the fishes of southwestern thailand, the andaman …

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Phuket mar. biol. Cent. Res. Bull. 70: 29–77 (2011) THE FISHES OF SOUTHWESTERN THAILAND, THE ANDAMAN SEA – A REVIEW OF RESEARCH AND A PROVISIONAL CHECKLIST OF SPECIES Ukkrit Satapoomin Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand Email: [email protected] ABSTRACT: A brief review of recent ichthyological research and compiled results on the records of fishes from the Andaman Sea coast of Thailand are presented. A total of 1,746 species in 198 families of fishes are currently known from the area. The 10 most speciose families are the Gobiidae (227 species), Labridae (78), Pomacentridae (71), Serranidae (61), Apogonidae (60), Blenniidae (52), Carangidae (52), Scorpaenidae (49), Lutjanidae (39) and Chaetodontidae (37), and these together account for 42% of the total fish fauna. The ichthyofauna is dominated by reef-associated fishes (983 species) and pelagic/benthic fishes inhabiting offshore habitats (971 species). A zoogeographic analysis reveals a peculiarity of the fauna of this marginal-sea region, viz., being one of the areas of sympatry of Indian and Pacific Ocean fishes, as well as harbouring regional endemics. The PMBC Reference Collection currently contains examples of only about 63% of the fishes known from the area, indicating a need for further local sampling campaigns and/or international collaborative exploratory research programs in the Andaman Sea to build up the Reference Collection. Keywords: Thailand, Andaman Sea, fishes, checklist, diversity. INTRODUCTION The Andaman Sea is an approximately 1,200 km long x 650 km wide body of water to the southeast of the Bay of Bengal. It is bounded by the Gulf of Mataban in the north, the Malay Peninsula in the east, the Andaman and Nicobar Islands in the west and the northern tip of Sumatra in the south. At its southeastern corner, the Andaman Sea narrows to form the Strait of Malacca, which separates the Malay Peninsula from the island of Sumatra. Along an approximate 1,000 km length of the Andaman Sea coast of Thailand, a variety of marine and coastal habitats are present, viz., mangroves, mud-flats, seagrass beds, rocky shores, intertidal sand-flats, coral reefs, open pelagic zones, and complex seabed systems on the continental shelf. The integrity of different peculiar environmental settings and/or characteristics among those habitat types provides suitable refuges for a vast variety of aquatic biota, including fishes. Although, biogeographically, it belongs to the eastern Indian Ocean, it is strongly influenced by the western Pacific Ocean biota via the northwesterly through-flow in the Strait of Malacca. It is thus an area of distributional confluence of the biotas of both the Indian Ocean and the Pacific Ocean (Satapoomin, 2002a, 2007; Kimura and Satapoomin, 2009). Being a marginal sea separated from the main Indian Ocean by the Andaman-Nicobar island arcs, it has been subjected to a profound effect of sea-level lowering, especially during major glaciations when it became a separated, enclosed basin. Such isolation processes are believed to have promoted speciation and/or endemism of the fish fauna in the Andaman Sea (e.g., McManus, 1985; Randall, 1998; Randall and Satapoomin, 1999). The area is thus characterized by its high diversity and the unique faunal composition of the fishes. However, knowledge and information on the overall diversity of marine fishes in the area is still limited, because very few attempts had been made to compile a list of all known fish species from the area. The purposes of the present paper are to provide a brief review of recent ichthyological research, and to establish a comprehensive checklist of all known marine and brackish-water

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Phuket mar. biol. Cent. Res. Bull. 70: 29–77 (2011)

THE FISHES OF SOUTHWESTERN THAILAND, THE ANDAMAN SEA – A REVIEW OFRESEARCH AND A PROVISIONAL CHECKLIST OF SPECIES

Ukkrit Satapoomin

Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, ThailandEmail: [email protected]

ABSTRACT:Abrief review of recent ichthyological research and compiled results on the recordsof fishes from the Andaman Sea coast of Thailand are presented. A total of 1,746 species in 198families of fishes are currently known from the area. The 10 most speciose families are theGobiidae (227 species), Labridae (78), Pomacentridae (71), Serranidae (61), Apogonidae (60),Blenniidae (52), Carangidae (52), Scorpaenidae (49), Lutjanidae (39) and Chaetodontidae (37),and these together account for 42% of the total fish fauna. The ichthyofauna is dominated byreef-associated fishes (983 species) and pelagic/benthic fishes inhabiting offshore habitats (971species). A zoogeographic analysis reveals a peculiarity of the fauna of this marginal-sea region,viz., being one of the areas of sympatry of Indian and Pacific Ocean fishes, as well as harbouringregional endemics. The PMBC Reference Collection currently contains examples of only about63% of the fishes known from the area, indicating a need for further local sampling campaignsand/or international collaborative exploratory research programs in the Andaman Sea to build upthe Reference Collection.

Keywords: Thailand, Andaman Sea, fishes, checklist, diversity.

INTRODUCTION

The Andaman Sea is an approximately1,200 km long x 650 km wide body of water tothe southeast of the Bay of Bengal. It is boundedby the Gulf of Mataban in the north, the MalayPeninsula in the east, the Andaman and NicobarIslands in the west and the northern tip of Sumatrain the south. At its southeastern corner, theAndaman Sea narrows to form the Strait ofMalacca, which separates the Malay Peninsula fromthe island of Sumatra. Along an approximate 1,000km length of the Andaman Sea coast of Thailand,a variety of marine and coastal habitats are present,viz., mangroves, mud-flats, seagrass beds, rockyshores, intertidal sand-flats, coral reefs, openpelagic zones, and complex seabed systems on thecontinental shelf. The integrity of different peculiarenvironmental settings and/or characteristicsamong those habitat types provides suitable refugesfor a vast variety of aquatic biota, including fishes.Although, biogeographically, it belongs to theeastern Indian Ocean, it is strongly influenced bythe western Pacific Ocean biota via the

northwesterly through-flow in the Strait ofMalacca. It is thus an area of distributionalconfluence of the biotas of both the Indian Oceanand the Pacific Ocean (Satapoomin, 2002a, 2007;Kimura and Satapoomin, 2009). Being a marginalsea separated from the main Indian Ocean by theAndaman-Nicobar island arcs, it has been subjectedto a profound effect of sea-level lowering, especiallyduring major glaciations when it became aseparated, enclosed basin. Such isolation processesare believed to have promoted speciation and/orendemism of the fish fauna in the Andaman Sea(e.g., McManus, 1985; Randall, 1998; Randall andSatapoomin, 1999). The area is thus characterizedby its high diversity and the unique faunalcomposition of the fishes. However, knowledgeand information on the overall diversity of marinefishes in the area is still limited, because very fewattempts had been made to compile a list of allknown fish species from the area.

The purposes of the present paper are toprovide a brief review of recent ichthyologicalresearch, and to establish a comprehensivechecklist of all known marine and brackish-water

Phuket mar. biol. Cent. Res. Bull.30

fishes from the Andaman Sea coast of Thailand.The checklist was compiled from various sourcesof information, including published literature,unpublished data made available under severalsampling programs/projects, and the fish collectiondata repository of the Phuket Marine BiologicalCenter (PMBC). It should be noted that the overalllist of species presented herein is only provisionalas a number of species records and identificationsin previously published literature (particularly thoseof inventory, non-taxonomic works), cannot beverified. Nonetheless, this provisional checklist willserve as a significant baseline for furtherestablishment of a conventional, annotatedchecklist or taxonomic review of the Andaman Seafishes.

METHODS

The compilation of the species list wasbased largely on accumulating information fromthe earlier literature, including inventory works and/or faunistic surveys, field guides, and taxonomicworks. Some unpublished data, including recentunpublished survey results made by the author,collection data of the Royal Ontario Museum’sproject on gobioid fishes of southwestern Thailandin 1993 (under auspices of Dr. RichardWinterbottom), and the PMBC fish collection datarepository were also incorporated into the currentchecklist. Selected taxonomic papers pertaining torecords of fishes from the Andaman Sea coast ofThailand used to compile the current checklist arelisted in Appendix B.

The systematic arrangement of familiesfollows Nelson (2006). Species in each family arearranged alphabetically. The scientific names offishes used in the cited literature have been updatedwhere possible, following recent usage, and hencejunior synonyms were omitted. Since thecompilation was also based, in part, on literaturethat provided only a list of names, the author hasonly been able to check or correctmisidentifications to a limited extent. In this regard,only correct names were retained, synonyms wereallocated according to the latest usage, whiledoubtful species records were excluded.

In addition to scientific names andclassification, the checklist provides basic

information on habitats and zoogeographicdistribution of each species. The availability ofvoucher specimens lodged at the ReferenceCollection of the PMBC was also emphasized inorder to draw the attention of ichthyologists whomay wish to examine these specimens. Thedesignation of habitat type(s) for each species wasbased largely on either quoted information fromthe compiled literature or from the author’sunpublished survey results and observations. Fourprincipal habitat categories adopted herein includedmangrove (Mg), coral reef (Cr), seagrass bed (Sg),and other offshore habitats (OShr). Thesecategories can be considered as goodrepresentations of the major marine and coastalfeatures along the southwestern Thailand coastline.Each of these habitat types, however, was moreor less loosely defined in order to accommodate arange of associated minor biotopes of each habitat.The offshore category, in particular, was the mostinclusive term as it encompassed a range ofbenthic- and pelagic-related habitats. Thisdesignation includes both near-shore and far-offshore fishes considered as either pelagic (i.e.,coastal pelagic, offshore pelagic, epipelagic, andmesopelagic) or demersal (i.e., coastal demersal,benthopelagic, and deep demersal) species. Themangrove habitat was considered to include certaininter-connected zones like mud-flats and the upperreaches of the estuarine areas. Coral reef wasdefined to include several nearby and/or associatedareas, namely intertidal rocky shore/rock platform,intertidal sand-flat, and subtidal bare-seafloor infront of the reef (so called as forereef).

Most of the well defined zoogeographicdistribution categories (see, e.g., Myers, 1991)were generally adopted or redefined with additionalexplanation, while some new categories were addedwith their definitions. The definitions for thosedistribution categories and their acronyms are asfollows: Indo-Pacific (IP), widespread across theIndian Ocean and western portion of the Pacificwith an eastern limit east of non-marginal areas ofthe Pacific Plate; Indo-west Pacific (IWP), a subsetof the Indo-Pacific with northeastern limit on thePacific Plate, marginally in the Carolines but notreaching the Marshalls; West Pacific (WP), limitedto the area bound by the Andaman Island andCocos-Keeling Island in the west to the margin of

31The fishes of southwestern Thailand

the Pacific Plate in the east, north to southernJapan, south to western and eastern Australia; Westcentral Pacific (WCP), west Pacific and PacificPlate distributions and may include the East Pacific;Indo-Australian Archipelago (IA), a subset of theWest Pacific, north to southern Taiwan, east tothe Solomons, south to the Great Barrier Reef;Indo-Malayan Archipelago (IM), a subset of Indo-Australian which is limited to the areas withinbounds of Malay Peninsula, Sumatra, Java, NewGuinea and Philippines; Indian Ocean (IO),widespread throughout the Indian Ocean; NorthernIndian Ocean (NIO), the extended range of easternIndian Ocean to encompass the northwesternlimits which include the west coast of India,Pakistan, Oman and Persian Gulf; Eastern IndianOcean (EIO), the eastern section of the IndianOcean westwards to Laccadive, Maldive andChagos archipelagoes; the eastern extremity of theIndian Ocean (EEIO), a narrow stripe or transitionarea along the joint margin of Indian-Australianand Asian lithospheric plates, including theAndaman Sea, Sumatra, Java, Bali, Flores, Cocos-Keeling/Christmas Islands and northwesternAustralian Shelf; Andaman Sea province (AS), thearea assigned to accommodate species havingrestricted distribution only to the Andaman Sea orranging southwards only to western Sumatra;Circumtropical (CIR), tropical seas worldwide; andCosmopolitan (COS), tropical and temperate seasworldwide. The boundary of some distributioncategories is presented (Fig. 1) for clarification ofthe definitions of those areas.

RESULTSAND DISCUSSION

1. Brief review of ichthyological researchThe fish fauna of the Andaman Sea coast

of Thailand was poorly documented until relativelyrecent times. There were several well-knownwestern ichthyologists/naturalists (e.g., Cuvier,Valenciennes, Bleeker, Günther, Fowler, Regan,Boulenger, etc.) who reported, listed or describedThai fishes between 1800 and the 1950s, but theirworks were restricted only to either the Gulf ofThailand (marine) or inland (freshwater) fishes(see Monkolprasit et al. (1997) for detailedhistorical review). The first marine fish of theAndaman Sea coast of Thailand described by a

westerner was a reef-associated goby, Pipidoniaquinquecincta Smith, 1931 [valid as Gobiopsisquinquecincta (Smith)]. During his term of service(1923–1935) as the Adviser in Fisheries to theDepartment of Fisheries of Thailand (formerlySiamese Bureau of Fisheries), Smith published anumber of articles pertaining to freshwater fishesand fishes of the Gulf of Thailand. Later, in 1945,he published a monograph entitled “The Fresh-water Fishes of Siam, or Thailand” (Smith, 1954).Suvatti (1950) was the first Thai ichthyologist topublish a list of Thai fishes. Several subsequentstudies on marine fishes in Thai waters by theThais were confined to groups of particularinterest, for instance Pleuronectiformes (byPunpoka, 1964), Lutjanidae (Banasopit, 1968),Serranidae (Banasopit, 1969; Monkolprasit, 1983),Chaetodontidae (Banasopit, 1971), Leiognathidae(Monkolprasit, 1973), Elasmobranchii(Monkolprasit, 1984), and Labridae (Monkolprasit,1989). The foremost comprehensive compiledchecklist of the fishes of Thailand, which includedboth freshwater and marine fishes, was publishedby Monkolprasit et al. in 1997. It listed 171 fishfamilies with a total of 1,741 species, about 30%(516 species) of which can be considered asfreshwater fishes. The records of marine fishesin the checklist were compiled from the literaturepublished before 1990, except for two papers (oneon mangrove fishes by Monkolprasit (1994) andone on ammodytid fishes by Ida et al. (1994)).That checklist recorded 710 species representing129 families of fishes from the Andaman Sea.

A few collections of reef fishes from theSimilan Islands were made during the R/V Te VegaCruise 2, eastern Indian Ocean cruise (fromSingapore to Colombo, Ceylon), in late 1963 (http://tevega.stanford.edu/cruises.htm). Thesecollections were deposited at the U.S. NationalMuseum of Natural History, and some weredesignated as types of new fish species describedsubsequently (i.e., Melichthys indicus Randall &Klausewitz, 1973, Gymnothorax pseudoherreiBöhlke, 2000, and Helcogramma lacuna Williams& Howe, 2003). In February 1979, Randall(Bishop Museum, Hawaii) and Allen (WesternAustralian Museum) made a few collections andphotographed reef fishes around Phuket and theSimilan Islands (see Allen et al., 2005). Although

Phuket mar. biol. Cent. Res. Bull.32

no comprehensive list of fishes resulted from thateffort, the collected materials had been used indescribing several new species (i.e., Leptojulischrysotaenia Randall & Ferraris, 1981, Chromisflavipectoralis Randall, 1988, Pomacentrus adelusAllen, 1991, P. azuremaculatus Allen, 1991, P.polyspinus Allen, 1991, P. similis Allen, 1991,Neopomacentrus sororius Randall & Allen, 2005,Plesiops auritus Mooi, 1995, Plectorhinchusmacrospilus Satapoomin & Randall, 2000,

Ctenochaetus truncatus Randall & Clements, 2001,Scorpaenopsis possi Randall & Eschmeyer, 2001,Helcogramma lacuna Williams & Howe, 2003, andOpistognathus cyanospilotus Smith-Vaniz, 2009).The most important collections of reef fishes forthe area were made in November 1993 under thecollaborative project between the Royal OntarioMuseum (ROM), Canada and the Phuket MarineBiological Center (PMBC). From well over 35sampling stations using the ichthyocide rotenone

Figure 1. Approximate range of some zoogeographic distribution categories. A: Indo-Pacific (IP),Indo-west Pacific (IWP), West central Pacific (WCP), and West Pacific (WP); B: Indo-Australianarchipelago (IA) and Indo-Malayan Archipelago (IM); C: Northern Indian Ocean (NIO), Eastern IndianOcean (EIO), and Eastern extremity of Indian Ocean (EEIO).

