

a fo

r 20

06.

Der

mo

Der

mo

Size

Size

Con

ditio

n I

ndex

Bed

Bus

hels

/Pe

rcen

t Oys

ter

Oys

ters

/Bus

hel

Spat

/ Bus

hel

Perc

ent M

orta

lity

Perc

ent P

reva

lenc

eW

eigh

ted

Prev

alen

ceN

o./b

u. >

2.5

in.

%oy

ster

s >2

.5 in

.D

ry M

eat/H

eigh

tH

aul

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

----

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Ben

nies

1.5

6134

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Ben

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2.7

39.7

3121

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24.8

19.7

Ben

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2.1

30.3

2318

Ben

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3.3

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18.5

13.8

Ben

nies

0.5

14.6

17.3

11.9

Ben

nies

0.3

1316

.84.

0486

4616

1717

1516

1524

100

6730

2.55

1.45

0.55

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5569

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710

.49.

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2.66

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8.06

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1.74

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1.36

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2.7

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18.5

Hog

Sho

al2.

724

.420

.15.

1661

6633

1422

2328

2839

9590

85.0

2.97

2.77

1.95

3428

2256

4366

.40.

018

0.02

20.

020

Hog

Sho

al3.

721

.320

4.41

Hog

Sho

al3.

010

10.1

3.02

Hog

Sho

al1.

87.

255.

3--

New

Bed

s2.

743

.921

.232

.8N

ew B

eds

0.7

17.1

18.3

22N

ew B

eds

0.1

16.5

14.8

17.9

New

Bed

s4.

312

.712

.56.

9N

ew B

eds

4.0

8.28

5.4

5.66

3030

286

2231

2414

3310

090

653.

052.

081.

3022

1214

7340

51.7

0.02

10.

026

0.02

5N

ew B

eds

0.3

1.8

4.5

5.44

New

Bed

s0.

21.

613.

94.

98N

ew B

eds

2.0

0.86

1.9

3.17

New

Bed

s0.

00

1.6

2.04

New

Bed

s--

0

�

Table �� page ��

Sum

mar

y of

the

200

6 R

ando

m S

ampl

ing

of t

he S

eed

Bed

sT

able

4.

Del

awar

e B

ay S

eed

Bed

dat

a fo

r 20

06.

Der

mo

Der

mo

Size

Size

Con

ditio

n I

ndex

Bed

Bus

hels

/P

erce

nt O

yste

rO

yste

rs/B

ushe

lSp

at/ B

ushe

lP

erce

nt M

orta

lity

Per

cent

Pre

vale

nce

Wei

ghte

d P

reva

lenc

eN

o./b

u. >

2.5

in.

%oy

ster

s >

2.5

in.

Dry

Mea

t/Hei

ght

Hau

l20

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

0420

0620

0520

04--

----

----

----

----

----

----

----

----

----

----

----

----

----

--

Str

awbe

rry

3.7

13.8

34.1

23.5

Str

awbe

rry

0.2

2.76

31.3

8.55

Str

awbe

rry

2.0

0.96

8.2

3.68

1247

322

1837

1721

2188

9670

2.13

2.27

1.85

1124

2092

5263

.40.

018

0.02

20.

017

Str

awbe

rry

0.8

06.

82.

72S

traw

berr

y2.

81.

88S

traw

berr

y2.

7--

Haw

ks N

est

4.0

53.9

48.3

14.7

Haw

ks N

est

3.0

35.8

147.

05H

awks

Nes

t0.

927

.56.

84.

2510

159

1924

.422

1326

1026

100

100

502.

632.

531.

2058

1714

5739

72.5

0.01

80.

025

0.01

9H

awks

Nes

t3.

017

.25.

22.

475

.717

.124

Haw

ks N

est

2.6

11.6

1.9

2.05

Haw

ks N

est

1.6

6.97

0--

Bea

dons

2.3

26.6

22.2

17.9

Bea

dons

3.3

17.4

20.8

8.62

Bea

dons

1.3

13.3

19.1

6.9

Bea

dons

1.0

7.79

18.4

6.02

Bea

dons

3.3

7.14

15.1

5.23

Bea

dons

4.0

5.41

43.

9742

4823

2222

3316

1730

7585

842.

051.

941.

8419

1812

4538

53.4

0.01

50.

029

0.02

2B

eado

ns2.

70.

673.

93.

43B

eado

ns3.

43.

29B

eado

ns0

1.8

Bea

dons

--1.

13

Vex

ton

3.3

15.1

29.5

10.6

Vex

ton

2.5

14.8

15.7

9.06

Vex

ton

2.8

13.4

12.8

6.39

1424

139

94

2230

3090

8065

2.25

1.05

1.60

1217

1186

7083

.90.

022

0.03

10.

025

Vex

ton

4.8

3.01

12.8

3.69

Vex

ton

1.7

012

.31

Vex

ton

0.8

0V

exto

n0.

3--

Egg

Isl

and

----

60.1

--E

gg I

slan

d--

--8.

3--

Egg

Isl

and

----

2--

Egg

Isl

and

----

1--

Egg

Isl

and

----

0.4

----

10--

--0

----

17--

--75

----

1.65

----

8--

--79

----

0.03

0--

Egg

Isl

and

----

0--

Egg

Isl

and

----

0--

Egg

Isl

and

----

0--

Egg

Isl

and

----

0--

Egg

Isl

and

----

0--

�

Table �� page ��

Sum

mar

y of

the

2006

Ran

dom

Sam

plin

g of

the

Seed

Bed

sT

able

4.