33The fishes of southwestern Thailand

around Phuket and adjacent islands, nearly eighteenthousand specimens representing about 570 speciesin 75 families of fishes were collected anddocumented. It was also the first time thatsignificant information on small cryptobenthic reeffishes for the area was obtained. The collectedmaterials have been used in the descriptions of 13new species to date (including 7 species in Gobiidaeand 1 in each of the following families:Apogonidae,Serranidae, Pomacentridae, Opistognathidae,Tripterygiidae, and Soleidae), with several otherdescriptions underway. Satapoomin (1993)established a first comprehensive checklist of coralreef fishes of the west coast of Thailand bycompiling information from both available literature(dated back to 1950) and the author’s surveyresults carried out during 1990–1992. The listrevealed a total of 597 species belonging to 66families of reef fishes in the area. Allen et al. (2005)provided an extensive list, consisting of 764 speciesin 70 families, of coral reef fishes being recordedand/or known for the area. The most recent list ofcoral reef fishes, with 888 species in 85 families,was published by Satapoomin (2007).

There are a limited number of studies onichthyofauna of either mangroves or seagrass bedsin the Andaman Sea coast of Thailand, most ofwhich were confined to particular sampling sites(e.g., Boonruang et al., 1994; Monkolprasit, 1994;Janekitkarn and Monkolprasit, 1994). Satapoominand Poovachiranon (1997) provided the firstcomprehensive list of the fish fauna from severalmajor mangrove and seagrass bed sites along thecoast of southwestern Thailand. That studyrevealed a total of 280 species representing 75families of fishes, which included 232 species (69families) and 149 species (51 families) found inmangroves and seagrass beds, respectively. In2002, a study by Fish Team of the Trang Project(2002) provided a list, together with illustrations,of 134 species in 49 families of fishes inhabiting amangrove estuary at Sikao, southwestern Thailand.Under the JSPS (Japan Society for the Promotionof Science) Multilateral Project on Coastal MarineScience, Matsuura and Kimura (2005) publisheda guide book with taxonomic accounts andillustrations of 128 species in 46 families of fishesinhabiting mangroves, seagrass beds and

surrounding area of Libong Island, Trang Province.The most recent list of the fish fauna in the seagrassbeds of the Andaman Sea coast of Thailand, with230 species in 62 families, was published bySatapoomin (2006).

The first publication on demersal and/orpelagic fishes of the Andaman Sea coast ofThailand was published by Lohakarn (1968). Thisincluded taxonomic accounts of 109 species in 57families. Fishes collected by bottom trawls duringthe 5th Thai-Danish Expedition to the AndamanSea in 1966 (the first Thai-Danish marineexpedition) were reported by Boonyanate andHylleberg (1993) with a total records of 80 speciesin 41 families. Other early studies on demersalfishes in the area were mostly focused on fisheriesaspects and, in most cases, faunal compositionwas reported mainly as higher taxonomic ranksrather than individual species (e.g., Pokapunt andSinoda, 1978; Boonragsa, 1988; Chantawong,1996). Some studies on experimental operationsof certain types of fishing gears reported not morethan 50 fish species each (e.g., Wongratana,1982b, 1983; Ruengpan, 1988). A morecomprehensive list of demersal fishes collected byotterboard trawl nets and deep sea shrimp trapsalong the Andaman Sea coast of Thailand at depthsof about 30–420 m was published by Wongratana(1982a), with 256 species in 91 families. Also,Pokapunt and Tantivala (1987) reported 182species of demersal fishes (in 66 families) in thearea from surveys using otterboard trawls at waterdepths less than 90 m. Deep-sea fishes of theAndaman Sea were poorly known. The results ofsurveys during 1975 and 1976 on fishes inhabitingthe continental shelf within the depth range of 200–550 m was reported by Pokapunt et al. (1983),with taxonomic accounts of 31 species in 20families. Under the Scientific CooperationProgramme between Denmark and Thailand during1996–2000, the Biodiversity of the Andaman SeaShelf (BIOSHELF) Project was undertaken by thestaff of the PMBC together with their Danishcounterparts. The benthic fauna, including fishes,was sampled at depths of 40–900 m. Preliminaryresults revealed a total of 55 families in 20 ordersof fishes, most of which were deep-water fishesthat have not yet been identified (Aungtonya et

Phuket mar. biol. Cent. Res. Bull.34

al., 2000). Under the JSPS project, acomprehensive guide book entitled “Fishes ofAndaman Sea, West Coast of Southern Thailand”covered 778 species in 106 families of fishes(Kimura et al., 2009).

The Reference Collection of the PMBC isone of the major sources of information on marinebiodiversity of theAndaman Sea coast of Thailand.The collection of fishes, in particular, was largelyestablished as a result of the FAO/DANIDASeminar on Fish Taxonomy in Southeast Asia heldat the PMBC in 1972. Apart from the AndamanSea fishes, a number of specimens obtained fromthe Gulf of Thailand, as well as from neighboringcountries (i.e., India, Malaysia, Singapore,Indonesia, Philippines, and Hong Kong) werebrought in during the seminar and subsequentlydeposited in the PMBC Reference Collection. Somedeposited material of fishes from Australia wasdonated by the Australian Museum under a pastexchange programme. One lot of fishes collectedfrom Myanmar during the first joint Myanmar-ThaiFishery Exploratory Survey in 1990 was alsodeposited in the Reference Collection(Nateewathana et al., 1983). A revised checklistof fishes in the PMBC Reference Collection waspublished by Nateewathana et al. (1983),comprising a total of 629 species in 119 families.The collection has gradually increased following anumber of subsequent researches (e.g.,Sirimontraporn et al., 1993; Satapoomin andPoovachiranon, 1997; Satapoomin, 1999, 2002b)and intensive sampling programmes (e.g., acollaborative project between ROM and PMBC in1993 and the PMBC’s past and ongoing projects,namely Coral Reef Management Project (1995–1999), Seagrass Resources Management andDugong Conservation Project (1997–2000), andAssessment of Coastal Marine Resources Project(since 2004)). According to the recent PMBC datarepository (http://pmbc.go.th/pmbc-rc/Taxonomy.php/), the most reliable records of fishes(excluding doubtful records and/ormisidentifications) deposited in the ReferenceCollection total at least 1,200 species belonging to170 families. TheAndaman Sea fishes, in particular,are well represented with a total number of 1,093species, comprising 1, 39, and 1,053 species in

Myxini, Chondrichthyes, and Actinopterygii,respectively.

2. Faunal composition and zoogeographyThe present checklist contains 1,746

species, representing 696 genera, 198 families, 40orders, and 3 classes, of marine and coastal fishesfrom theAndaman Sea coast of Thailand (AppendixA). The figure is relatively high as compared tothose reported from several other localities in theIndian Ocean. For example, 533 species werereported from the Cocos (Keeling) Islands (Allenand Smith-Vaniz, 1994), 603 from the LaccadiveArchipelago (Jones and Kumaran, 1980), 622 fromChristmas Island (Hobbs et al., 2010), 784 fromthe Chagos Archipelago (Winterbottom andAnderson, 1999), 880 from Seychelles andadjacent areas (Smith and Smith, 1969), 885 fromthe Reunion Island (Letourneur et al., 2004), 930species from Oman (Randall, 1995), 1,078 fromthe Red Sea (Golani and Bogorodsky, 2010), 1,090from the Maldives (Anderson et al., 1998), and1,800 (as an estimate) from southern Africa(Heemstra and Heemstra, 2004). There may bethe real differences in fish diversity among thoselocalities, but it should be noted that the numberof species recorded probably reflect biases ordifferences in any or all of the following aspects:sampling effort, habitat/depth representation, andgeographical coverage.

The 10 most speciose families (Gobiidae,Labridae, Pomacentridae, Serranidae,Apogonidae,Blenniidae, Carangidae, Scorpaenidae, Lutjanidaeand Chaetodontidae) collectively account for about42% of the total fish fauna. The cumulativepercentage contribution when the top 20 familiesare taken into account (Table 1) is over half thetotal (56%), suggesting that the fauna is notpredominated by only a few families but by a setof several families. This seems to reflect thediversity of habitats encompassed by the currentfaunal list. Each habitat usually contains a uniquecomposition of the predominant fish families,except for the Gobiidae, which is the most diversegroup in nearly all coastal habitats (e.g., coral reefs,seagrass beds and mangroves – Satapoomin, 2006,2007; Fish Team of the Trang Project, 2002). Thecurrent checklist, which includes species from

35The fishes of southwestern Thailand

various habitats, thus evens out the significancecontribution of certain predominant families in theanalysis. The relative abundance of these familiesis similar, although with variable ranking, to thosein studies of inshore fishes at many other Indo-Pacific localities, for example, Reunion Island(Letourneur et al., 2004), Maldive Islands (Randalland Anderson, 1993), Chagos Archipelago(Winterbottom and Anderson, 1999), northwesternAustralia (Allen, 1993), Calamianes Islands,Palawan Province (Allen, 2000), southern Taiwan(Shao et al., 2008), Yaku-shima Island, southernJapan (Motomura et al., 2010), and MarianaIslands (Myers and Donaldson, 2003). TheGobiidae, in particular, was usually the leadingfamily when it was adequately sampled. Other less

diverse families include 25 families with a rangeof species numbers of 10–19 species, 27 familieswith 5–9 species, 59 families with 2–4 species,and the remaining 66 families each with singlespecies.

With respect to habitat association, the fishfauna consists overwhelmingly of reef fishes,which comprise about 56% (983 species) of thetotal species. The coastal species from mangrovesand seagrass beds represent about 17% (296species) and 14% (246 species) of the total fauna,respectively. The other major component, whichis loosely categorized to accommodate fishesinhabiting a variety of offshore habitats (spanningpelagic to benthic zones at various depths),comprises about 55% (971 species) of the total

Table 1. Twenty most speciose families of fishes of southwestern Thailand, the Andaman Sea.

Family Number of genera Number of species % of total fish fauna

Gobiidae 62 227 13.0Labridae 30 78 4.5Pomacentridae 18 71 4.1Serranidae 17 61 3.5Apogonidae 14 60 3.4Blenniidae 21 52 3.0Carangidae 20 52 3.0Scorpaenidae 23 49 2.8Lutjanidae 10 39 2.2Chaetodontidae 7 37 2.1Acanthuridae 5 32 1.8Muraenidae 7 32 1.8Syngnathidae 11 26 1.5Carcharhinidae 7 25 1.4Nemipteridae 3 25 1.4Engraulidae 5 24 1.4Lethrinidae 5 22 1.3Leiognathidae 8 21 1.2Tetraodontidae 8 21 1.2Scaridae 6 20 1.1

Subtotal 287 974 55.8

Other 176 families 409 772 44.2

Total 696 1,746 100.0

Phuket mar. biol. Cent. Res. Bull.36

fauna. The figures accounting for the proportionof species shared between habitats varyconsiderably and are relatively low, i.e., 42%between mangroves and seagrass beds, 20.5%between offshore habitats and coral reefs, 16.8%between offshore habitats and seagrass beds,15.6% between offshore habitats and mangroves,9.4% between coral reefs and seagrass beds, and5.3% between coral reefs and mangroves. Thespecies shared among the 4 biotopes account foronly 2.3% of the total species recorded. Speciesshared between or among habitats, however,comprise a number of species which are stronglydependent on more than a single habitat type, andoften exhibit a segregated habitat-use pattern withrespect to their different life-history stages (e.g.,Mumby et al., 2004; Shibuno et al., 2008). Thisseems to suggest an inter-system connection infish populations. Because unique assemblages ofspecies are restricted to each particular habitattype, while others depend upon the existence ofmultiple habitat-types through connectivity, specialattention must be paid to holistic approaches ofmulti-ecosystems management and conservationif the overall diversity of the fishes is to bemaintained. At present, there is growing concernover the science of connectivity becauseunderstanding of the linkage among ecosystemsis considered as a key to the success ofconservation and management of marine andcoastal areas, particularly when emphasizing thedesign of marine protected areas (MPAs) and MPAnetwork (Botsford et al., 2009; Sale et al., 2010).

The zoogeographic analysis of specieswith adequately known distribution ranges ispresented in Table 2. The analysis reveals that thefish fauna consists primarily of species which arewidely distributed either in the Indo-Pacific or Indo-west Pacific (50.7%). Approximately 17.1% of thefauna are found within portions of the westernPacific and northeastern Indian Ocean (i.e., NIO–WCP, NIO–WP, NIO–IA, NIO–IM, EIO–WCP,EIO–WP, EIO–IA, and EIO–IM). Similarly 15.5%of species have a distribution range in the westernPacific or portions of this region (i.e., WCP, WP,IA, and IM). This clearly confirms that easternmost boundary of the Indian Ocean including theAndaman Sea overlaps the biogeographic regionof the West Pacific. About 12% of the fauna has a

primarily Indian Ocean range, with lesser values(Table 3) for either the fauna confined to the easternextremity of the Indian Ocean (about 1%) or thefauna restricted only to the Andaman Sea province(1.7%). The remaining species (4.9%) have pan-oceanic ranges, including circumtropical andcosmopolitan distributions. The presence of faunahaving both an Indian Ocean distribution range andalso a western Pacific distribution range in the areacan be considered as compelling evidence of adistribution confluence of both Indian Ocean andPacific Ocean faunas in the region. The boundarywhere sympatry of the eastern (= Pacific Ocean)and western (= Indian Ocean) populations is mostevident seems to encompass Christmas Island andthe Java–Bali region (see, Randall, 1998; Allen andAdrim, 2003; Hobbs et al., 2010). The stronginfluence of the Indian Ocean fauna in the areacan be further highlighted by considering thenumber of sibling species that involve Indian andPacific Ocean species pairs, and some geographicvariants that contain different Indian and PacificOcean colour morphs. While inconsistentconsiderations among ichthyologists overtaxonomic status (i.e., variants, subspecies, or fullspecies) of the latter exist, concrete morphologicaland/or genetic evidence is required to ascertaintheir taxonomic status (Randall, 1998; Hastingsand Springer, 2009). As exemplified in Table 4,which includes 68 sibling species-pairs (in 17families) and 18 species (in 10 families) possessinggeographic variation in colour morphs, theAndaman Sea fauna is predominantly representedby the Indian Ocean forms.

Since establishment of the PMBCReference Collection about 25 years ago, a greatdeal of knowledge has accumulated and well over190 new species (see also Aungtonya et al., 2006)have been described from the Andaman Sea. Atpresent, the Reference Collection maintains typematerials of 21 fish species, including 9 holotypesand 27 lots of paratypes (Table 5). Reliable recordsof Andaman Sea fishes currently deposited at theReference Collection account for about 63% ofthe total fish fauna. The fish collection thus stilldoes not include a comprehensive representationof theAndaman Sea fish fauna. In order to maintainits significant role in supporting biodiversityresearch of the ichthyofauna of this particular

37The fishes of southwestern Thailand

region, the fish collection needs to be upgradedby additional sampling so that the missing 27% ofspecies can be made available as voucher orreference materials for future researchers. Alsoworth mentioning here are the figures reflectingbiotope representation of the fishes catalogued inthe Reference Collection, which vary considerably,i.e., 64% of total offshore fishes and 65% of thereef-associated fishes, but 81% of the mangrovefishes and 88% of the seagrass fishes. Thesefigures can form a basis for targeting futuresampling programs for the Reference Collection.Small and cryptic reef fishes and particularly thoseinhabiting the deep-reef habitats are in need ofextensive sampling. For fishes dwelling amongvarious offshore habitats, deep demersal,benthopelagic, and mesopelagic fishes are amongthe groups that have not been adequately sampled.Deep sea exploratory research programs aregenerally costly and particularly require specifictechniques, equipment as well as taxonomicexpertise. The establishment of such programs islikely to require international collaborative effortsand funding.

Although the present checklist providesthe most extensive species records for the area, itshould be considered as a provisional rather thana complete faunal list. Also, there are a number ofidentifications that still need verification throughexamination of voucher specimens. Furthermore,the sampling efforts for several fish groups and/or in certain habitats remain deficient. The presentprovisional checklist, however, can at least helpto draw the attention of ichthyologists who areinterested in fish fauna of the Andaman Sea towhere there are knowledge gaps. Also, it can serveas a baseline for a comprehensive taxonomicreview and/or annotated checklist of the AndamanSea fishes. It is hoped that, joint research orcollaborative efforts will be possible in the nearfuture to accomplish such goals. While the currentlimited effort and interest, especially with respectto taxonomy, has slowed down the progress inthis research area, a concomitant obstacleregarding decreasing personnel trained inichthyology in Thailand seems to be more criticaland needs urgent action.

Table 2. Zoogeographic analysis of the fish fauna of southwestern Thailand, the Andaman Sea.

Distribution No. species %

Indo-Pacific (IP) 417 24.5Indo-West Pacific (IWP) 448 26.3West central Pacific (WCP) 27 1.6West Pacific (WP) 137 8.0Indo-Australian Archipelago (IA) 48 2.8Indo-Malayan Archipelago (IM) 53 3.1Indian Ocean (IO) 99 5.8Northern Indian Ocean (NIO) 10 0.6Eastern Indian Ocean (EIO) 46 2.7Northern Indian Ocean to West central Pacific (NIO–WCP) 2 0.1Northern Indian Ocean to West Pacific (NIO–WP) 37 2.2Northern Indian Ocean to Indo-Australian Archipelago (NIO–IA) 21 1.2Northern Indian Ocean to Indo-Malayan Archipelago (NIO–IM) 29 1.7Eastern Indian Ocean to West central Pacific (EIO–WCP) 12 0.7Eastern Indian Ocean to West Pacific (EIO–WP) 112 6.6Eastern Indian Ocean to Indo-Australian Archipelago (EIO–IA) 28 1.6Eastern Indian Ocean Indo-Malayan Archipelago (EIO–IM) 50 3.0Eastern extremity of Indian Ocean (EEIO) 15 0.9Andaman Sea (AS) 28 1.7Circumtropical (CIR) 38 2.2Cosmopolitan (COS) 46 2.7

Phuket mar. biol. Cent. Res. Bull.38

Table 3. List of fishes currently known to have restricted distribution in the Andaman Sea and adjacentareas: A) Andaman Sea endemics, including the species ranges southward to western Sumatra; B) Thespecies found in Andaman Sea and some other localities along eastern edge of the eastern Indian Ocean(= eastern extremity of Indian Ocean (EEIO))

Family: Species Distribution

A)Andaman Sea endemicsMyxinidae:Eptatretus indrambaryai Wongratana, 1983 Andaman SeaProscylliidae:Proscyllium magnificum Last & Vongpanich, 2004 Andaman SeaRajidae:Anacanthobatis sp.1 Andaman SeaAcropomatidae:Acropoma argentistigma Okamoto & Ida, 2002 Andaman SeaAploactinidae:Cocotropus echinatus (Cantor, 1850) Andaman SeaApogonidae:Apogon sp.1 West coast of ThailandArchamia ataenia Randall & Satapoomin, 1999 Andaman Sea–W. SumatraBlenniidae:Blenniella leopardus (Fowler, 1904) Andaman Sea–W. SumatraMeiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001 Andaman Sea–W. SumatraBythitidae:Alionematichthys phuketensis Møller & Schwarzhans, 2008 West coast of ThailandGobiidae:Callogobius andamanensis Menon & Chatterjee, 1974 Andaman SeaScartelaos cantoris (Day, 1871) Andaman SeaTrimma fucatum Winterbottom & Southcott, 2007 West coast of ThailandTrimma sanguinellus Winterbottom & Southcott, 2007 Andaman Sea–W. SumatraHaemulidae:Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000 Andaman SeaPlectorhinchus macrospilus Satapoomin & Randall, 2000 West coast of ThailandPomadasys andamanensis McKay & Satapoomin, 1994 West coast of ThailandLabridae:Cirrhilabrus joanallenae Allen, 2000 Andaman Sea–W. SumatraHalichoeres kallochroma (Bleeker, 1853) Andaman Sea–W. SumatraMonacanthidae:Paraluteres sp.1 Andaman SeaOphichthidae:Allips concolor McCosker, 1972 Andaman SeaPhallostethidae:Phenacostethus posthon Roberts, 1971 Andaman SeaPlesiopidae:Plesiops thysanopterus Mooi, 1995 N. Sumatra (Andaman Sea)Pomacentridae:Chromis xouthos Allen & Erdmann, 2005 Andaman Sea–W. SumatraPomacentrus polyspinus Allen, 1991 West coast of ThailandSerranidae:Cephalopholis polyspila Randall & Satapoomin, 2000 Andaman Sea–W. SumatraMeganthias filiferus Randall & Heemstra, 2007 Andaman Sea

39The fishes of southwestern Thailand

Table 3. (continued)

Family: Species Distribution

Soleidae:Aseraggodes satapoomini Randall & Desoutter-Meniger, 2007 West coast of ThailandSternoptychidae:Polyipnus asper Harold, 1994 Andaman SeaTripterygiidae:Helcogramma lacuna Williams & Howe, 2003 West coast of Thailand

B) Limited to eastern extremity of Indian Ocean (EEIO)Urolophidae:Urolophus javanicus (Martens, 1864) Andaman Sea, Java Sea (W. Java)Blenniidae:Ecsenius lubbocki Springer, 1988 Andaman Sea, BaliEcsenius paroculus Springer, 1988 Andaman Sea., W. Sumatra, Java

Sea (Seribu Is.& N. Java)Praealticus triangulus (Chapman, 1951) Andaman Sea, Java Sea (N. Java)Labridae:Cirrhilabrus cyanopleura (Bleeker, 1851) Andaman Sea, W. Sumatra,

Christmas Is., Java Sea (Pulau Putri& N. Java), Bali

Malacanthidae:Hoplolatilus luteus Allen & Kuiter, 1989 Andaman Sea, Bali, FloresOpistognathidae:Opistognathus cyanospilotus Smith-Vaniz, 2009 Andaman Sea, BaliPolynemidae:Filimanus perplexa Feltes, 1991 Andaman Sea, W. Sumatra,

S. Java, BaliPomacentridae:Pomacentrus alleni Burgess, 1981 Andaman Sea, Christmas Is.,

Java Sea (Sunda Strait &Pulau Putri)

Pomacentrus azuremaculatus Allen, 1991 Andaman Sea, Java Sea (Seribu Is.)Pomacentrus xanthosternus Allen, 1991 Andaman Sea, W. Sumatra,

Java Sea (Seribu Is.), BaliPseudochromidae:Pseudochromis andamanensis Lubbock, 1980 Andaman Sea, W. Sumatra,

Bali, Timor Sea (Ashmore Reef &Cartier Is., N. Australia)

Scaridae:Chlorurus troschelii (Bleeker, 1853) Andaman Sea, Java Sea (Seribu Is.)

Scarus maculipinna Westneat, Satapoomin & Randall, 2007 Andaman Sea, W. Sumatra, JavaSea (Pulau Putri)

Siganidae:Siganus magnificus (Burgess, 1977) Andaman Sea, W. Sumatra, Java

Sea (W. Java)

Phuket mar. biol. Cent. Res. Bull.40

Table 4. Examples of Indian and Pacific Ocean species pairs and geographic variants that the AndamanSea fishes represented by the Indian Ocean forms.

A) Sibling species: Andaman Sea fish (vs. its Pacific sibling)

Acanthuridae PinguipedidaeAcanthurus tristis (A. pyroferus) Parapercis maculata (P. pulchella)Ctenochaetus truncatus (C. strigosus) PomacanthidaeNaso elegans (N. lituratus) Centropyge eibli (C. vroliki)Zebrasoma desjardinii (Z. veliferum) Genicanthus caudovittatus (G. melanospilos)Apogonidae PomacentridaeArchamia ataenia (A. zosterophora) Amphiprion akallopisos (A. sandarcinos)Blenniidae Amphiprion ephippium (A. melanopus)Aspidontus tractus (A. taeniatus) Amphiprion sebae (A. polymnus)Atrosalarias fuscus (A. holomelas) Chromis dimidiata (C. iomelas)Blenniella cyanostigma (B. caudolineata) Chromis opercularis (C. xanthura)Blenniella leopardus (B. interrupta) Dascyllus carneus (D. reticulatus)Meiacanthus urostigma (M. ditrema) Neopomacentrus sororius (N. azysron)Plagiotremus phenax (P. laudandus) Pomacentrus similis (P. coelestis)Chaetodontidae Pomacentrus xanthosternus (P. simsiang)Chaetodon andamanensis (C. plebeius) PtereleotridaeChaetodon triangulum (C. baronessa) Ptereleotris arabica (P. hanae)Chaetodon falcula (C. ulietensis) ScaridaeChaetodon gardineri (C. selene) Chlorurus capistratoides (C. japanensis)Chaetodon interruptus (C. unimaculatus) Chlorurus rhakoura (C. oedema)Chaetodon madagaskariensis (C. mertensii) Chlorurus sordidus (C. spilurus)Chaetodon melannotus (C. ocellicaudus) Chlorurus strongylocephalus (C. microrhinos)Chaetodon trifasciatus (C. lunulatus) Chlorurus troschelii (C. bleekeri)Hemitaurichthys zoster (H. polylepis) Hipposcarus harid (H. longiceps)Heniochus pleurotaenia (H. varius) Scarus maculipinna (S. flavipectoralis)Gobiidae Scarus russelii (S. schlegeli)Amblygobius semicinctus (A. phalaena) Scarus scaber (S. oviceps)Trimma flammeum (T. macrophthalma) Scarus viridifucatus (S. spinus)Labridae ScorpaenidaeAnampses lineatus (A. melanurus) Pterois miles (P. volitans)Cirrhilabrus cyanopleura (C. lyukyuensis) SerranidaeCoris cuvieri (C. gaimard) Cephalopholis polyspila (C. microprion)Bodianus diana (B. dictynna) Pogonoperca ocellata (P. punctata)Gomphosus caeruleus (G. varius) Pseudanthias ignitus (P. dispar)Halichoeres cosmetus (H. claudia) Rabualichthys stigmaticus (R. altipinnis)Halichoeres kallochroma (H. leucurus) SiganidaeHalichoeres leucoxanthus (H. chrysus) Siganus puelloides (S. puellus)Halichoeres vrolokii (H. melanurus) Siganus stellatus (S. punctatus)Halichoeres zeylonicus (H. hartzfeldii) SyngnathidaeParacheilinus mccoskeri (P. carpenteri) Doryramphus multiannulatus (D. pessuliferus)Pseudojuloides kaleidos (P. cerasinus) TetraodontidaeMullidae Canthigaster smithae (C. epilampra)Parupeneus trifasciatus (P. crassilabris)

41The fishes of southwestern Thailand

B) Geographic variants: Andaman Sea fish possessing Indian Ocean colour morph

Dasyatidae Nemipteridae*(4)Dasyatis kuhlii *(6, 7)Scolopsis bilineatusAcanthuridae Scolopsis vosmeri*(7)Paracanthurus hepatus PomacanthidaeBalistidae *(7, 8)Pomacanthus imperator*(5, 7)Balistapus undulatus *(5, 7)Pygoplites diacanthusCaesionidae PomacentridaeCaesio cuning *(2)Chrysiptera rollandiLabridae *(7)Neoglyphidodon nigroris*(9)Epibulus insidiator *(6)Pomacentrus chrysurus*(5, 7)Halichoeres hortulanus *(2, 8)Pomacentrus pavo*(5)Labroides dimidiatus Serranidae*(7)Thalassoma jansenii *(3)Cephalopholis urodetaLutjanidae*(1, 7)Lutjanus russelli

*Note: Recognition of allopatric colour morphs in certain species can be referred to (1) Allen andTalbot, 1985; (2) Allen, 1991; (3) Randall and Heemstra, 1991; (4) Randall, 1995; (5) Randall, 1998;(6) Satapoomin, 2000; (7) Kuiter and Tonozuka, 2001; (8) Lieske and Myers, 2001; and (9) Carlson etal., 2008.

Table 4. (continued)

Table 5. List of fishes for which their type materials are deposited at the Reference Collection of thePhuket Marine Biological Center. Number in bracket denotes the number of catalogued lots for paratypes.

Family: Species Type materials

Apogonidae:Archamia ataenia Randall & Satapoomin, 1999 Paratypes (1)Blenniidae:Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001 Paratypes (2)Gobiidae:Cryptocentrus crocatus Wongratana, 1975 Holotype

[valid as Myersina crocatus (Wongratana, 1975)]Gladiogobius rex Shibukawa & Allen, 2007 Holotype, Paratypes (3)Myersina adonis Shibukawa & Satapoomin, 2006 Holotype, Paratypes (1)Periophthalmus walailakae Darumas & Tantichodok, 2002 Paratypes (2)Trimma fucatum Winterbottom & Southcott, 2007 Paratypes (1)Trimma sanguinellus Winterbottom & Southcott, 2007 Paratypes (1)Haemulidae:Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000 Paratypes (2)Plectorhinchus macrospilus Satapoomin & Randall, 2000 Paratypes (1)Pomadasys andamanensis McKay & Satapoomin, 1994 HolotypeNemipteridae:Nemipterus aurora Russell, 1993 Paratypes (1)Nemipterus thosaporni Russell, 1991 Paratypes (1)

Phuket mar. biol. Cent. Res. Bull.42

Table 5. (continued)

Family: Species Type materials

Opistognathidae:Opistognathus variabilis Smith-Vaniz, 2009 Paratypes (2)Proscylliidae:Proscyllium magnificum Last & Vongpanich, 2004 Holotype, Paratypes (3)Pomacentridae:Pomacentrus azuremaculatus Allen, 1991 HolotypePomacentrus polyspinus Allen, 1991 HolotypePomacentrus similis Allen, 1991 Paratypes (1)Scaridae:Scarus maculipinna Westneat, Satapoomin & Randall, 2007 Holotype, Paratypes (1)Serranidae:Cephalopholis polyspila Randall & Satapoomin, 2000 Paratypes (4)Meganthias filiferus Randall & Heemstra, 2007 Holotype

ACKNOWLEDGEMENTS

This paper was initially prepared as acontribution to the concluding conference of the10-year JSPS Multilateral Core UniversityProgramme on “Coastal Marine Science” in Tokyoduring 26–29 October 2010, but I finally couldnot make the trip. It was then presented as a plenarytalk at the seminar on “Biodiversity of SouthernThailand in Focus: Status and Research Direction”at Walailak University, Nakorn Sri Thammarat,Thailand, during 10–12 November 2010. I amgrateful for the generous support made availableby the JSPS for me to be able to join several fishworkshops conducted over the years, whichultimately gave me a wonderful opportunity to learnmore about the fish faunas of SE Asia. Special

thanks are due to Dr. Keiichi Matsuura [KM] ofthe National Museum of Nature and Science,Tokyo, for his considerable effort in leading thefish diversity project under the JSPS multilateralprogramme and encouraging me to write up thispaper. I am very grateful to Dr. RichardWinterbottom [RW] of the Royal Ontario Museumfor sharing the research outcomes from the 1993expedition to Phuket and inspiring my interest insmall cryptobenthic reef fishes. Both KM and RWkindly gave their time for reviewing and improvingthe first draft of this manuscript. Several past andongoing projects at PMBC which contribute to theinformation and knowledge on the Andaman Seafish fauna have been continually supported by theThai government budget. I am also grateful to themany staff members of the PMBC for theirassistance in the field.

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Sale, P.E., H. Van Lavieren, M.C.A. Lagman, J. Atema, M. Butler, C. Fauvelot, J.D. Hogan, G.P. Jones,K.C. Lindeman, C.B. Paris, R. Steneck and H.L. Stewart. 2010. Preserving Reef Connectivity:A Handbook for Marine Protected Area Managers. Connectivity Working Group, Coral ReefTargeted Research and Capacity Building for Management Program. United Nations University– Institute for Water, Environment and Health (UNU–INWEH). 79 p.

Satapoomin, U. 1993. Updated list of reef fishes and their distribution along the west coast of Thailand,Andaman Sea. Phuket mar. biol. Cent. Spec. Publ. 12: 67–91.

Satapoomin, U. 1999. A survey of fish fauna at the Cape Panwa reef, southeastern Phuket. Phuketmar. biol. Cent. Res. Bull. 62: 9–31.

Phuket mar. biol. Cent. Res. Bull.46

Satapoomin, U. 2000. A preliminary checklist of coral reef fishes of the Gulf of Thailand, South ChinaSea. Raffles Bull. Zool. 48(1): 31–53.