Del

awar

e B

ay S

eed

Bed

dat

a fo

r 20

06.

Der

mo

Der

mo

Size

Size

Con

ditio

n In

dex

Bed

Bus

hels

/P

erce

nt O

yste

rO

yste

rs/B

ushe

lSp

at/ B

ushe

lP

erce

nt M

orta

lity

Per

cent

Pre

vale

nce

Wei

ghte

d P

reva

lenc

eN

o./b

u. >

2.5

in.

%oy

ster

s >2

.5 in

.D

ry M

eat/H

eigh

tH

aul

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

2006

2005

2004

----

----

----

----

----

----

----

----

----

----

----

----

----

----

Led

ge4.

01.

95--

1.96

Led

ge0.

90.

35--

1.57

Led

ge1.

30

--1.

091.

17--

1.31

0.2

--3

40--

1767

--40

0.92

--0.

701

--1

99--

83.3

----

----

Led

ge0.

40

--0.

28L

edge

0.7

0--

0L

edge

--0

Tra

nspl

ants

Ben

San

d3.

5550

.222

.229

741

240

187

102.

9B

en S

and

3.68

2835

.419

020

085

4615

33.6

Shel

l Roc

k4.

357

.167

255

284

5192

189.

73Sh

ell R

ock

1.5

52.9

37.3

264

143

1326

712

.8Sh

ell R

ock

20.3

521

10.4

Shel

ling

Shel

l Roc

k1.

4359

.460

.633

326

111

627

197.

2Sh

ell R

ock

5.58

46.6

58.7

216

276

6214

722

10.7

Shel

l Roc

k3.

9244

.156

.323

320

470

105

219.

36Sh

ell R

ock

37.1

180

809.

88H

awks

Nes

t4.

0253

.923

061

29N

antu

xent

3.54

46.4

209

4929

��

Table �� Oyster size frequency on the natural oyster beds in �� Frequencies areexpressed as the number in each size class per ��qt bushel�

Tab

le 5

. O

yste

r si

ze f

requ

ency

on

the

natu

ral o

yste

r be

ds in

200

6. F

requ

enci

es a

re e

xpre

ssed

as

the

num

ber

in e

ach

size

cla

ss p

er b

ushe

lS

ize

(mm

)R

IU

Arn

Arn

Up

Mid

Mid

Coh

Ship

JSe

abSh

ell R

Ben

SnB

enN

ant

Hog

New

Stra

wH

awN

Bea

dV

ext

Led

ge0

00

0.0

0.0

00

00.

00.

00.

00.

00.

00.

00.

00.

00.

00.

00.

00.

05

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.1

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

100.

00.

00.

00.

00.

00.

00.

00.

00.

10.

00.

10.

00.

00.

00.

00.

00.

00.

00.

015

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.1

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

200.

00.

00.

00.

00.

00.

00.

00.

00.

50.

00.

00.

00.

00.

00.

00.

00.

00.

00.

025

0.6

0.7

2.8

0.0

2.9

1.5

3.5

3.5

6.5

4.0

0.4

3.0

0.4

0.1

0.0

3.2

3.6

0.2

0.0

302.

61.

56.

30.

05.

93.

67.

21.

95.

61.

90.

73.

51.

20.

00.

02.

02.

70.

00.

035

7.8

2.4

9.5

0.0

12.4

3.3

11.1

4.0

12.8

2.7

1.8

3.8

2.1

0.1

0.0

3.9

1.9

0.0

0.0

408.

24.

611

.30.

915

.36.

512

.68.

320

.69.

64.

28.

32.

30.

30.

45.

93.

10.

00.

045

8.7

1.9

8.0

0.9

11.7

4.5

13.0

8.0

22.4

14.5

9.6

8.7

3.9

1.1

0.0

5.7

3.6

0.4

0.0

5010

.42.

85.

60.

08.

14.

59.

710

.820

.516

.610

.110

.75.

51.

80.

46.

52.

90.

60.

055

17.3

5.0

11.0

1.7

11.5

6.7

11.0

9.4

15.1

16.8

10.5

9.9

5.7

2.1

0.0

6.9

2.9

0.8

0.2

6026

.86.

218

.04.

317

.18.

012

.74.

213

.511

.811

.69.

35.

72.

80.

79.

31.

90.

20.

065

32.4

6.3

20.6

0.0

18.7

11.8

17.5

6.8

15.4

10.6

7.6

11.4

4.1

2.4

0.7

8.9

2.4

0.8

0.2

7027

.77.

818

.01.

716

.914

.220

.08.

317

.110

.26.

08.

94.

12.

91.

19.

61.

90.

60.

075

19.2

5.3

15.1

0.9

18.5

15.6

17.6

8.7

15.2

10.0

3.8

12.5

5.7

2.2

0.7

8.0

2.9

0.0

0.3

8010

.83.

19.

70.

914

.911

.815

.69.

214

.610

.14.

49.

24.

12.

82.

57.

60.

50.

20.

085

2.8

2.6

4.3

0.9

14.0

12.2

11.6

10.1

10.8

8.3

2.8

6.9

2.8

1.1

0.7

6.3

1.9

0.4

0.2

902.

40.

52.

40.

012

.210

.510

.89.

79.

44.

92.

85.

03.

91.

61.

14.

12.

41.

10.