Satapoomin, U. 2002a. Comparative study of reef fish fauna in Thai waters: the Gulf of Thailandversus the Andaman Sea. Phuket Marine Biological Center, Technical Paper No. 1/2002. 41p.

Satapoomin, U. 2002b. Patterns of fish assemblages on coral reefs of the Adang-Rawi Islands, theAndaman Sea, with comment on management implications for coral reef reserves. Nat. Hist.Bull. Siam Soc. 50(1): 25–55.

Satapoomin, U. 2006. Fishes. In: A Guide to Fauna of Seagrass Beds in the Andaman Sea. Departmentof Marine and Coastal Resources, Ministry of Natural Resources and Environment, Thailand.pp. 97–229. (bilingual)

Satapoomin, U. 2007. A Guide to Reef Fishes of the Andaman Sea, Thailand. Phuket Marine BiologicalCenter, Department of Marine and Coastal Resources. 231 p. (in Thai)

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Manuscript received: 10 January 2011Accepted: 1 August 2011

47The fishes of southwestern Thailand

Appendix A

Checklist of marine fishes of southwesternThailand, the Andaman Sea

Abbreviations:1. Habitat types – Cr (coral reefs); Mg (mangroves); Sg

(seagrass beds); OShr (other coastal and offshore areas)2. Distribution categories – IP (Indo-Pacific); IWP (Indo-

West Pacific); WCP (west central Pacific); WP (westPacific); IA (Indo-Australian Archipelago); IM (Indo-Malayan Archipelago); IO (Indian Ocean); NIO (northernIndian Ocean); EIO (eastern Indian Ocean); EEIO (easternextremity of Indian Ocean); AS (Andaman Sea); CIR(circumtropical); COS (cosmopolitan) [see methodssection for definitions]

3. Voucher specimens – PMBC (denotes availability ofvoucher specimens at the Reference Collection of thePhuket Marine Biological Center)

CLASS MYXINIOrder MyxiniformesMyxinidaeEptatretus indrambaryai Wongratana, 1983: OShr; [AS];

PMBCCLASS CHONDRICHTHYESOrder ChimaeriformesChimaeridaeHydrolagus sp.: OShr; PMBCOrder OrectolobiformesParascylliidaeParascyllium sp.: OShr; PMBCHemiscylliidaeChiloscyllium griseum Müller & Henle, 1838: OShr; [NIO-

IM]; PMBCChiloscyllium hasselti Bleeker, 1852: Sg, OShr; [IM];

PMBCChiloscyllium indicum (Gmelin, 1788): OShr; [IWP];

PMBCChiloscyllium plagiosum (Bennett, 1830): OShr; [IWP]Chiloscyllium punctatum Müller & Henle, 1838: Cr, Sg,

OShr; [EIO-WP]StegostomatidaeStegostoma fasciatum (Hermann, 1793): Cr, OShr; [IP];

PMBCGinglymostomatidaeNebrius ferrugineus (Lesson, 1830): Cr, OShr; [IWP]RhincodontidaeRhincodon typus Smith, 1828: Cr, OShr; [COS]Order LamniformesAlopiidaeAlopias pelagicus Nakamura, 1935: OShr; [CIR]Alopias superciliosus (Lowe, 1840): OShr; [CIR]Alopias vulpinus (Bonnaterre, 1788): OShr; [COS]LamnidaeIsurus oxyrinchus Rafinesque, 1810: OShr; [COS]

Order CarcharhiniformesScyliorhinidaeAtelomycterus marmoratus (Bennett, 1830): OShr; [IWP]Cephaloscyllium umbratile Jordan & Fowler, 1903: OShr;

[WP]; PMBCHalaelurus buergeri (Müller & Henle, 1838): OShr; [WP]Halaelurus hispidus (Alcock, 1891): OShr; [EIO-WP];

PMBCProscylliidaeProscyllium magnificum Last & Vongpanich, 2004: OShr;

[AS]; PMBCTriakidaeIago omanensis (Norman, 1939): OShr; [IO]; PMBCMustelus manazo Bleeker, 1854: OShr; [IWP]; PMBCMustelus mosis Hemprich & Ehrenberg, 1899: OShr; [IO];

PMBCHemigaleidaeChaenogaleus macrostoma (Bleeker, 1852): OShr; [IWP]Hemigaleus microstoma Bleeker, 1852: OShr; [EIO-WP];

PMBCHemipristis elongatus (Klunzinger, 1871): OShr; [IWP];

PMBCCarcharhinidaeCarcharhinus albimarginatus (Rüppell, 1837): Cr, OShr;

[IP]Carcharhinus altimus (Springer, 1950): OShr; [COS]Carcharhinus amblyrhynchoides (Whitley, 1934): OShr;

[IWP]Carcharhinus amblyrhynchos (Bleeker, 1856): OShr; [IP]Carcharhinus amboinensis (Müller & Henle, 1839): OShr;

[IWP]Carcharhinus brachyurus (Günther, 1870): OShr; [COS]Carcharhinus brevipinna (Müller & Henle, 1839): OShr;

[CIR]Carcharhinus dussumieri (Valenciennes, 1839): OShr;

[NIO-WP]Carcharhinus falciformis (Müller & Henle, 1839): OShr;

[CIR]Carcharhinus leucas (Valenciennes, 1839): OShr; [CIR]Carcharhinus limbatus (Valenciennes, 1839): OShr; [COS]Carcharhinus longimanus (Poey, 1861): OShr; [COS]Carcharhinus macloti (Müller & Henle, 1839): OShr; [IWP];

PMBCCarcharhinus melanopterus (Quoy & Gaimard, 1824): Cr,

OShr; [IP]; PMBCCarcharhinus obscurus (LeSueur, 1818): OShr; [COS]Carcharhinus plumbeus (Nardo, 1827): OShr; [CIR];

PMBCCarcharhinus sealei (Pietschmann, 1916): OShr; [IWP]Carcharhinus sorrah (Valenciennes, 1839): OShr; [IWP]Galeocerdo cuvier (Péron & LeSueur, 1822): OShr; [CIR]Loxodon macrorhinus Müller & Henle, 1839: OShr;

[IWP]Negaprion acutidens (Rüppell, 1837): OShr; [IP]Rhizoprionodon acutus (Rüppell, 1837): OShr; [CIR];

PMBCRhizoprionodon oligolinx Springer, 1964: OShr; [IWP]

Phuket mar. biol. Cent. Res. Bull.48

Scoliodon laticuadus Müller & Henle, 1839: OShr; [IWP]Triaenodon obesus (Rüppell, 1835): Cr, OShr; [IP]SphyrnidaeEusphyra blochii (Cuvier, 1817): OShr; [NIO-WP]; PMBCSphyrna lewini (Griffith & Smith, 1834): OShr; [COS];

PMBCSphyrna mokarran (Rüppell, 1837): OShr; [CIR]Sphyrna zygaena (Linnaeus, 1758): OShr; [CIR]; PMBCOrder HexanchiformesHexanchidaeHeptranchias perlo (Bonnaterre, 1788): OShr; [COS]Order SqualiformesSqualidaeEtmopterus brachyurus Smith & Radcliffe, 1912: OShr;

[WP]Squalus acanthias Linnaeus, 1758: OShr; [COS]; PMBCSqualus mitzukurii Jordan & Snyder, 1903: OShr; [CIR];

PMBCOrder SquantiformesSquatinidaeSquatina sp.1: OShrOrder TorpediniformesNarcinidaeNarcine brunnea Annandale, 1909: OShr; [NIO-IM];

PMBCNarcine indica Henle, 1834: OShr; [WP]Narcine maculata (Shaw, 1804): OShr; [WP]; PMBCNarcine prodosalis Bessednov, 1966: OShr; [EIO-WP]Narcine timlei (Bloch & Schneider, 1801): OShr; [EIO-

WP]Narke dipterygia (Bloch & Schneider, 1801): OShr; [NIO-

WP]Temera hardwickii Gray, 1831: OShr; [IM]; PMBCOrder PristiformesPristidaePristis microdon Latham, 1794: OShr; [CIR]; PMBCOrder RajiformesRhinidaeRhina ancylostoma (Bloch & Schneider, 1801): Cr, OShr;

[IP]; PMBCRhynchobatidaeRhynchobatus palpebratus Compagno & Last, 2008: Cr,

OShr; [IA (presently known from west coast of Thailandand northern Australia)]

RhinobatidaeRhinobatos granulatus Cuvier, 1829: OShr; [NIO-IM];

PMBCRhinobatos thouin (Anonymous in Lacepède, 1798): OShr;

[IWP]Rhinobatos typus Bennett, 1830: OShr; [EIO-WP]RajidaeAnacanthobatis sp.1 : OShr, PMBCRaja sp.: OShrOrder MyliobatiformesPlesiobatidaePlesiobatis daviesi Wallace, 1967: OShr; [IP]; PMBC

UrolophidaeUrolophus javanicus (Martens, 1864): OShr; [EEIO]DasyatidaeDasyatis kuhlii (Müller & Henle, 1841): Cr, OShr; [IWP];

PMBCDasyatis zugei (Müller & Henle, 1841): OShr; [EIO-WP]Himantura fai Jordan & Seale, 1906: OShr; [IWP]Himantura gerrardi (Gray, 1851): Sg, OShr; [IP]; PMBCHimantura imbricata (Bloch & Schneider, 1801): Mg, Sg,

OShr; [IWP]; PMBCHimantura jenkinsii (Annadale, 1909): Cr, OShr; [EIO]Himantura undulata (Bleeker, 1852): OShr; [EIO-WP]Himantura walga (Müller & Henle, 1841): Sg, OShr; [EIO-

IM]; PMBCPastinachus sephen (Forsskål, 1775): Cr, OShr; [IWP]Taeniura lymma (Forsskål, 1775): Cr; [IWP]; PMBCTaeniura meyeni Müller & Henle, 1841: Cr; [IP]Urogymnosus asperrimus (Bloch & Schneider, 1801): Cr;

[IP]GymnuridaeGymnura poecilura (Shaw, 1804): Mg, OShr; [IP]; PMBCMyliobatidaeAetobatus narinari (Euphrasen, 1790): Cr, OShr; [COS]Aetomylaeus maculatus (Gray, 1832): OShr; [EIO-WP]Aetomylaeus milvus (Valenciennes, 1841): OShr; [IWP]Aetomylaeus nichofii (Bloch & Schneider, 1801): OShr;

[IWP]; PMBCAetomylaeus vespertilio (Bleeker, 1852): OShr; [IWP]Manta birostris (Donndorff, 1798): Cr, OShr; [CIR]Mobula kuhlii (Valenciennes, in Müller & Henle, 1841):

OShr; [IWP]Rhinoptera javanica Müller & Henle, 1841: OShr; [IWP];

PMBCRhinoptera sewelli Misra, 1946: OShr; [EIO]CLASS ACTINOPTERYGIIOrder ElopiformesElopidaeElops hawaiiensis Regan, 1909: Mg, Sg, OShr; [WCP];

PMBCMegalopidaeMegalops cyprinoides (Broussonet, 1782): Mg, OShr; [IP];

PMBCOrder AlbuliformesAlbulidaeAlbula glossodonta (Forsskål, 1775): OShr; [IP]Albula oligolepis Iwatsuki & Hidaka, 2008: OShr; [IO];

PMBCNotacanthidaeNotacanthus sp.: OShr; PMBCOrder AnguilliformesAnguillidaeAnguilla bicolor bicolor McClelland, 1844: Mg; [IWP];

PMBCAnguilla marmorata Quoy & Gaimard, 1824: OShr; [IP];

PMBCChlopsidae

49The fishes of southwestern Thailand

Kaupichthys sp.: CrMuraenidaeEchidna delicatula (Kaup, 1856): OShr; [EIO-WP]; PMBCEchidna nebulosa (Ahl, 1789): Cr; [IP]; PMBCEchidna polyzona (Richardson, 1844): Cr; [IP]; PMBCEnchelycore bayeri (Schultz, 1953): Cr; [IP]Gymnomuraena zebra (Shaw, 1797): Cr; [IP]; PMBCGymnothorax annulatus (Smith & Bohlke, 1997): OShr;

[IM]Gymnothorax buroensis (Bleeker, 1857): Cr; [IP]Gymnothorax chilospilus Bleeker, 1865: Cr; [IP]; PMBCGymnothorax favagineus (Bloch & Schneider, 1801): Cr;

[IWP]; PMBCGymnothorax fimbriatus (Bennett, 1832): Cr; [IP]; PMBCGymnothorax flavimarginatus (Rüppell, 1828): Cr; [IP]Gymnothorax javanicus (Bleeker, 1859): Cr; [IP]; PMBCGymnothorax margaritophorus Bleeker, 1864: Cr; [IP]Gymnothorax melatremus Schultz, 1953: Cr; [IP]Gymnothorax meleagris (Shaw & Nodder, 1795): Cr; [IP]Gymnothorax moluccensis (Bleeker, 1864): Cr; [IWP]Gymnothorax monochrous (Bleeker, 1856): Cr; [IP]Gymnothorax permistus (Smith, 1962): Cr; [IO]; PMBCGymnothorax pindae Smith, 1962: Cr; [IP]; PMBCGymnothorax pseudoherrei Böhlke, 2000: Cr; [IWP]Gymnothorax pseudothyrsoideus (Bleeker, 1852): Cr;

[NIO-WP]; PMBCGymnothorax reticularis Bloch, 1795: OShr; [IO]; PMBCGymnothorax richardsoni (Bleeker, 1852): Cr; [IWP];

PMBCGymnothorax rueppelliae (Mcclelland, 1845): Cr; [IP]Gymnothorax thyrsoideus (Richardson, 1844): Cr, OShr;

[EIO-WCP]; PMBCGymnothorax undulatus (Lacepède, 1803): Cr; [IP]; PMBCGymnothorax zonipectis Seale, 1906: Cr; [IP]; PMBCRhinomuraena quaesita Garman, 1888: Cr; [IWP]Scuticaria tigrina (Lesson, 1828): Cr; [IP]Uropterygius concolor Rüppell, 1838: Cr; [IP]; PMBCUropterygius inornatus Gosline, 1958: Cr; [WCP]Uropterygius xanthopterus Bleeker, 1859: Cr; [IP]OphichthidaeAllips concolor McCosker, 1972: Cr, Mg; [AS]Apterichthus sp.: OShr; PMBCBrachysomophis crocodilinus (Bennet, 1833): Cr, OShr;

[IP]Echelus uropterus (Temminck & Schlegel, 1846): OShr;

[IWP]; PMBCMuraenichthys gymnopterus (Bleeker, 1853): OShr; [EIO-

WP]Myrichthys colubrinus (Boddaert, 1781): Cr; [IP]; PMBCNeenchelys sp.: OShr; PMBCOphichthus evermanni Jordan & Richardson, 1909: OShr;

[WP]Ophichthus sp.: OShr; PMBCPisonodophis cancrivorus (Richardson, 1844): Mg, Sg,

OShr; [IWP]; PMBCScolecenchelys gymnota (Bleeker, 1854): Sg; [IWP]

Scolecenchelys macroptera (Bleeker, 1857): Sg, OShr;[IWP]; PMBC

Xyrias revulsus Jordan & Snyder, 1901: OShr; [IWP]ColocongridaeColoconger scholesi Chan, 1967: OShr; [IWP]; PMBCColoconger sp.: OShr; PMBCMuraenesocidaeCongresox talabon (Cuvier, 1829): OShr; [EIO-IM];

PMBCCongresox talabonoides (Bleeker, 1853): OShr; [IWP];

PMBCMuraenosox bagio (Hamilton, 1822): OShr; [IWP]Muraenosox cinereus (Forsskål, 1775): OShr; [IWP];

PMBCCongridaeAriosoma anago (Temminck & Schlegel, 1846): OShr;

[IWP]; PMBCBathycongrus guttulatus (Günther, 1887): OShr; [IP]Bathyuroconger sp.: OShr; PMBCConger cinereus Rüppell, 1828: OShr; [IP]; PMBCGnathophis sp.: OShr; PMBCGorgasia preclara Böhlke & Randall, 1981: Cr; [EIO-IA]Heteroconger hassi (Klausewitz & Eibl-Eibesfeldt, 1959):

Cr; [IP]Rhynchoconger squaliceps (Alcock, 1894): OShr; [EIO]Uroconger lepturus (Richardson, 1845): OShr; [IWP];

PMBCNettastomatidaeNettastoma sp.: OShrOrder ClupeiformesPristigasteridaeIlisha elongata (Bennett, 1830): OShr; [EIO-IM]; PMBCIlisha kampeni (Weber & Beaufort, 1913): Sg, OShr; [EIO-

IM]Ilisha megaloptera (Swainson, 1839): Mg, OShr; [EIO-

IM]; PMBCIlisha melastoma (Schneider, 1801): Mg, OShr; [EIO-WP];

PMBCIlisha sirishai Seshagiri Rao, 1975: OShr; [NIO-IM]Opisthopterus tardoore (Cuvier, 1829): Mg, OShr; [NIO-

IM]; PMBCPellona ditchela Valenciennes, 1847: Mg, OShr; [IWP];

PMBCRaconda russeliana Gray, 1831: OShr; [EIO-IM]EngraulidaeCoilia dussumieri Valenciennes, 1848: OShr; [EIO-IM];

PMBCCoilia macrognathos Bleeker, 1852: OShr; [IM]Coilia neglecta Whitehead, 1968: OShr; [EIO-IM]Enchasicholina punctifer Fowler, 1938: OShr; [IP]Encrasicholina devisi (Whitley, 1940): OShr; [IWP]; PMBCEncrasicholina heteroloba (Rüppell, 1837): OShr; [IWP];

PMBCSetipinna breviceps (Cantor, 1850): OShr; [EIO-IM]Setipinna taty (Valenciennes, 1848): Mg, OShr; [EIO-IM]Setipinna tenuifilis (Valenciennes, 1848): OShr; [EIO-WP]

Phuket mar. biol. Cent. Res. Bull.50

Stolephorus andhraensis Babu Rao, 1966: OShr; [EIO-IA]; PMBC

Stolephorus baganensis Hardenberg, 1933: Mg, OShr;[EIO-IM]

Stolephorus chinensis (Günther, 1880): Mg, OShr; [IM]Stolephorus commersonii Lacepède, 1803: Mg, Sg, OShr;

[IWP]; PMBCStolephorus dubiosus Wongratana, 1983: OShr; [EIO-IM]Stolephorus indicus (van Hasselt, 1823): Mg, Sg, OShr;

[IWP]; PMBCStolephorus insularis Hardenberg, 1933: Mg, OShr; [NIO-

WP]Stolephorus waitei Jordan & Seale, 1926: OShr; [EIO-IA];

PMBCThryssa baelama (Forster, 1775): OShr; [IWP]Thryssa dussumieiri (Valenciennes, 1848): OShr; [NIO-

IM]Thryssa hamiltonii (Gray, 1835): Mg, Sg, OShr; [NIO-

IA]; PMBCThryssa kammalensis (Bleeker, 1849): Mg, OShr; [IM]Thryssa mystax (Schneider, 1801): OShr; [NIO-IM];

PMBCThryssa setirostris (Broussonet, 1782): Mg, Sg, OShr;

[IWP]; PMBCThryssa spinidens (Jordan & Seale, 1925): OShr; [EIO]ChirocentridaeChirocentrus dorab (Forsskål, 1775): Mg, Sg, OShr; [IWP];

PMBCChirocentrus nudus Swainson, 1839: OShr; [IWP]ClupeidaeAmblygaster clupeoides Bleeker, 1849: OShr; [EIO-WP]Amblygaster leiogaster (Valenciennes, 1847): OShr; [IWP]Amblygaster sirm (Walbaum, 1792): OShr; [IWP]; PMBCAnodontostoma chacunda (Hamilton, 1822): Mg, Sg, OShr;

[NIO-WP]; PMBCAnodontostoma thailandiae Wongratana, 1983: OShr; [EIO-

IM]Dussumieria acuta Valenciennes, 1847: OShr; [NIO-IM];

PMBCDussumieria elopsoides Bleeker, 1849: Mg, OShr; [IWP]Escualosa thoracata (Valenciennes, 1847): Mg, Sg, OShr;

[NIO-IA]; PMBCHerklotsichthys quadrimaculatus (Rüppell, 1837): Cr,

OShr; [IP]; PMBCHilsa kelee (Cuvier, 1829): Mg, Sg, OShr; [IWP]Nematalosa galatheae Nelson & Rothman, 1973: OShr;

[EIO-IM]Nematalosa nasus (Bloch, 1795): OShr; [EIO-WP]Sardinella albella (Valenciennes, 1847): Mg, Sg, OShr;

[IWP]; PMBCSardinella fimbriata (Valenciennes, 1847): OShr; [EIO-IA];

PMBCSardinella gibbosa (Bleeker, 1849): OShr; [IWP]; PMBCSardinella lemura Bleeker, 1853: OShr; [WP]Spratelloides delicatulus (Bennett, 1831): Cr, OShr; [IP];

PMBC

Spratelloides gracilis (Temminck & Schlegel, 1846): Cr,Mg, Sg, OShr; [IP]

Tenualosa reevesii (Richardson, 1846): OShr; [WP]Tenualosa toli (Valenciennes, 1847): OShr; [NIO-IM];

PMBCOrder GonorynchiformesChanidaeChanos chanos (Forsskål, 1775): Cr, Mg, OShr; [IP]Order SiluriformesPlotosidaePlotosus canius Hamilton, 1822: Mg, Sg, OShr; [EIO-IA];

PMBCPlotosus lineatus (Thunberg, 1787): Cr, Mg, Sg, OShr;

[IWP]; PMBCAriidaeArius arius (Hamilton, 1822): OShr; [EIO-IM]; PMBCArius caelatus Valenciennes, 1840: Mg; [EIO-IM]Arius gagora (Hamilton, 1822): OShr; [EIO]Arius oetik Bleeker, 1846: OShr; [WP]Arius venosus Valenciennes, 1840: OShr; [EIO-IM]Nemapteryx nenga (Hamilton, 1822): OShr; [EIO-IM]Nemuta thalassinus (Rüppell, 1837): OShr; [IWP]; PMBCOsteogeneiosus militaris (Linnaeus, 1758): OShr; [EIO-

IM]Plicofollis nella Bleeker, 1846: OShr; [WP]BagridaeMystus gulio (Hamilton, 1822): Mg; [EIO-IM]; PMBCOrder ArgentiniformesAlepocephalidaeAlepocephalus bicolor Alcock, 1891: OShr; [IWP]; PMBCOrder StomiiformesSternoptychidaeArgyropelecus affinis Garman, 1899: OShr; [CIR]; PMBCArgyropelecus sladeni Regan, 1908: OShr; [CIR]; PMBCPolyipnus asper Harold, 1994: OShr; [AS]; PMBCSternoptyx obscura Garman, 1899: OShr; [IWP]; PMBCStomiidaeAstronesthes lucifer Gilbert, 1905: OShr; [IP]Chauliodus sloanei Bloch & Schneider, 1801: OShr; [COS]Eustomias sp.: OShr; PMBCOrder AteleopodiformesAteleopodidaeAteleopus indicus Alcock, 1891: OShr; [EIO]; PMBCOrder AulopiformesParaulopidaeParaulopus brevirostris (Fourmanoir, 1981): OShr; [WP]SynodontidaeHarpadon nehereus (Hamilton, 1822): OShr; [IWP]Saurida elongata (Temminck & Schlegel, 1846): OShr;

[WP]; PMBCSaurida gracilis (Quoy & Gaimard, 1824): Cr, OShr; [IP];

PMBCSaurida longimanus Norman, 1939: OShr; [NIO]; PMBCSaurida micropectoralis Shindo & Yamada, 1972: Mg,

OShr; [WP]; PMBCSaurida nebulosa Valenciennes, 1849: Cr, Mg, Sg, OShr;

[IP]; PMBC

51The fishes of southwestern Thailand

Saurida tumbil (Bloch, 1795): OShr; [IWP]; PMBCSaurida undosquamis (Richardson, 1848): Cr, OShr;

[IWP]; PMBCSynodus dermatogenys Fowler, 1912: Cr, OShr; [IP];

PMBCSynodus hoshinonis Tanaka, 1917: OShr; [IWP]; PMBCSynodus indicus (Day, 1873): OShr; [IWP]; PMBCSynodus jaculum Russell & Cressey, 1979: Cr; [IP]; PMBCSynodus rubromarmoratus Russell & Cressey, 1979: Cr;

[WP]Synodus variegatus (Lacepède, 1801): Cr, OShr; [IP];

PMBCTrachinocephalus myops (Bloch & Schneider, 1801): Mg,

Sg, OShr; [CIR]; PMBCChlorophthalmidaeChlorophthalmus corniger Alcock, 1894: OShr; [EIO-WP]AlepisauridaeAlepisaurus ferox Lowe, 1833: OShr; [COS]ParalepididaeLestolepis luetkeni (Ege, 1933): OShr; [IP]Maculisudis longipinnis Kotthaus, 1967: OShrParalepis philippinus Fowler, 1934: OShrOrder MyctophiformesNeoscopelidaeNeoscopelus macrolepidotus Jonhson, 1863: OShr; [COS]Neoscopelus microchir Matsubara, 1943: OShr; [CIR];

PMBCMyctophidaeBenthosema pterotum (Alcock, 1890): OShr; [IWP]Diaphus coeruleus (Kluzinger, 1871): OShr; [IWP]Diaphus splendidus (Brauer, 1904): OShr; [CIR]Diaphus thiollierei Fowler, 1934: OShr; [IWP]Order LampriformesLophotidaeLophotus lecepede Giorna, 1809: OShr; [CIR]; PMBCTrachipteridaeTrachipterus jacksonensis (Ramsay, 1881): OShr; [COS]Zu cristatus (Bonelli, 1820): OShr; [CIR]; PMBCOrder PolymixiiformesPolymixiidaePolymixia berndti Gilbert, 1905: OShr; [IP]Order GadiformesBregmacerotidaeBregmaceros mcclellandi Thompson, 1840: OShr; [EIO];

PMBCMacrouridaeCoelorinchus argentatus Smith & Radcliffe, 1912: OShr;

[IWP]Coelorinchus macrorhynchus Smith & Radcliffe, 1912:

OShr; [WP]Coelorinchus radcliffei Gilbert & Hubbs, 1920: OShr; [WP]Coryphaenoides asprellus (Smith & Radcliff, 1912): OShr;

[IO]; PMBCMalacocephalus laevis (Lowe, 1843): OShr; [COS]MoridaePhysiculus roseus Alcock, 1891: OShr; [EIO-WP]Carapidae

Encheliophis homei (Richardson, 1846): Sg; [IWP]; PMBCOnuxodon fowleri (Smith, 1955): Cr; [IP]Onuxodon parvibranchium (Fowler, 1927): Cr; [IP];

PMBCPyramodon ventralis Smith & Radcliffe, 1913: OShr; [IP]Order OphidiiformesOphidiidaeGlyptophidium argenteum Alcock, 1889: OShr; [IM];

PMBCGlyptophidium macropus Alcock, 1894: OShr; [NIO]Hypopleuron caninum Smith & Radcliffe, 1913: OShr;

[IWP]; PMBCNeobythites fasciatus Smith & Radcliffe, 1913: OShr; [WP]BythitidaeAlionematichthys phuketensis Møller & Schwarzhans,

2008: Cr; [AS]Alionematichthys piger (Alcock, 1890): Cr; [WCP]Alionematichthys riukiuensis (Aoyaki, 1954): Cr; [WP]Brosmophyciops pautzkei Schultz, 1960: Cr; [IWP]; PMBCDinematichthys iluocoeteoides Bleeker, 1855: Cr; [IWP];

PMBCOrder BatrachoidiformesBatrachoididaeAllenbatrachus grunniens (Linnaeus, 1758): Mg, OShr;

[NIO-IM]; PMBCOrder LophiiformesLophiidaeLophiodes lugubris (Alcock, 1894): OShr; [EIO]Lophiodes mutilus (Alcock, 1893): OShr; [IWP]; PMBCAntennariidaeAntennarius coccineus (Cuvier, 1831): Cr; [IP]Antennarius commerson (Latreille, 1804): Cr, OShr; [IP];

PMBCAntennarius hispidus (Bloch & Schneider, 1801): Cr, OShr;

[IWP]Antennarius maculatus (Desjardins, 1840): Cr; [IP]Antennarius nummifer (Cuvier, 1817): Cr; [IP]; PMBCAntennarius pictus (Shaw & Nodder, 1794): Cr; [IP]; PMBCChaunacidaeChaunax pictus Lowe, 1846: OShr; [COS]OgcocephalidaeHalieutaea coccinea Alcock, 1889: OShr; [IO]Halieutaea indica Annandale & Jenkins, 1910: OShr; [IWP]Halieutaea stellata (Vahl, 1797): OShr; [WP]; PMBCHalieutopsis micropa (Alcock, 1891): OShr; [IWP]Order MugiliformesMugilidaeChelon macrolepis (Smith, 1846): Mg; [IP]; PMBCChelon parsia (Hamilton, 1822): Mg; [EIO]Chelon subviridis (Valenciennes, 1836): Mg, Sg; [IWP];

PMBCCrenimugil crenilabris (Forsskål, 1775): Cr; [IP]Ellochelon vaigiensis (Quoy & Gaimard, 1825): Cr, Mg,

Sg; [IP]; PMBCMoolgarda cunnesius (Valenciennes, 1836): Mg, Sg; [IWP];

PMBCMoolgarda engeli (Bleeker, 1859): Mg; [IP]

Phuket mar. biol. Cent. Res. Bull.52

Moolgarda pedaraki (Valenciennes, 1836): Mg, Sg; [IWP];PMBC

Moolgarda perusii (Valenciennes, 1836): Mg, Sg; [IWP];PMBC

Moolgarda seheli (Forsskål, 1775): Cr, Mg; [IP]; PMBCOedalechilus labiosus (Valenciennes, 1836): Cr; [IP];

PMBCParamugil parmatus (Cantor, 1849): Mg; [IM]; PMBCOrder AtheriniformesPhallostethidaeNeostethus bicornis Regan, 1916: Mg; [IM]Neostethus lankesteri Regan, 1916: Mg; [IM]; PMBCPhenacostethus posthon Roberts, 1971: Mg; [AS]AtherinidaeAtherinomorus duodecimalis (Valenciennes, 1835): Cr, Mg,

Sg, OShr; [IP]; PMBCAtherinomorus lacunosus (Forster & Schneider, 1801):

OShr; [EIO-WP]; PMBCAtherinomorus pinguis (Lacepède, 1803): Sg, OShr; [IWP]Atherion elymus Jordan & Starks, 1901: Cr, OShr; [WP];

PMBCHypoatherina temminckii (Bleeker, 1853): Cr, OShr [IP];

PMBCHypoatherina valenciennei (Bleeker, 1853): Mg, Sg, OShr;

[WP]; PMBCOrder BeloniformesAndrianichthyidaeOryzias javanicus (Bleeker, 1854): Mg; [IM]; PMBCExocoetidaeCheilopogon arcticeps (Günther, 1866): OShr; [WP]Cheilopogon spilopterus (Valenciennes, 1846): OShr; [WP];

PMBCExocoetus volitans (Linnaeus, 1758): OShr; [COS]; PMBCHirundichthys speculiger (Valenciennes, 1847): OShr; [CIR]Parexocoetus brachypterus (Richardson, 1846): OShr; [IP];

PMBCHemiramphidaeDermogenys siamensis Fowler, 1934: Mg; [IM]Hemiramphus far (Forsskål, 1775): Cr, Sg, OShr; [IWP];

PMBCHyporhamphus balinensis (Bleeker, 1858): OShr; [IWP]Hyporhamphus dussumieri (Valenciennes, 1846): Sg, OShr;

[IP]; PMBCHyporhamphus limbatus (Valenciennes, 1846): Mg, Sg,

OShr; [NIO-WP]Hyporhamphus melanopterus Collette & Parin, 1978: Mg,

OShr; [IM]; PMBCHyporhamphus quoyi Valenciennes, 1847: Mg, Sg, OShr;

[IWP]; PMBCHyporhamphus unicuspis Collette & Parin, 1978: OShr;

[NIO]Zenarchopterus buffornis (Valenciennes, 1847): Mg, Sg,

OShr; [EIO-IA]; PMBCZenarchopterus dispar (Valenciennes, 1847): Mg, Sg, OShr;

[IWP]; PMBCZenarchopterus dunckeri Mohr, 1926: Mg, Sg, OShr; [WP];

PMBC

BelonidaeAblennes hians (Valenciennes, 1846): Cr, OShr; [COS];