095

0.2

0.0

0.7

0.0

8.6

7.3

6.4

5.4

6.2

5.4

2.5

3.3

1.8

1.8

0.4

3.7

1.5

1.5

0.0

100

0.0

0.3

0.7

0.9

4.5

4.5

3.7

4.2

3.1

3.1

1.6

3.5

3.4

2.1

1.1

3.5

1.7

1.5

0.2

105

0.0

0.0

0.0

0.0

2.9

1.8

3.2

4.2

2.4

1.8

1.4

2.3

1.9

1.7

1.1

3.2

1.5

0.9

0.0

110

0.0

0.0

0.0

0.0

1.1

1.3

1.4

0.5

0.5

0.8

0.8

0.8

1.4

0.8

0.0

0.7

0.9

2.5

0.0

115

0.0

0.0

0.0

0.0

0.9

1.3

0.5

1.4

0.5

0.4

1.7

0.8

0.9

0.9

0.4

1.3

0.9

0.6

0.0

120

0.0

0.0

0.0

0.0

0.2

0.0

0.0

0.5

0.1

0.4

0.6

0.6

0.2

0.5

0.4

0.6

0.7

0.8

0.0

125

0.0

0.0

0.0

0.0

0.2

0.0

0.0

0.5

0.0

0.0

0.3

0.2

0.2

0.5

0.0

0.2

0.2

0.0

0.0

130

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.2

0.0

0.0

0.2

0.0

0.0

0.4

0.0

0.0

0.0

0.4

0.0

135

0.0

0.0

0.0

0.0

0.2

0.0

0.0

0.0

0.1

0.0

0.2

0.0

0.0

0.0

0.0

0.0

0.0

0.8

0.0

140

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.4

0.0

0.0

0.0

0.0

145

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

150

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.1

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

155

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

0.0

Tot

al/B

u.17

851

144

1319

913

118

912

021

314

486

122

6130

1210

142

141

No.

Mea

sure

d82

337

877

515

882

721

1189

509

2335

1112

848

813

345

228

3354

524

574

6G

reat

er th

an 3

"35

1233

378

6671

5563

4523

4526

169

3915

111

> 3

" 20

059

1021

096

7010

475

7150

2227

2110

2015

1616

- >

3"

2004

5-

130

4968

7463

1647

9-

1611

149

99

1

Gre

ater

than

2.5

"96

2672

511

492

108

7095

6637

6534

2211

5819

121

> 2

.5"

2005

3457

720

144

112

147

103

107

7725

4428

1224

2318

17-

> 2

.5"

2004

23-

500

9810

212

931

104

7711

-22

1420

1412

111

��

Table �� Average ��year growth increment for animals reaching market �� size�the average minimal size of animals reaching market size in one year� and ages�to�market size for oysters from three bay regions�

Average AverageGrowth Minimal Size Age to

Bed Group Data Source Increment Reaching Market MarketLow mortality Arnolds �� mm �� mm � to � yrMedium mortality Middle� Cohansey �� mm � �� mm � to � yr�Shell Rock Shell Rock �� mm �� mm � to � yrHigh mortality Bennies Sand� New Beds �� mm �� mm � to � yr

�Shell Rock numbers are less certain because of relatively heavy �shing on this bed and thetransplant of relatively large numbers of animals to this bed from upbay�

��

Table �� Surplus production estimates for �� for the oyster stock on the NewJersey natural oyster beds in Delaware Bay� Projections were conducted using the

��th and ��th percentiles of natural mortality� SAW� and ��updated growthestimates and their average� and a conversion from numbers to market�equivalent�bushels using the �� average of �� oystersbushel from dock�side monitoringof landings�

�� Growth Estimate

��th Percentile Estimate ��th Percentile EstimateSurplus Production Surplus Production

Bay Region �market�equivalent bushels� �market�equivalent bushels�Low mortality �� Medium mortality �� Shell Rock �� High mortality � ��

Total ��

Average Growth �� and Updated ��


Bay Region �market�equivalent bushels� �market�equivalent bushels�Low mortality �� Medium mortality �� Shell Rock �� High mortality ��

Total ��

Updated �� Growth Estimate


Bay Region �market�equivalent bushels� �market�equivalent bushels�Low mortality �� Medium mortality �� Shell Rock �� High mortality ��

Total ��

��

Table �� The ratio of spat to oysters by bay region since the beginning of thedirect�market program� Bay regions are de�ned in Figure �� Parentheses show theratio taking into account recruitment enhancement through shell planting�

Low mortality Medium mortality Shell Rock High mortality��

��

Table �� Summary of shell�planting activities for �� Shell�planting was carriedout in late June�early July� �� Six ��acre grids received direct plants� Hawk�sNest �� Nantuxent Point �� Bennies Sand �� and Shell Rock �� and �� Twoothers were replants of shell planted o� Reeds Beach and moved upbay in lateAugust to Bennies Sand � and �� Ocean quahog shell and surf�clam shell wereused� Projections of marketable bushels assumed a ��year time to market size andnatural mortality at the juvenile rate in year � and at the adult rate in years � and

�� The mortality estimates used were the ��th percentiles of the � �� timeseries� for Shell Rock� �� for years �� and �� respectively� for theremainder� �� Bushel conversions assume �� oysters per bushel�

Bushels Spat ProjectedLocation Type of Shell Planted Planted Collected SpatBu HarvestHawk�s Nest � Surf Clam �� Nantuxent �� Quahog mix� �� Bennies Sand � Surf Clam replant �� Bennies Sand � Quahog mix� �� Bennies Sand �� Surf Clam replant �� Shell Rock �� Quahog mix� � �� Shell Rock �� Quahog mix� �� Shell Rock �� Quahog mix� ��

Total �� Quahog mix � Quahog and surf clam processed to same small size

��

Table �� Average half�lives for sur�cial oyster shell on Delaware Bay oyster beds�for the �� time period�

Location Half�life �yr�Round Island ��Upper Arnolds �� Arnolds ��Upper Middle ��Middle ��Cohansey ��Ship John ��Sea Breeze �� Shell Rock ��Bennies Sand ��Bennies ��Nantuxent Point ��Hog Shoal ��Hawk�s Nest ��Strawberry �� New Beds ��Beadons �� Vexton ��Egg Island �� Ledge ��

��

Table �� shell balance �net change from �� for Delaware Bay oyster beds�in kg per bed��

NetChange

Location ShellUpper Arnolds ��Arnolds ��Middle �� Cohansey ��Ship John ��Shell Rock �� Bennies Sand ��Bennies �� Nantuxent Point �� New Beds ��Hawk�s Nest �� Hog Shoal �� Strawberry � � �� Beadons ��Vexton �� Egg Island ��Ledge ��

��

Table �� Harvest statistics for �� Fraction covered indicates the fraction ofbed area swept by industry dredges during the �shing season� Fractions above �indicate a total swept area greater than the bed area�

Fraction Bushels Percent ofOyster Bed Bed Area �m�� Covered Harvested HarvestRound Island �� Upper Arnolds �� Arnolds �� Upper Middle �� Middle �� Cohansey �� Sea Breeze �� Ship John �� Shell Rock �� Bennies Sand �� Bennies �� Nantuxent Point �� New Beds �� Hawk�s Nest �� Hog Shoal �� Strawberry �� Beadons �� Vexton �� Egg Island �� Ledge ��

Total ��

�

Table � � Statistics for oysters going to market� obtained from dock�side monitor�ing of landings� Sizes are given in inches� Percentiles refer to the percentile sizes ofthe size�frequency distribution�

��th ��th ��th Mean Number Number � ��

Mean size percentile percentile percentile per bushel per bushel��

�

Table �� Percentiles of the real and apparent exploitation rates for oysters ��

based on the �shing record for �� The SARC recommends using the realexploitation rates for setting harvest provisions�

percentile

Shell Rock Shell Rock High MortalityBeds

High MortalityBeds

Percentile Real Apparent Real Apparent

��

��

��

��

��

��

Table �� Percentiles of the real exploitation rates for all oysters and for onebay region for oysters �� based on the �shing record for �� The uppermedium�mortality bed group is Middle and Upper Middle� The lower medium�mortality bed group is Cohansey� Ship John� and Sea Breeze� The all�oyster uppermedium�mortality percentiles are also used for the low�mortality beds� Arnolds�Upper Arnolds� and Round Island�

percentile

All Oysters All Oysters All Oysters Oysters ��

Percentile MediumMortality

Upper MediumMortality

Lower MediumMortality

Lower MediumMortality

��

��

� ��

��

��

��

Figure �� The footprint of the Delaware Bay natural oyster beds showing thelocations of the high�quality �dark shade� and medium�quality �light shade� grids�Each grid is a rectangle �� latitude � �� longitude� equivalent to approximately�� acres�

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Round Island

Arnolds

Upper Arnolds

Upper Middle

MiddleSea Breeze

Shell Rock

Ledge

Vexton

Ship John

Beadons

New Jersey

Egg Island

Delaware Bay

Nantuxent Point

Delaware

Bennies

Bennies Sand

Cohansey

New Beds

StrawberryHawk’s Nest

Hog Shoal

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��

Figure �� Average annual bay�wide oyster and spat abundance per ��qt� bushel�with �� Least Signi�cant Di�erence con�dence intervals� Underlined values arenot signi�cantly di�erent� Mean � average of annual values for � ��

Delaware Bay Seed BedsYear 1989 1992 1996 1991 2000 1993 1997 1990 1995 1999 2001 2003 2005 2002 1998 1994 2006 2004 MeanOysters 189 178 172 172 153 153 152 151 148 121 119 115 114 110 107 101 99 95 135

"------------------------------------------------------------------------------------------------------------------------------------------------------------------------"---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

"--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

Year 1991 1999 1997 1998 1995 1994 1990 1989 2000 2002 1993 2005 2004 1992 1996 2003 2006 2001 MeanSpat 268 191 151 128 127 124 112 69 55 44 44 29 27 25 22 22 21 15 79

"----------------------"----------------------------------------------------------------------------

"-----------------------------------------------------------------------------------------"------------------------------------------------------------------------------------------------------

"----------------------------------------------------------------------------------------------------------------------------------------------"--------------------------------------------------------------------------------------------------------------------------------------------

��

Figure � Delaware Bay natural oyster beds showing the locations of the ��random sampling sites �white stars� and the revised bed footprint de�ned by thehigh�quality �dark shade� and medium�quality grids �light shade�� The updatedfootprints are shown for Middle� Ship John� Sea Breeze� Cohansey� Shell Rock�Bennies Sand� Nantuxent Point� Hawk�s Nest� Hog Shoal� Vexton� Beadons� andStrawberry� The original footprints are shown for the remaining beds� Low�qualitygrids are shown in white�

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39˚ 25' 39˚ 25'Round Island

Arnolds

Upper Arnolds

Upper Middle

Middle

Sea Breeze

Shell Rock

Ledge

Vexton

Ship John

Beadons

New Jersey

Egg Island

Delaware Bay

Nantuxent Point

Delaware

Bennies

Bennies Sand

Cohansey

New Beds

StrawberryHawk’s Nest

Hog Shoal

75˚ 30'

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75˚ 20'

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39˚ 10' 39˚ 10'

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39˚ 25' 39˚ 25'