PMBCStrongylura incisa (Valenciennes, 1846): OShr; [EIO-WP];

PMBCStrongylura leiura (Bleeker, 1850): OShr; [IWP]; PMBCStrongylura strongylura (van Hasselt, 1823): Mg, Sg, OShr;

[NIO-IA]; PMBCTylosurus acus melanotus (Bleeker, 1850): Cr, OShr; [IP]Tylosurus crocodilus crocodilus (Peron & Lesueur, 1821):

Cr, Mg, Sg, OShr; [CIR]Order CyprinodontiformesAplocheilidaeAplocheilus panchax Day, 1875: Mg; [EIO-IM]; PMBCOrder BeryciformesDiretmidaeDiretmoides veriginae Kotlyar, 1987: OShr; [EIO-WP];

PMBCMonocentridaeMonocentris japonica (Houttuyn, 1782): OShr; [IWP];

PMBCTrachichthyidaeGephyroberyx darwinii (Johnson, 1866): OShr; [COS];

PMBCBerycidaeBeryx splendens Lowe, 1834: OShr; [COS]Centroberyx druzhinini (Busakhin, 1981): OShr; [IWP];

PMBCHolocentridaeMyripristis adustus (Bleeker, 1853): Cr, OShr; [IP]Myripristis berndti (Jordan & Evermann, 1903): Cr, OShr;

[IP]; PMBCMyripristis botche Cuvier, 1829: Cr, OShr; [IWP]; PMBCMyripristis chryseres Jordan & Evermann, 1903: Cr, OShr;

[IP]Myripristis hexagona (Lacepède, 1802): Cr, OShr; [IWP];

PMBCMyripristis kuntee Cuvier, 1831: Cr, [IP]Myripristis murdjan (Forsskål, 1775): Cr; [IP]Myripristis violacea Bleeker, 1851: Cr; [IP]Myripristis vittatus Valenciennes, 1831: Cr; [IP]Neoniphon sammara (Forsskål, 1775): Cr, OShr; [IP];

PMBCOstichthys japonicus (Cuvier, 1829): OShr; [WP]; PMBCSargocentron caudimaculatum (Rüppell, 1835): Cr; [IP];

PMBCSargocentron cornutum (Bleeker, 1853): Cr; [WP]; PMBCSargocentron diadema (Lacepède, 1802): Cr; [IP]Sargocentron melanospilos (Bleeker, 1858): Cr; [IWP]Sargocentron rubrum (Forsskål, 1775): Cr, OShr; [IWP];

PMBCSargocentron spiniferum (Forsskål, 1775): Cr; [IP]Order ZeiformesGrammicolepididaeXenolepidichthys dalgleishi Gilchrist, 1922: OShr; [CIR]ZeidaeCyttopsis rosea (Lowe, 1843): OShr; [COS]; PMBC

53The fishes of southwestern Thailand

Zenopsis nebulosa (Temminck & Schlegel, 1845): OShr;[IP]

Order GasterosteiformesPegasidaeEurypegasus draconis (Linnaeus, 1766): Cr, OShr; [IP];

PMBCPegasus volitans Linnaeus, 1758: Cr, Sg, OShr; [IWP];

PMBCSolenostomidaeSolenostomus cyanopterus Bleeker, 1852: Cr; [IP]Solenostomus paradoxus (Pallas, 1770): Cr, Sg; [IWP];

PMBCSyngnathidaeBhanotia fasciolata (Dumeril, 1870): Cr, OShr; [IA];

PMBCChoeroichthys brachysoma (Bleeker, 1855): Cr; [IP];

PMBCChoeroichthys sculptus (Günther, 1870): Cr, OShr; [IP]Corythoichthys amplexus Dowson & Randall, 1906: Cr,

OShr; [IWP]Corythoichthys flavofasciatus (Rüppell, 1838): Cr; [IP]Corythoichthys haematopterus (Bleeker, 1851): Cr, OShr;

[IWP]; PMBCCorythoichthys schultzi Herald, 1953: Cr; [IP]; PMBCDoryrhamphus janssi (Herald & Randall, 1972): Cr, OShr;

[WP]; PMBCDunckerocampus multiannulatus (Regan, 1903): Cr; [IO]Halicampus grayi Kaup, 1856: OShr; [NIO-WP]; PMBCHalicampus mataafae (Jordan & Seale, 1906): Cr, OShr;

[IP]Halicampus spinirostris (Dawson & Allen, 1981): Cr;

[EIO-WP]Hippichthys cyanospilus (Bleeker, 1854): Sg, OShr; [IWP];

PMBCHippichthys heptagonus (Bleeker, 1853): Sg, OShr; [IWP];

PMBCHippichthys penicillus (Cantor, 1849): Mg, Sg, OShr; [NIO-

WP]; PMBCHippichthys spicifer (Rüppell, 1840): Mg, OShr; [IP];

PMBCHippocampus comes Cantor, 1850: Cr, Sg; [IM]; PMBCHippocampus kelloggi Jordan & Snyder, 1901: OShr;

[IWP]; PMBCHippocampus kuda Bleeker, 1852: Mg, Sg, OShr;[NIO-

WCP]; PMBCHippocampus spinosissimus Weber, 1913: OShr; [EIO-IA];

PMBCHippocampus trimaculatus Leach, 1814: OShr; [EIO-WP];

PMBCPhoxocampus belcheri (Kaup, 1856): Cr, OShr; [IWP]Syngnathoides biaculeatus (Bloch, 1785): Sg, OShr; [IWP];

PMBCTrachyrhamphus bicoarctatus (Bleeker, 1857): Cr, Sg,

OShr; [IWP]; PMBCTrachyrhamphus longirostris Kaup, 1856: OShr; [IWP];

PMBC

Trachyrhamphus serratus (Temminck & Schlegel, 1850):OShr; [EIO-WP]; PMBC

AulostomidaeAulostomus chinensis (Linnaeus, 1766): Cr; [IP]; PMBCFistulariidaeFistularia commersonii Rüppell, 1835: Cr, Sg; [IP]; PMBCFistularia petimba Lacepède, 1803: OShr; [CIR]; PMBCCentriscidaeAeoliscus strigatus (Günther, 1860): Cr, Sg, OShr; [IWP]Centriscus cristatus De Vis, 1885: OShr; [WP]Centriscus scutatus Linnaeus, 1758: Cr, Sg, OShr; [IWP];

PMBCOrder SybranchiformesSybranchidaeMacrotrema caligans (Cantor, 1849): Mg; [IM]Order ScorpaeniformesDactylopteridaeDactyloptena macracantha (Bleeker, 1854): OShr; [EIO-

WP]; PMBCDactyloptena orientalis (Cuvier, 1829): Cr, OShr; [IP];

PMBCDactyloptena perterseni (Nystrom, 1887): OShr; [IWP];

PMBCScorpaenidaeAblabys taenianotus (Cuvier, 1829): Cr; [EIO-WP]Apistus carinatus (Bloch & Schneider, 1801): OShr; [IWP]Choridactylus multibarbus Richardson, 1848: OShr; [IWP]Dendrochirus biocellatus (Fowler, 1938): Cr; [IP]Dendrochirus brachypterus (Cuvier, 1829): Cr, OShr; [IP];

PMBCDendrochirus zebra (Cuvier, 1829): Cr, Sg, OShr; [IWP];

PMBCInimicus cuvieri (Gray, 1832): Cr, OShr; [IM]Lioscorpius longiceps Günther, 1880: OShr; [WP]Minous coccineus Alcock, 1890: OShr; [IWP]Minous inermis Alcock, 1889: OShr; [IO]Minous monodactylus (Bloch & Schneider, 1801): OShr;

[IWP]; PMBCMinous trachycephalus (Bleeker, 1854): OShr; [EIO-WP]Neomerinthe amplisquamiceps (Fowler, 1938): OShr; [WP]Paracentropogon longispinis (Bleeker, 1829): Cr, OShr;

[EIO-WP]Parapterois heterurus (Bleeker, 1856): OShr; [IWP]Parascorpaena aurita (Rüppell, 1838): Cr, OShr; [IWP];

PMBCParascorpaena mossambica (Peters, 1855): Cr; [IP]Pontinus macrocephalus (Sauvage, 1882): OShr; [WP];

PMBCPterois antennata (Bloch, 1787): Cr, OShr; [IP]; PMBCPterois miles (Bennett, 1828): Cr, Sg, OShr; [IO]; PMBCPterois mombasae (Smith, 1957): Cr, OShr; [IO]Pterois radiata Cuvier, 1829: Cr, OShr; [IP]; PMBCPterois russellii Bennett, 1831: Sg, OShr; [IWP]; PMBCRhinopias eschmeyeri Conde, 1977: Cr; [IWP]Rhinopias frondosa (Günther, 1891): Cr, OShr; [IWP]Richardsonichthys leucogaster (Richardson, 1848): OShr;

[IWP]

Phuket mar. biol. Cent. Res. Bull.54

Scorpaenodes albaiensis (Evermann & Seale, 1907): Cr;[IWP]; PMBC

Scorpaenodes guamensis (Quoy & Gaimard, 1824): Cr,OShr; [IP]; PMBC

Scorpaenodes hirsutus (Smith, 1957): Cr; [IP]Scorpaenodes minor (Smith, 1958): Cr; [IP]Scorpaenodes parvipinnis (Garrett, 1863): Cr, OShr; [IP];

PMBCScorpaenodes scaber (Ramsay & Ogilby, 1886): Cr; [EIO-

WCP]Scorpaenodes varipinnis Smith, 1957: Cr; [IWP]Scorpaenopsis diabolus Cuvier, 1829: Cr, OShr; [IP];

PMBCScorpaenopsis longispina Randall & Eschmeyer, 2001: Cr;

[IO]Scorpaenopsis neglecta Heckel, 1837: Sg, OShr; [EIO-WP];

PMBCScorpaenopsis oxycephala (Bleeker, 1849): Cr, OShr;

[IWP]; PMBCScorpaenopsis possi Randall & Eschmeyer, 2001: Cr; [IP]Scorpaenopsis ramaraoi Randall & Eschmeyer, 2001: Cr,

OShr; [NIO-WP]Scorpaenopsis vesosa (Cuvier, 1829): OShr; [IWP]Sebastapistes fowleri (Pietschmann, 1934): Cr; [IP]Sebastapistes mauritiana (Cuvier, 1829): Cr, OShr; [IP]Sebastapistes strongia (Cuvier, 1829): Cr; [IP]Setarches guentheri Johnson, 1862: OShr; [CIR]Synanceia horrida (Linnaeus, 1766): Cr, OShr; [EIO-WP];

PMBCSynanceia verrucosa (Smith, 1956): Cr, OShr; [IP]Taenianotus triacanthus Lacepède, 1802: Cr; [IP]Trachicephalus uranoscopus (Bloch & Schneider, 1801):

Mg, Sg, OShr; [EIO-WP], PMBCVespicula trachinoides (Cuvier & Valenciennes, 1829): Mg,

Sg, OShr; [WP]; PMBCCaracanthidaeCaracanthus unipinna (Gray, 1831): Cr; [IWP]; PMBCAploactinidaeAcanthosphex leurynnis (Jordan & Seale, 1905): Mg, Sg,

OShr; [EIO-IM]; PMBCCocotropus echinatus (Cantor, 1850): Mg, Sg; [AS]Cocotropus larvatus Poss & Allen, 1987: OShr; [WP]Paraploactis obbesi (Weber, 1913): Cr, OShr; [WP]Sthenopus mollis Richardson, 1848: OShr; [WP]Xenaploactis cautes Poss & Eschmeyer, 1980: OShr; [IM

(presently known from AS and Gulf of Thailand)]TriglidaeLepidotrigla punctipectoralis Fowler, 1938: OShr; [IWP]Lepidotrigla spiloptera Günther, 1880: OShr; [IWP];

PMBCPeristediidaePeristedion liorhynchus Günther, 1871: OShr; [IWP];

PMBCPeristedion moluccense Bleeker, 1850: OShr; [WP]; PMBCSatyrichthys adeni (Lloyd, 1907): OShr; [IWP]; PMBCSatyrichthys sp.: OShr; PMBC

BembridaeBrachybembras sp.: OShrPlathycephalidaeCociella punctata (Cuvier, 1829): Cr, Mg, Sg, OShr; [IWP];

PMBCElates ransonneti (Steindachner, 1877): OShr; [IA]; PMBCGrammoplites scaber (Linnaeus, 1758): Mg, Sg, OShr;

[NIO-IM]; PMBCInegocia japonica (Tilesius, 1812): Mg, Sg, OShr; [EIO-

WP]; PMBCKumococius rodericensis Cuvier in Cuvier & Valenciennes,

1829: OShr; [NIO-IA]; PMBCPlatycephalus indicus (Linnaeus, 1758): Mg, Sg, OShr;

[IWP]; PMBC]Rogadias pristiger (Cuvier, 1829): OShr; [IWP]; PMBCRogadias serratus (Cuvier, 1829): OShr; [IWP]; PMBCSorsogona melanoptera (Knapp & Wongratana, 1987);

OShr; [NIO]Sorsogona tuberculata (Cuvier, 1829): OShr; [NIO-IA];

PMBCSuggrundus macracanthus (Bleeker, 1869): OShr; [EIO-

IA]; PMBCSunagocia carbunculus (Valenciennes, 1833): Mg, Sg, OShr;

[EIO-IM]; PMBCSunagocia otaitensis (Cuvier, 1829): Cr, OShr; [IP]; PMBCThrysanophrys celebicus (Bleeker, 1854): Cr; [IWP]Thrysanophrys chitonae Schultz, 1966: Cr, OShr; [IP]HoplichthyidaeHoplichthys citrinus Gilbert, 1905: OShr; [WP]Order PerciformesAmbassidaeAmbassis buruensis Bleeker, 1857: Mg; [WP]; PMBCAmbassis dussumieri Cuveir, 1828: Sg; [IWP]Ambassis gymnocephalus (Lacepède, 1802): Mg; [IWP];

PMBCAmbassis interruptus Bleeker, 1852: Mg, Sg; [IA]; PMBCAmbassis macracanthus Bleeker, 1849: Mg, Sg; [EIO-IM];

PMBCAmbassis nalua (Hamilton, 1822): Mg, Sg; [EIO-IA];

PMBCAmbassis urotaenia Bleeker, 1852: Mg, Sg; [IWP]; PMBCAmbassis vachellii Richardson, 1846: Mg, Sg; [IWP];

PMBCLatidaeLates calcalifer (Bloch, 1790): Mg; [NIO-WP]; PMBCAcropomatidaeAcropoma argentistigma Okamoto & Ida, 2002: OShr; [AS]Synagrops japonicus (Döderlein, 1883): OShr; [IP]; PMBCSynagrops malayanus Weber, 1913: OShr; [WCP]Synagrops philippinensis (Günther, 1880): OShr; [EIO-

WP]; PMBCSerranidaeAethaloperca rogaa (Forsskål, 1775): Cr, OShr; [IP];

PMBCAnyperodon leucogrammicus (Valnciennes, 1828): Cr, OShr;

[IP]; PMBC

55The fishes of southwestern Thailand

Cephalopholis argus Bloch & Schneider, 1801: Cr, OShr;[IP]; PMBC

Cephalopholis aurantia (Valenciennes, 1828): OShr; [IWP]Cephalopholis boenak (Bloch, 1790): Cr, Sg, OShr; [IWP];

PMBCCephalopholis formosa (Shaw & Nodder, 1812): Cr, OShr;

[EIO-WP]; PMBCCephalopholis leopardus (Lacepède, 1801): Cr; [IP]Cephalopholis miniata (Forsskål, 1775): Cr, OShr; [IP];

PMBCCephalopholis polyspila Randall & Satapoomin, 2000: Cr,

OShr; [AS]; PMBCCephalopholis sexmaculata (Rüppell, 1830): Cr; [IP]Cephalopholis sonnerati (Valenciennes, 1828): Cr, OShr;

[IP]; PMBCCephalopholis urodeta (Bloch & Schneider, 1801): Cr,

OShr; [IP]; PMBCChelidoperca margaritifera Weber, 1913: OShr; [WP];

PMBCCromileptes altivelis (Valenciennes, 1828): Cr, OShr; [WP];

PMBCDiploprion bifasciatum Cuvier, 1828: Cr, OShr; [EIO-WP];

PMBCEpinephelus amblycephalus (Bleeker, 1857): OShr; [WP];

PMBCEpinephelus areolatus (Forsskål, 1775): Cr, OShr; [IWP];

PMBCEpinephelus bleekeri (Vaillant, 1877): Cr, Mg, Sg, OShr;

[NIO-IA]; PMBCEpinephelus caeruleopunctatus (Bloch, 1790): Cr, OShr;

[IWP]; PMBCEpinephelus coioides (Hamilton, 1822): Cr, Mg, Sg, OShr:

[IWP]; PMBCEpinephelus epistictus (Temminck & Schlegel, 1842): OShr;

[IWP]; PMBCEpinephelus erythrurus (Valenciennes, 1828): Cr, Sg, OShr;

[NIO-IM]; PMBCEpinephelus fasciatus (Forsskål, 1775): Cr, OShr; [IP];

PMBCEpinephelus flavocaeruleus (Lacepède, 1801): Cr, OShr;

[IO]Epinephelus fuscoguttatus (Forsskål, 1775): Cr, OShr; [IP];

PMBCEpinephelus heniochus Fowler, 1904: OShr; [WP]Epinephelus hexagonatus (Bloch & Schneider, 1801): Cr;

[IP]Epinephelus lanceolatus (Bloch, 1790): Cr, OShr; [IP];

PMBCEpinephelus latifasciatus (Temminck & Schlegel, 1842):

OShr; [IWP]Epinephelus longispinis (Kner, 1864): Cr, OShr; [IO]Epinephelus macrospilos (Bleeker, 1855): Cr; [IP]; PMBCEpinephelus malabaricus (Bloch & Schneider, 1801): Cr,

Mg, OShr; [IWP]; PMBCEpinephelus merra Bloch, 1793: Cr, OShr; [IP]; PMBCEpinephelus morrhua (Valenciennes, 1833): OShr; [IP];

PMBC

Epinephelus octofasciatus Griffin, 1926: OShr; [IP]; PMBCEpinephelus ongus (Bloch, 1790): Cr, OShr; [IWP]; PMBCEpinephelus polyphekadion (Bleeker, 1849): Cr, OShr; [IP];

PMBCEpinephelus quoyanus (Valenciennes, 1830): Cr, Sg, OShr;

[WP]; PMBCEpinephelus radiatus (Day, 1867): OShr; [IWP]; PMBCEpinephelus sexfasciatus (Valenciennes, 1828): Mg, Sg,

OShr; [IA]; PMBCEpinephelus spilotoceps Schultz, 1953: Cr; [IP]Epinephelus tauvina (Forsskål, 1775): Cr, OShr; [IP];

PMBCEpinephelus tukula Morgan, 1959: Cr, OShr; [IWP]Epinephelus undulosus (Quoy & Gaimard, 1824): Cr, OShr;

[IWP]; PMBCGrammistes sexlineatus (Thunberg, 1792): Cr; [IP]Grammistops ocellatus Schultz, 1953: Cr; [IP]Liopropoma africanum (Smith, 1954): Cr; [IO]; PMBCLiopropoma susumi (Jordan & Seale, 1906): Cr; [IP];

PMBCMeganthias filiferus Randall & Heemstra, 2007: OShr; [AS];

PMBCPlectropomus areolatus (Rüppell, 1830): Cr, OShr; [IP]Plectropomus laevis (Lacepède, 1801): Cr, OShr; [IP]Plectropomus pessuliferus Fowler, 1901: Cr, OShr; [IWP]Pogonoperca ocellata Günther, 1859: Cr, OShr; [IO];

PMBCPseudanthias evansi (Smith, 1954): Cr; [IO]; PMBCPseudanthias ignitus (Randall & Lubbock, 1981): Cr; [EIO]Pseudanthias rubrizonatus (Randall, 1983): Cr; [WP]Pseudanthias squamipinnis (Peters, 1855): Cr; [IWP];

PMBCPseudogramma polyacanthum (Bleeker, 1856): Cr; [IP];

PMBCRabaulichthys stigmaticus Randall & Pyle, 1989: Cr, OShr;

[IO]Variola albimarginata Baissac, 1953: Cr, OShr; [IWP]Variola louti (Forsskål, 1775): Cr, OShr; [IP]; PMBCOstracoberycidaeOstracoberyx dorygenys Fowler, 1934: OShr; [IWP]PseudochromidaeAmsichthys knighti (Allen, 1987): Cr; [WP]Blennodesmus scapularis Günther, 1871: Cr; [IA]; PMBCPseudochromis andamanensis Lubbock, 1980: Cr; [EEIO];

PMBCPseudochromis caudalis Boulenger, 1898: Cr; [IO]Pseudochromis coccinicauda Tickell in Day, 1888: Cr;

[EIO]Pseudochromis fuscus Müller & Troschel, 1849: Cr; [EIO-

IA]; PMBCPseudochromis tapeinosoma Bleeker, 1853: Cr; [IA];

PMBCPseudoplesiops rosae Schultz, 1953: Cr; [EIO-WCP];

PMBCPlesiopidaeCalloplesiops altivelis (Steindachner, 1903); Cr; [IP]Plesiops auritus Mooi, 1995: Cr; [EIO]

Phuket mar. biol. Cent. Res. Bull.56

Plesiops caeruleolineatus Rüppell, 1835: Cr; [IP]; PMBCPlesiops corallicola Bleeker, 1853: Cr; [WCP]; PMBCOpistognathidaeOpistognathus cyanospilotus Smith-Vaniz, 2009: Cr;

[EEIO]Opistognathus nigromarginatus Rüppell, 1830: Cr, OShr;

[IWP]; PMBCOpistognathus rosenbergii (Bleeker, 1856): Cr, OShr;

[EIO-WP]; PMBCOpistognathus variabilis Smith-Vaniz, 2009: Cr; [EIO-WP];

PMBCPriacanthidaeHeteropriacanthus cruentatus (Lacepède, 1801): Cr, OShr;

[COS]; PMBCPriacanthus blochii Bleeker, 1853: OShr; [IWP]; PMBCPriacanthus hamrur (Forsskål, 1775): Cr, OShr; [IP];

PMBCPriacanthus macracanthus Cuvier, 1829: Cr, OShr; [WP];

PMBCPriacanthus sagittarius Starnes, 1988: OShr; [IWP]Priacanthus tayenus Richardson, 1846: OShr; [NIO-WP];

PMBCPristigenys niphonia (Cuvier, 1829): OShr; [IWP]ApogonidaeApogon abrogramma Fraser & Lachner, 1985: Cr; [IWP]Apogon angustatus (Smith & Radcliffe, 1911): Cr; [IP];

PMBCApogon apogonides (Bleeker, 1856): Cr; [IWP]Apogon aureus (Lacepède, 1802): Cr, OShr; [IWP]Apogon bryx Fraser, 1998: OShr; [IWP]Apogon campbelli Smith, 1949: Cr; [IO]Apogon compressus (Smith & Radcliffe, 1911): Cr; [WP];

PMBCApogon cookii Macleay, 1881: Cr; [IWP]; PMBCApogon cyanosoma Bleeker, 1853: Cr; [IWP]; PMBCApogon endekataenia Bleeker, 1852: Cr; [IA]; PMBCApogon fasciatus (White, 1790): OShr; [IWP]; PMBCApogon fleurieu (Lacepède, 1802): Cr, OShr; [IWP]Apogon frenatus Valenciennes, 1832: Cr; [IP]; PMBCApogon hyalosoma Bleeker, 1852: Mg, Sg; [WP]; PMBCApogon kallopterus Bleeker, 1856: Cr; [IP]; PMBCApogon kalosoma (Bleeker, 1852): Cr; [EIO-WP]Apogon lateralis Valenciennes, 1832: Cr, Mg, Sg; [IWP];

PMBCApogon moluccensis Valenciennes, 1832: Cr, Sg; [IA];

PMBCApogon nigrofasciatus Lachner, 1953: Cr; [IP]; PMBCApogon novaeguinae Valenciennes, 1832: Mg, Sg; [IWP];

PMBCApogon ocellicaudus Allen, Kuiter & Randall, 1994: Cr;

[IA]Apogon pleuron Fraser, 2005: Sg, OShr; [EIO-WP]; PMBCApogon poecilopterus Cuvier, 1828: OShr; [EIO-WP]Apogon semiornatus Peters, 1876: Cr; [IWP]Apogon smithi (Kotthaus, 1970): OShr; [IWP]; PMBCApogon sp.1: Cr; [AS]

Apogon striatodes Gon, 1996: OShr; [IM]; PMBCApogon striatus (Smith & Radcliffe, 1912): OShr; [NIO-

WP]; PMBCApogon taeniophorus Regan, 1908: Cr; [IP]; PMBCApogon talboti Smith, 1961: Cr; [IWP]Apogon thermalis Cuvier, 1829: Cr, Sg; [IWP]; PMBCApogon trimaculatus Cuvier, 1828: Cr, OShr; [WP]; PMBCApogon truncatus Bleeker, 1854: OShr; [IWP]; PMBCApogonichthyoides nigripinnis (Cuvier, 1828): OShr; [EIO-

IA]; PMBCApogonichthyoides sialis (Jordan & Thompson, 1914): Cr;

[EIO-WP]Apogonichthys sp.: OShrArchamia ataenia Randall & Satapoomin, 1999: Cr; [AS];

PMBCArchamia bleekeri Günther, 1859: Cr; [IWP]Archamia fucata (Cantor, 1850): Cr, Sg, OShr; [IP]; PMBCArchamia macroptera (Cuvier, 1828): Cr; [EIO-WP]Cheilodipterus artus Smith, 1961: Cr; [IP]; PMBCCheilodipterus macrodon (Lacepède, 1802): Cr; [IP];

PMBCCheilodipterus quinquelineatus Cuvier, 1828: Cr; [IP];

PMBCFoa fo (Jordan & Seale, 1906): Sg, OShr; [IWP]; PMBCFowleria aurita (Valenciennes, 1831): Cr; [IWP]; PMBCFowleria isostigma (Jordan & Seale, 1960): Cr; [IWP];

PMBCFowleria vaiulae (Jordan & Seale, 1906): Cr; [IWP];

PMBCFowleria variegata (Valenciennes, 1832): Cr, Mg, Sg;

[IWP]; PMBCGymnapogon urospilotus Lachner, 1953: Cr; [WCP]Neamia octospina Smith & Radcliffe, 1912: Cr; [IWP]Nectamia fusca (Quoy & Gaimard, 1824): Cr; [IWP];

PMBCNectamia luxuria Fraser, 2008: Cr; [EIO-WP]; PMBCNectamia savayensis (Günther, 1872): Cr; [IP]Pseudamia hayashii Randall, Lachner, & Fraser, 1985: Cr;

[IP]Pseudamia sp.1: CrRhabdamia cypselurus Weber, 1909: Cr; [IWP]; PMBCRhabdamia gracilis (Bleeker, 1856): Cr; [IWP]; PMBCSiphamia tubifer Weber, 1909: Cr; [WP]Zoramia leptacantha (Bleeker, 1856): Cr; [IWP]; PMBCZoramia perlita (Fraser & Lachner, 1985): Cr; [IM]SillaginidaeSillaginopsis panijus (Hamilton, 1822): OShr; [EIO];

PMBCSillago aeolus Jordan & Evermann, 1902: Cr, Mg, Sg, OShr;

[IWP]; PMBCSillago chondropus Bleeker, 1849: OShr; [IWP]; PMBCSillago intermedius Wongratana, 1977: OShr; [EIO-IM]Sillago lutea McKay, 1985: OShr; [EIO-IA]Sillago sihama (Forsskal, 1775): Mg, Sg, OShr; [IWP];

PMBCMalacanthidae

57The fishes of southwestern Thailand

Branchiostegus sawakinensis Amirthalingam, 1969: OShr;[IWP]; PMBC

Hoplolatilus cuniculus Randall & Dooley, 1974: Cr; [IP]Hoplolatilus luteus Allen & Kuiter, 1989: Cr, OShr; [EEIO];

PMBCMalacanthus brevirostris Guichenot, 1848: Cr, OShr; [IP];

PMBCMalacanthus latovittatus (Lacepède, 1801): Cr; [IP]LactariidaeLactarius lactarius (Bloch & Schneider, 1801): OShr;

[NIO-WP]; PMBCCoryphaenidaeCoryphaena equiselis Linnaeus, 1758: OShr; [COS]; PMBCCoryphaena hippurus Linnaeus, 1758: OShr; [COS];

PMBCRachycentridaeRachycentron canadum (Linnaeus, 1766): Cr, Sg, OShr;

[CIR]; PMBCEcheneidaeEcheneis naucrates Linnaeus, 1758: Cr, OShr; [COS];

PMBCPhtheirichthys lineatus (Menzies, 1791): OShr; [COS]Remora remora (Linnaeus, 1758): OShr; [COS]CarangidaeAlectis ciliaris (Bloch, 1788): Cr, OShr; [CIR]; PMBCAlectis indica (Rüppell, 1830): Cr, Sg, OShr; [IWP]; PMBCAlepes djedaba (Forsskål, 1775): Mg, OShr; [IWP]; PMBCAlepes kleinii (Bloch, 1793): Mg, Sg, OShr; [NIO-WP];

PMBCAlepes melanoptera (Swainson, 1839): OShr; [NIO-IM];

PMBCAlepes vari (Cuvier, 1833): Cr, Mg, OShr; [IWP]Atropus atropus (Schneider, 1801): OShr; [NIO-WP]Atule mate (Cuvier, 1833): Cr, Mg, Sg, OShr; [IP]; PMBCCarangoides armatus (Rüppell, 1830): Mg, Sg, OShr;

[IWP]; PMBCCarangoides caeruleopinnatus (Rüppell, 1830): Cr, Sg,

OShr; [IWP]; PMBCCarangoides chrysophrys (Cuvier, 1833): OShr; [IWP]Carangoides equula (Temminck & Schlegel, 1844): Cr,

OShr; [IP]Carangoides ferdua (Forsskål, 1775): Cr, OShr; [IP];

PMBCCarangoides fulvoguttatus (Forsskål, 1775): OShr; [IWP];

PMBCCarangoides gymnostethus (Cuvier, 1833): OShr; [IWP];

PMBCCarangoides hedlandensis (Whitley, 1933): Cr, OShr;

[IWP]; PMBCCarangoides malabaricus (Bloch & Schneider, 1801):

OShr; [IWP]; PMBCCarangoides oblongus (Cuvier, 1833): OShr; [IWP];

PMBCCarangoides orthogrammus (Jordan & Gilbert, 1882): Cr,

OShr; [IP]Carangoides plagiotaenia Bleeker, 1857: Cr, OShr; [IP];

PMBC

Carangoides praeustus (Bennett, 1830): Mg, Sg, OShr;[NIO-IM]; PMBC

Carangoides talamparoides Bleeker, 1852: OShr; [NIO-IA]

Caranx heberi (Bennett, 1830): Cr, OShr; [NIO-WP]Caranx ignobilis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP];

PMBCCaranx melampygus Cuvier, 1833: Cr, OShr; [IP]; PMBCCaranx papuensis Alleyne & Macleay, 1877: Cr, OShr;

[IP]Caranx sexfasciatus Quoy & Gaimard, 1824: Cr, Mg, Sg,

OShr; [IP]; PMBCDecapterus kurroides Bleeker, 1855: OShr; [IWP]; PMBCDecapterus macarellus (Cuvier, 1833): OShr; [CIR];

PMBCDecapterus macrosoma Bleeker, 1851: Cr, OShr; [IP];

PMBCDecapterus maruadsi (Temminck & Schlegel, 1843): OShr;

[WP]; PMBCDecapterus russelli (Rüppell, 1830): Cr, OShr; [IWP];

PMBCElagatis bipinnulata (Quoy & Gaimard, 1825): Cr, OShr;

[CIR]; PMBCGnathanodon speciosus (Forsskål, 1775): Cr, OShr; [IP];

PMBCMegalaspis cordyla (Linnaeus, 1758): Cr, OShr; [IP];

PMBCNaucrates ductor (Linnaeus, 1758): OShr; [CIR]; PMBCParastromateus niger (Bloch, 1795): OShr; [IWP]; PMBCScomberoides commersonianus Lacepède, 1801: Cr, Mg,

Sg, OShr; [IWP]; PMBCScomberoides lysan (Forsskål, 1775): Cr, Mg, Sg, OShr;