��

Figure �� Time series of oyster abundance� by bay region� High mortality�Beadons� Nantuxent Point� Strawberry� Hog Shoal� Vexton� Hawk�s Nest� New Beds�Egg Island� Ledge� Bennies� Bennies Sand� mediummortality �less Shell Rock�� ShipJohn� Cohansey� Sea Breeze� Middle� Upper Middle� low mortality� Arnolds� UpperArnolds� Round Island�

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

5.0x109

1.0x1010

1.5x1010

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2.5x1010

3.0x1010

3.5x1010

4.0x1010

Num

ber

of o

yste

rs

Year

High Mortality Beds

Shell Rock


Low Mortality Beds

��

Figure �� Average annual oyster and spat abundance per ��qt bushel forthe medium�mortality and high�mortality region for the � �� time period�Medium mortality � Upper Middle� Middle� Ship John� Cohansey� Shell Rock�High�mortality � Bennies� Bennies Sand� Nantuxent Point� Hog Shoal� New Beds�Strawberry� Hawk�s Nest� Beadons� Vexton� Underlined values are not signi�cantlydi�erent according to �� Least Signi�cant Di�erence con�dence intervals� Mean� average of annual values� � � means that are signi�cantly di�erent�


Year 1997 2000 1996 1991 1989 2003 1992 1993 1990 2005 2001 1998 1999 2002 2006 2004 1995 1994 MeanOysters 265 262 247 222 214 213 195 190 184 158 155 155 153 149 144 141 138 124 181*

"----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

High Mortality Beds

Year 1996 1989 1995 1997 1992 1990 1991 1999 1998 2006 2000 2005 2001 1993 1994 2003 2004 2002 MeanOysters 116 99 85 80 78 77 77 72 65 61 60 60 59 51 43 40 34 25 65*

"------------------------------------------------------------------------------------"--------------------------------------------------------------------------------------------------------------------------------------------

"-------------------------------------------------------------------------------------------------------------------------------------------------------"-------------------------------------------------------------------------------------------------------------------------------------------------------

"--------------------------------------------------------------------------------------------------------------------------------------------------------


Year 1999 1991 1995 1998 1994 1997 1990 2000 2002 1989 1993 2006 2004 2003 2005 1996 1992 2001 MeanSpat 351 307 179 169 140 113 103 78 75 70 60 36 36 29 28 28 28 17 95

"----------------------------"----------------------------

"-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

High Mortality Beds

Year 1991 1997 1999 1994 1995 1998 1990 1989 2000 1993 2004 2005 2002 2003 1992 2006 2001 1996 MeanSpat 255 209 146 129 121 112 98 68 52 44 26 24 22 21 20 16 14 13 73

"--------------------------------------------------"------------------------------------------------------------

"--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

��

Figure �� Time series of the fractional distribution of oyster abundance� amongbay regions� Bed distributions by region are given in Figure ��

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0

0.1

0.2

0.3

0.4

0.5

0.6

0.7

0.8

0.9

1

Oys

ter

Fra

ctio

n

High Mortality BedsShell RockMedium Mortality BedsLow Mortality Beds

��

Figure �� Time series of oyster abundance� by bay region� for the Dermo era�� Bed regions are de�ned in Figure ��

1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109

3.5x109

4.0x109

4.5x109

Num

ber

of O

yste

rs

High Mortality Beds

Shell Rock


Low Mortality Beds

�

Figure �� Time series of spawning stock biomass by bay region� Bed distributionsby region are given in Figure ��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

2.0x108

4.0x108

6.0x108

8.0x108

1.0x109

1.2x109

1.4x109

1.6x109

1.8x109

Spa

wni

ng S

tock

Bio

mas

s (g

)

High Mortality Beds

Shell Rock


Low Mortality Beds

�

Figure �� Time series of the fractional distribution of spawning stock biomassamong the bay regions� Bed distributions by region are given in Figure ��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060

0.1

0.2

0.3

0.4

0.5

0.6

0.7

0.8

0.9

1

Spa

wni

ng S

tock

Bio

mas

s F

ract

ion

High Mortality Beds

Shell Rock


Low Mortality Beds

��

Figure �� The abundance of small� submarket and market�size animals since��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109

3.5x109

4.0x109

Num

ber

of O

yste

rs

<2.5 inches

2.5-3 inches

>3 inches

��

Figure �� The abundance of small� submarket and market�size animals since ��by bay region� Bed distributions by region are given in Figure ��

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

1.0x108

2.0x108

3.0x108

4.0x108

5.0x108

6.0x108

7.0x108

8.0x108

<2.5 inches 2.5-3 inches >3 inches

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

5.0x107

1.0x108

1.5x108

2.0x108

2.5x108

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

1.0x108

2.0x108

3.0x108

4.0x108

5.0x108

6.0x108

7.0x108

Shell Rock High Mortality

Low Mortality

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109Medium Mortality

��

Figure �� The fraction of marketable animals�� that are of market�size ��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060

0.1

0.2

0.3

0.4

0.5

0.6

Fra

ctio

n of

Mar

keta

ble

Oys

ters

> 3

Inch

es

��

Figure � � Annual average condition index �dry meat weight �g�hinge�to�lip

dimension �mm��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060

0.002

0.004

0.006

0.008

0.01

0.012

0.014

0.016

0.018

Ave

rage

Con

ditio

n In

dex

��

Figure �� Annual average condition index �dry meat weight �g�hinge�to�lip

dimension �mm�� by bay group� Bed distributions by region are given in Figure��

1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060

0.005

0.01

0.015

0.02

0.025

0.03

Ave

rage

Con

ditio

n In

dex

Low Mortality

Medium Mortality

Shell Rock

High Mortality

��

Figure �� Number of spat recruiting per year for the �� time series�cumulatively by bay region� Bay regions are de�ned in Figure ��

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

1.0x109

2.0x109

3.0x109

4.0x109

5.0x109

6.0x109

7.0x109

8.0x109

9.0x109

1.0x1010

Num

ber

of S

pat

High Mortality Beds

Shell Rock


Low Mortality Beds

��

Figure �� Number of spat recruiting per year for the � �� time series�

1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.0x100

1.0x109

2.0x109

3.0x109

4.0x109

5.0x109

6.0x109

Num

ber

of S

pat

��

Figure �� The number of spat recruiting per ��mm oyster per year�19

53

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0

1

2

3

4

5

6

7

8

Rat

io o

f Spa

t to

Old

er O

yste

rs

�

Figure �� Location of �� shell plants� denoted by yellow stars� New Jerseydownbay plants are on leased grounds ��B� ��A�� Transplant locations for thesedownbay plants are denoted by purple stars� Selected high�quality oyster grounds inNew Jersey are denoted by shaded ��acre grids� Red delineates State of Delawarebeds�

75˚ 25'

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75˚ 25'

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74˚ 55'

74˚ 55'

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39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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39˚ 10' 39˚ 10'

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39˚ 20' 39˚ 20'

75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

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39˚ 10' 39˚ 10'

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75˚ 25'

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75˚ 05'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

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39˚ 20' 39˚ 20'

75˚ 25'

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75˚ 05'

75˚ 00'

75˚ 00'

74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

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39˚ 20' 39˚ 20'

75˚ 25'

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75˚ 05'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

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39˚ 20' 39˚ 20'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

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39˚ 20' 39˚ 20'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

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39˚ 20' 39˚ 20'

75˚ 25'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

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75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

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39˚ 20' 39˚ 20'

75˚ 25'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

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75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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39˚ 00' 39˚ 00'

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75˚ 25'

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75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

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39˚ 20' 39˚ 20'

75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

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75˚ 25'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

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39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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74˚ 55'

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39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

75˚ 25'

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75˚ 15'

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Shell RockNantuxent Point

Bennies Sand

Hawk’s Nest

Jigger HillLower Middle

Tongers bed

101-A 5-B

Pleansonton’s Rock Drum Bed

Silver Bed

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2006 Shell Plants

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�

Figure �� Cumulative number of spat recruiting to ��oyster�shell bags deployedin the last week of June and collected bi�weekly through September� Colors identifythe month of settlement� Increment in circle diameter indicates the number ofspat that settled during that time period� Total diameter indicates the cumulativenumber of spat� Note that circle diameter bears a nonlinear relationship to totalspat counts�

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39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

10 spatJuly set

50 spatAugust set

100 spatSeptember set

New Jersey

Delaware BayDelaware

MiddleCohansey

Shell Rock

Bennies

New Beds

554D 99C

Bayside

Money Island

Beadons Point

Bivalve

Thompson’s Beach

Cape Shore

Arnolds

Ship John

Nantuxent

Egg Island

Sea Breeze

Over the Bar

Lower Middle

Silver

Ridge

75˚ 30'

75˚ 30'

75˚ 20'

75˚ 20'

75˚ 10'

75˚ 10'

75˚ 00'

75˚ 00'

74˚ 50'

74˚ 50'

38˚ 55' 38˚ 55'

39˚ 00' 39˚ 00'

39˚ 05' 39˚ 05'

39˚ 10' 39˚ 10'

39˚ 15' 39˚ 15'

39˚ 20' 39˚ 20'

39˚ 25' 39˚ 25'

39˚ 30' 39˚ 30'

��

Figure �� Cumulative number of spat recruiting to ��oyster�shell bags deployedin the last week of June and collected bi�weekly through September since ��Station locations are depicted in Figure ��

Arn

olds

Bay

side

Mid

dle

Coh

anse

y

Sea

Bre

eze

Shi

p Jo

hn

She

ll R

ock

Ben

nies

Nan

tuxe

nt P

oint

Bea

dons

Poi

nt

New

Bed

s

Mon

ey Is

land

Egg

Isla

nd

554D

99C

Biv

alve

Tho

mps

on's

Bea

ch

Cap

e S

hore

Ove

r th

e B

ar

Low

er M

iddl

e

Silv

er

Rid

ge

0

100

200

300

400

500

600

700

Tot

al N

umbe

r of

Spa

t Col

lect

ed

2004

2005

2006

��

Figure �� Estimated number of bushels of shell lost from the New Jersey oysterbeds for the time period �� Lower estimates in �� and �� re�ect theaddition of shell through shell planting to o�set the shell loss�

1999 2000 2001 2002 2003 2004 2005 20060

100,000

200,000

300,000

400,000

500,000

600,000

700,000

Bus

hels

Los

t per

Yea

r

Year

��

Figure �� Estimated net change in sur�cial shell content in bushels by bay regionfor the New Jersey oyster beds for the time period �� Positive values onShell Rock in �� and �� re�ect the addition of shell through shell planting too�set the shell loss�

1999 2000 2001 2002 2003 2004 2005 2006-350000

-300000

-250000

-200000

-150000

-100000

-50000

0

50000

100000

Net

Cha

nge

in B

ushe

ls

Low Mortality Beds


Shell Rock

High Mortality Beds

��

Figure � � Relationship between Fall Dermo infection levels �WP�weighted

prevalence� and Fall mortality as measured by box counts� Each point correspondsto a measurement from one bed for one year�

All Beds y = 9.1435x + 9.9472R2 = 0.4079p < 0.0001

0

20

40

60

80

100

0 1 2 3 4 5 6

Dermo WP

% M

ort

alit

y

Low Mortality Beds y = 0.9151x + 11.391R2 = 0.0099p = 0.5636

0

20

40

60

80

100

0 1 2 3 4 5

Dermo WP

% M

ort

alit

y

Medium Mortality Beds y = 3.9141x + 10.285R2 = 0.2281p < 0.0001

0

20

40

60

80

100

0 1 2 3 4 5

Dermo WP

% M

ort

alit

y

High Mortality Beds y = 7.9566x + 18.034R2 = 0.2386p < 0.0001

0

20

40

60

80

100

0 1 2 3 4 5

Dermo WP

% M

ort

alit

y

��

Figure �� Mean and �� Dermo prevalence in oysters on New Jersey DelawareBay oyster beds� Error bars are �� con�dence intervals� Upper bar in each pairrepresents the �� mean� Lower bar is the �� value� Abscissa is prevalencein percent� ND� no data�

��

Figure �� Mean and �� weighted prevalence of Dermo disease on New JerseyDelaware Bay oyster beds� Error bars are �� con�dence intervals� Upper bar ineach pair represents the �� mean� Lower bar is the �� value� Abscissa isweighted prevalence in Mackin�s units� ND� no data�

��

Figure �� Time series showing the cyclic nature of Dermo disease weightedprevalence� Note the tendency for epizootics �weighted prevalences �� to be ofa number of years in duration and to occur about every � years� Error bars are ��con�dence intervals�

��

Figure �� Time series of box�count mortality on New Jersey Delaware Bay oysterbeds prorated by bay section� The height of each shaded area is proportional to thetotal number of deaths contributed by that bay region� The cumulative sum of thefour bay regions measures the bay�wide mortality rate for that year�

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

0.45

0.5

Box

Cou

nt F

ract

ion

Year

High Mortality Beds

Shell Rock


Low Mortality Beds

�

Figure �� Time series of box�count mortality on New Jersey Delaware Bay oysterbeds by bay section� The height of each shaded area measures the mortality ratein that bay region� The bay�region value can be obtained by the di�erence betweenthe top and bottom ordinate values for the region�

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0

0.2

0.4

0.6

0.8

1

1.2

1.4

1.6

1.8

Box

Mor

talit

y

Year

High Mortality Beds

Shell Rock


Low Mortality Beds

�

Figure �� Broodstock�recruitment relationship for the �� time period forthe natural oyster beds of Delaware Bay� Latest year listed as �� because theplot compares end�of�� oyster abundance with �� recruitment�

0.0x100

5.0x109

1.0x1010

1.5x1010

2.0x1010

2.5x1010

4.0x10104.5x10105.0x1010

0.0x

100

5.0x

109

1.0x

1010

1.5x

1010

2.0x

1010

3.0x

1010

3.5x

1010

4.0x

1010

Num

ber

of S

pat

Number of Adult Oysters

1950s

1960s

1970s

1980s

1990s

2000s

2005

All Strata

�

Figure �� The relationship between oyster abundance and box�count mortalityfor the �� time period for the natural oyster beds of Delaware Bay� Latestyear listed as �� because the plot compares end�of�� oyster abundance with�� mortality�

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

0.45

0.5

0.0x100 5.0x109 1.0x1010 1.5x1010 2.0x1010 2.5x1010 3.0x1010 3.5x1010 4.0x1010

Box

Cou

nt F

ract

ion

Broodstock

1950s

1960s

1970s

1980s

1990s

2000s

2005

�

Figure �� The relationship between recruitment and box�count mortality for the�� time period for the natural oyster beds of Delaware Bay�

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

0.45

0.5

0.0x

100

5.0x

109

1.0x

1010

1.5x

1010

2.0x

1010

2.5x

1010

3.0x

1010

4.5x

1010

5.0x

1010

Box

Cou

nt F

ract

ion

Number of Spat

1950s

1960s

1970s

1980s

1990s

2000s

2006

�

Figure �� Number of bushels harvested from the natural oyster beds of DelawareBay since the inception of the direct�market program�

1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 20060

20000

40000

60000

80000

100000

120000

140000

Bus

hels

Har

vest

ed

�

Figure � Number of oysters harvested from the natural oyster beds of DelawareBay� Prior to �� the bay�season �shery removed oysters from the beds andtransplanted them downbay to leased grounds� The direct�market �shery began in�� In �� an intermediate transplant program began� In this �gure� since ��the total stock manipulation� including transplant and direct�market is identi�ed asthe apparent harvest� those oysters landed are identi�ed as the real harvest� Zerosrepresent years of �shery closure�

1953

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0.0x100

5.0x107

1.0x108

1.5x108

2.0x108

2.5x108

3.0x108

3.5x108

4.0x108

4.5x108

5.0x108

5.5x108

6.0x108

Num

ber of Oysters H

arvested

Year

Real Numbers

Apparent Numbers

�

Figure �� Catch �in bushels� per boat�day�

Delaware Bay Market Beds

0

10

20

30

40

50

60

70

80

90

100

110

120

130

1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 2006

Catc

h pe

r Bo

at D

ay

Single Dredge Dual Dredge

�

Figure �� Size frequency of oysters landed in �� Size class values are the meanof the size class�

0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 4.25 4.75 5.25 5.750

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

0.45

Siz

e F

requ

ency

Size (inches)

�

Figure �� Fishing mortality rates by bay region during the �� timeperiod� After �� the total re�ects both the direct�market removals and thosetransplanted by the intermediate transplant program� Bed groups de�ned in Figure�� Negative numbers indicate bay regions in which the addition of animals bytransplant exceeded the loss due to �shing�

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

-0.1

0

0.1

0.2

0.3

0.4

0.5

Rea

l Fis

hing

Mor

talit

y

Low Mortality Beds


Shell Rock

High Mortality Beds

�

Figure �� Fishing mortality rates by bay region during the �� timeperiod� The total re�ects both the direct�market removals and those transplantedby the intermediate transplant program� Bed groups de�ned in Figure �� Negativenumbers indicate bay regions in which the addition of animals by transplantexceeded the loss due to �shing�

1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 2006-0.04

-0.02

0

0.02

0.04

0.06

0.08

0.1

Rea

l Fis

hing

Mor

talit

y

Low Mortality Beds


Shell Rock

High Mortality Beds

Figure �� Fishing mortality rates during the �� time period� In this�gure� since �� the total stock manipulation� including transplant and direct�market is identi�ed as the apparent rate� those oysters landed are identi�ed as thereal rate� Zeros represent years of �shery closure�

1955

1957

1959

1961

1963

1965

1967

1969

1971

1973

1975

1977

1979

1981

1983

1985

1987

1989

1991

1993

1995

1997

1999

2001

2003

2005

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

0.45

0.5

Fis

hing

Mor

talit

y R

ate

Fishing Mortality Rate--Apparent

Fishing Mortality Rate--Real

Figure �� Fishing mortality rate during the �� time period based on SSB�

1997 1998 1999 2000 2001 2002 2003 2004 2005 20060.00

0.01

0.02

0.03

0.04

Fis

hing

Mor

talit

y R

ate

�

Figure �� Fishing mortality rate during the �� time period based onmarketable abundance �animals ��

1997 1998 1999 2000 2001 2002 2003 2004 2005 20060

0.01

0.02

0.03

0.04

0.05

0.06

Exp

loita

tion

Rat

e

�

Figure �� Summary status of the stock for �� Green indicates variables judgedto be above average relative to the � �� time period or having an improvingtrend relative to the previous year� Orange indicates variables judged to be belowaverage relative to the � �� time period or having a degrading trend relative tothe previous year� Light green indicates near�average conditions� generally de�ned

as conditions falling within the ��th�to��th percentiles of the � �� time period�but sometimes determined by scienti�c judgment� Fraction of stock refers to thedispersion of the stock across the salinity gradient in the four bay regions� Allpercentiles are relative to the � �� time series� Parentheses are values thatinclude the �� shell plants�

Low Medium High

Mortality Beds Mortality Beds Shell Rock Mortality Beds

Fraction of Stock 0.14 0.64 0.07 0.15

Total Abundance

2006 Percentile 0.08 0.42 0.40 0.25


Spawning Stock Biomass

2006 Percentile 0.27 0.67 0.67 0.67


Recruitment

2006 Percentile 0.18 0.42 0.18 (0.45) 0.08 (0.55)


Spat per Adult

2006 Ratio 0.16 0.34 0.32 (0.64) 0.42 (1.00)

2006 Percentile 0.36 0.47 0.25 (0.50) 0.18 (0.60)

2006 Juveniles (fract.<2.5") 0.63 0.46 0.53 0.55

2006 Percentile 0.06 0.13 0.25 0.38

Dermo Infection Status

2005-2006 trend increasing increasing increasing increasing

2006 Mortality Rate 0.06 0.16 0.18 0.21

2006 Percentile 0.08 0.58 0.47 0.31

Abundance Position vs

Target Below Below Below Below

Threshold Below Above Above Below

SSB Position vs

Target Below Above Above Above

Threshold Near Above Above Above

2007 Surplus Production



�

Figure �� Position of the oyster stock in �� with respect to biomass andabundance targets and thresholds� The target is taken as the median of abundanceor biomass during the � �� time period� The threshold is taken as half thesevalues�

0.0x100

6.0x107

1.2x108

1.8x108

2.4x108

3.0x108

3.6x108

4.2x108

0.0x100

2.0x108

4.0x108

6.0x108

8.0x108

1.0x109

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance

Target Threshold 2003 2004 2005 2006

Low Mortality Beds

0.0x100

2.0x108

4.0x108

6.0x108

8.0x108

1.0x109

0.0x100

5.0x108

1.0x109

1.5x109

2.0x109

2.5x109

3.0x109

3.5x109

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance


0.0x100

4.0x107

8.0x107

1.2x108

1.6x108

2.0x108

0.0x100

4.0x107

8.0x107

1.2x108

1.6x108

2.0x108

2.4x108

2.8x108

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance

Shell Rock

0.0x100

9.0x107

1.8x108

2.7x108

3.6x108

4.5x108

5.4x108

0.0x100

9.0x107

1.8x108

2.7x108

3.6x108

4.5x108

5.4x108

6.3x108

7.2x108

Spa

wni

ng S

tock

Bio

mas

s (g

)

Abundance

High Mortality Beds

�

Figure � � Relationship between the long�term mean of natural mortality esti�mated from Fall box counts and the long�term mean intensity of Dermo infectionssince �� Data are individual bed estimates� Note that the increase in mortal�ity appears to be a step function with thresholds at weighted prevalences of about� and � on the Mackin scale� Note that the inshore high�mortality beds tend towithstand higher infection intensities for a given degree of mortality�

�

status of the sto c - rutgers university · status of the sto c k figure summarizes the condition...

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