[IP]; PMBCScomberoides tala (Cuvier, 1832): OShr; [EIO-WP];

PMBCScomberoides tol (Cuvier, 1832): Cr, Mg, Sg, OShr; [IWP];

PMBCSelar boops (Cuvier, 1833): Cr, OShr; [IWP]; PMBCSelar crumenophthalmus (Bloch, 1793): Cr, OShr; [CIR];

PMBCSelaroides leptolepis (Cuvier, 1833): Cr, Sg, OShr; [NIO-

WP]; PMBCSeriola rivoliana Valenciennes, 1833: OShr; [CIR]; PMBCSeriolina nigrofasciata (Rüppell, 1829): Cr, OShr; [IWP];

PMBCTrachinotus bailloni (Lacepède, 1801): Cr, OShr; [IP];

PMBCTrachinotus blochii (Lacepède, 1801): Cr, OShr; [IP]Trachinotus botla (Shaw, 1803): OShr; [IO]; PMBCUlua mentalis (Cuvier, 1833): Cr, OShr; [IWP]; PMBCUraspis helvola (Forster, 1801): OShr; [IP]; PMBCUraspis uraspis (Günther, 1860): OShr; [IWP]; PMBCMenidaeMene maculata (Bloch & Schneider, 1801): OShr; [IWP];

PMBCLeiognathidaeEquulites berbis (Valenciennes, 1835): OShr; [IWP]; PMBC

Phuket mar. biol. Cent. Res. Bull.58

Equulites elongatus (Günther, 1860): OShr; [IWP]; PMBCEquulites leuciscus (Günther, 1860): Mg, Sg, OShr; [IWP];

PMBCEquulites oblongus (Valenciennes, 1835): Mg, OShr; [IWP];

PMBCEquulites stercorarius (Everman & Seale, 1907): Mg, Sg,

OShr; [WP]; PMBCEubleekeria jonesi (James, 1971): Mg, Sg, OShr; [EIO-

WP]; PMBCGazza dentex (Valenciennes, 1835): OShr; [IO]; PMBCGazza minuta (Bloch, 1797): Mg, Sg, OShr; [IWP]; PMBCGazza rhombea Kimura, Yamashita & Iwatsuki, 2000:

OShr; [EIO-WP]; PMBCKaralla daura (Cuvier, 1829): Mg, OShr; [IWP]; PMBCLeiognathus equulus (Forsskål, 1775): Mg, Sg, OShr;

[IWP]; PMBCLeiognathus fasciatus (Lacepède, 1803): Mg, OShr; [IWP];

PMBCLeiognathus longispinis (Valenciennes, 1835): Mg, Sg,

OShr; [EIO-IA]; PMBCNuchequula blochii (Valenciennes, 1835): Mg, OShr; [NIO-

WP]; PMBCNuchequula gerroides (Bleeker, 1851): Mg, Sg, OShr; [EIO-

WP]; PMBCPhotopectoralis bindus (Valenciennes, 1835): Mg, OShr;

[IWP]; PMBCSecutor hanedai Moshizuki & Hayashi, 1989: Mg, Sg,

OShr; [IM]; PMBCSecutor indicius Monkolprasit, 1973: OShr; [IM]; PMBCSecutor interruptus (Valenciennes, 1835): Mg, OShr; [EIO-

WP]Secutor insidiator (Bloch, 1787): Mg, Sg, OShr; [IWP];

PMBCSecutor megalolepis Moshizuki & Hayashi, 1989: Mg, Sg,

OShr; [IA]; PMBCBramidaeBrama dussumieri Cuvier, 1831: OShr; [CIR]; PMBCLutjanidaeAphareus furca (Lacepède, 1801): OShr; [IP]; PMBCAphareus rutilans Cuvier, 1830: Cr, OShr; [IP]; PMBCAprion virescens Valenciennes, 1830: Cr, OShr; [IP];

PMBCLipocheilus carnolabrum (Chan, 1970): OShr; [NIO-WP];

PMBCLutjanus argentimaculatus (Forsskål, 1775): Cr, Mg, Sg,

OShr; [IP]; PMBCLutjanus bengalensis (Bloch, 1790): Cr, OShr; [IO]Lutjanus biguttatus (Valenciennes 1830): Cr, Mg, Sg, OShr;

[EIO-IA]; PMBCLutjanus bohar (Forsskål, 1775): Cr, OShr; [IP]; PMBCLutjanus decussatus (Cuvier, 1828): Cr, OShr; [EIO-IA];

PMBCLutjanus ehrengergii (Peters, 1869): OShr; [IWP]Lutjanus erythropterus Bloch, 1790: Cr, OShr; [NIO-IA]Lutjanus fulviflamma (Forsskål, 1775): Cr, Mg, Sg, OShr;

[IWP]; PMBC

Lutjanus fulvus (Schneider, 1801): Cr, OShr; [IP]; PMBCLutjanus gibbus (Forsskål, 1775): Cr, OShr; [IP]; PMBCLutjanus johnii (Bloch, 1792): Cr, Mg, Sg, OShr; [IWP];

PMBCLutjanus kasmira (Forsskål, 1775): Cr, OShr; [IP]; PMBCLutjanus lemniscatus (Valenciennes, 1828): Cr, Mg, OShr;

[EIO-IA]; PMBCLutjanus lunulatus (Park, 1797): Cr, Mg, Sg; [NIO-IA];

PMBCLutjanus lutjanus Bloch, 1790: Cr, OShr; [IWP]; PMBCLutjanus madras (Valenciennes, 1831): Cr, Mg, OShr;

[IWP]; PMBCLutjanus malabaricus Schneider, 1801: Cr, OShr; [NIO-

WP]; PMBCLutjanus monostigma (Cuvier, 1828): Cr, OShr; [IP];

PMBCLutjanus quinquelineatus Bloch, 1790: Cr, OShr; [NIO-

WP]; PMBCLutjanus rivulatus (Cuvier, 1828): Cr, OShr; [IP]; PMBCLutjanus rufolineatus (Valenciennes, 1830): OShr; [WP]Lutjanus russelli (Bleeker, 1849): Cr, Mg, Sg, OShr; [IWP];

PMBCLutjanus sebae (Cuvier, 1828): Cr, OShr; [IWP]; PMBCLutjanus timorensis (Quoy & Gaimard, 1824): OShr; [WP]Lutjanus vitta (Quoy & Gaimard, 1824): Cr, Sg, OShr;

[IWP]; PMBCMacolor macularis Fowler, 1931: Cr, OShr; [EIO-WP]Macolor niger (Forsskål, 1775): Cr, OShr; [IP]; PMBCParacaesio xanthurus (Bleeker, 1869): OShr; [INP]Pinjalo lewisi Randall, Allen & Anderson, 1987: Cr; [NIO-

WP]Pinjalo pinjalo (Bleeker, 1850): OShr; [IWP]; PMBCPristipomoides flavipinnis Shinohara, 1963: OShr; [WCP]Pristipomoides multidens (Day, 1871): OShr; [IWP]; PMBCPristipomoides typus Bleeker, 1852: OShr; [IA]; PMBCSymphorichthys spilurus (Günther, 1874): Cr, OShr; [WP]Symphorus nematophorus (Bleeker, 1860): Cr, Sg, OShr;

[WP]; PMBCCaesionidaeCaesio caerulaurea Lacepède, 1801: Cr, Mg, Sg, OShr;

[IP]; PMBCCaesio cuning (Bloch, 1791): Cr, Sg, OShr; [EIO-WP];

PMBCCaesio lunaris Cuvier, 1830: Cr, OShr; [IWP]; PMBCCaesio teres Seale, 1906: Cr, OShr; [IP]Caesio varilineata Carpenter, 1987: Cr, OShr; [IWP]Caesio xanthonota Bleeker, 1853: Cr, OShr; [IO]; PMBCDipterygonotus balteatus (Valenciennes, 1830): Cr, OShr;

[IWP]; PMBCGymnocaesio gymnoptera (Bleeker, 1856): Cr; [IWP];

PMBCPterocaesio chrysozona (Cuvier, 1830): Cr, OShr; [IWP];

PMBCPterocaesio pisang (Bloch, 1853): Cr, OShr; [IWP]; PMBCPterocaesio randalli Carpenter, 1985: Cr; [EIO-IM]Pterocaesio tessellata Carpenter, 1987: Cr, OShr; [EIO-

IA]; PMBC

59The fishes of southwestern Thailand

Pterocaesio tile (Cuvier, 1830): Cr, OShr; [IP]; PMBCLobotidaeLobotes surinamensis (Bloch, 1790): Mg, Sg, OShr; [COS];

PMBCGerreidaeGerres erythrourus (Bleeker, 1791): Mg, Sg, OShr; [EIO-

WP]; PMBCGerres filamentosus (Cuvier, 1829): Cr, Mg, Sg, OShr;

[IP]; PMBCGerres limbatus Cuvier, 1830: Mg, OShr; [EIO-WP];

PMBCGerres longirostris (Lacepède, 1801): Cr, Mg, OShr; [IWP]Gerres lucidus (Cuvier, 1830): Cr; [EIO-WP]Gerres macracanthus Bleeker, 1854: Mg, Sg, OShr; [IWP];

PMBCGerres oblongus (Cuvier, 1830): Cr, Mg, Sg, OShr; [IP];

PMBCGerres oyena (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP];

PMBCGerres phaiya Iwatsuki & Heemstra, 2001: Mg, Sg, OShr;

[EIO]; PMBCGerres setifer (Hamilton, 1822): OShr; [EIO]Gerres shima Iwatsuki, Kimura & Yoshino, 2007: OShr;

[WP]Pentaprion longimanus (Cantor, 1850): OShr; [EIO-WP];

PMBCHaemulidaeDiagramma pictum pictum (Thunberg, 1792): Cr, Sg, OShr;

[WP]; PMBCHapalogenys merguiensis Iwatsuki, Satapoomin &

Amaoka, 2000: OShr; [AS]; PMBCPlectorhinchus chaetodonoides Lacepède, 1800: Cr, OShr;

[IWP]; PMBCPlectorhinchus gibbosus (Lacepède, 1802): Cr, Mg, Sg,

OShr; [IP]; PMBCPlectorhinchus macrospilus Satapoomin & Randall, 2000:

Cr; [AS]; PMBCPlectorhinchus vittatus (Linnaeus, 1758): Cr, OShr; [IWP]Pomadasys andamanensis Mckay & Satapoomin, 1994:

Cr; [AS]; PMBCPomadasys argenteus (Forsskål, 1775): OShr; [IWP];

PMBCPomadasys argyreus (Valenciennes, 1833): Mg, OShr;

[NIO-IM]; PMBCPomadasys auritus (Cuvier, 1830): OShr; [IA]Pomadasys furcatus (Bloch & Schneider, 1801): OShr;

[IM]; PMBCPomadasys kaakan (Cuvier, 1830): Mg, Sg, OShr; [IWP];

PMBCPomadasys maculatus (Bloch, 1793): OShr; [IWP]; PMBCPomadasys olivaceus (Day, 1875): OShr; [IO]Pomadasys unimaculatus Tian, 1982: Mg, Sg, OShr; [WP];

PMBCNemipteridaeNemipterus balinensoides (Popta, 1918): OShr; [WP]Nemipterus bathybius Snyder, 1911: OShr; [IA]; PMBC

Nemipterus bipunctatus (Ehrenberg, 1830): Cr, OShr; [IO];PMBC

Nemipterus furcosus (Valenciennes, 1830): Cr, OShr; [EIO-WP]; PMBC

Nemipterus hexodon (Quoy & Gaimard, 1824): OShr; [WP];PMBC

Nemipterus japonicus (Bloch, 1791): OShr; [IWP]; PMBCNemipterus marginatus (Valenciennes, 1830): OShr; [WP]Nemipterus mesoprion (Bleeker, 1853): OShr; [IM];

PMBCNemipterus nematophorus (Bleeker, 1853): OShr; [EIO-

IM]; PMBCNemipterus peronii (Valenciennes, 1830): Cr, Mg, OShr;

[IWP]; PMBCNemipterus tambuloides (Bleeker, 1853): OShr; [IM];

PMBCNemipterus thosaporni Russell, 1991: OShr; [IM]; PMBCNemipterus zysron (Bleeker, 1856-57): Cr, OShr; [IWP];

PMBCParascolopsis aspinosa (Rao & Rao, 1981): OShr; [NIO]Parascolopsis eriomma (Jordan & Richardson, 1909):

OShr; [IWP]; PMBCParascolopsis inermis (Schlegel, 1843): OShr; [EIO-WP];

PMBCScolopsis affinis Peters, 1877: Cr, Sg, OShr; [WP]; PMBCScolopsis bilineatus (Bloch, 1793): Cr, Sg, OShr; [EIO-

WP]; PMBCScolopsis ciliatus (Lacepède, 1802): Cr, Mg, Sg, OShr;

[WP]; PMBCScolopsis lineatus Quoy & Gaimard, 1824: Cr, OShr;

[WCP]; PMBCScolopsis margaritifer (Cuvier, 1830): Cr, OShr; [WP]Scolopsis monogramma (Cuvier, 1830): Cr, Sg, OShr;

[WP]; PMBCScolopsis taeniopterus (Valenciennes, 1830): Mg, Sg, OShr;

[IA]; PMBCScolopsis vosmeri (Bloch, 1792): Cr, OShr; [IWP]; PMBCScolopsis xenochrous Günther, 1872: Cr, OShr; [EIO-WP]LethrinidaeGnathodentex aurolineatus (Lacepède, 1802): Cr; [IP]Gymnocranius elongatus Senta, 1973: OShr; [IWP]Gymnocranius grandoculis (Valenciennes, 1830): Cr, OShr;

[IP]; PMBCGymnocranius griseus (Schlegel, 1844): Cr, OShr; [EIO-

WP]; PMBCGymnocranius microdon (Bleeker, 1851): Cr, OShr; [WCP]Lethrinus conchyliatus (Smith, 1959): OShr; [IO]Lethrinus crocineus Smith, 1959: Cr, OShr; [IO]; PMBCLethrinus erythracanthus Cuvier, 1830: Cr, OShr; [IP]Lethrinus erythropterus Valenciennes, 1830: Cr, OShr;

[IWP]Lethrinus harak (Forsskål, 1775): Cr, OShr; [IP]; PMBCLethrinus lentjan (Lacepède, 1802): Cr, Sg, OShr; [IWP];

PMBCLethrinus mahsena (Forsskål, 1775): Cr; [IO]; PMBCLethrinus microdon Valenciennes, 1830: Cr, OShr; [IWP]

Phuket mar. biol. Cent. Res. Bull.60

Lethrinus nebulosus (Forsskål, 1775): Cr, OShr; [IP];PMBC

Lethrinus obsoletus (Forsskål, 1775): Cr, OShr; [IP]Lethrinus olivaceus Valenciennes, 1830: Cr, OShr; [IP]Lethrinus ornatus Valenciennes, 1830: Cr, Sg, OShr; [EIO-

WP]; PMBCLethrinus rubrioperculatus Sato, 1978: Cr; [IP]; PMBCLethrinus variegatus Valenciennes, 1830: OShr; [IWP];

PMBCLethrinus xanthochilus Klunzinger, 1870: Cr, OShr; [IP]Monotaxis grandoculis (Forsskål, 1775): Cr, OShr; [IP];

PMBCWattsia mossambrica (Smith, 1957): OShr; [IWP]; PMBCSparidaeAcanthopagrus berda (Forsskål, 1775): Mg, OShr; [IWP]Argyrops spinifer (Forsskål, 1775): Cr, OShr; [IWP]; PMBCPolynemidaeEleutheronema tetradactylum (Shaw, 1804): Mg, OShr;

[NIO-IA]; PMBCEleutheronema tridactylum (Bleeker, 1849): OShr; [IM]Filimanus perplexa Feltes, 1991: OShr; [EEIO]; PMBCFilimanus similis Feltes, 1991: OShr; [NIO]Filimanus xanthonema (Valenciennes, 1831): OShr; [EEIO]Leptomelanosoma indicum (Shaw, 1804): Mg, OShr; [NIO-

IM]; PMBCPolydactylus microstomus (Bleeker, 1851): Mg, OShr; [EIO-

WP]; PMBCPolydactylus plebeius (Broussonet, 1782): Mg, OShr; [IWP];

PMBCPolydactylus sexfilis (Valenciennes, 1831): OShr; [IP]Polydactylus sextarius (Bloch, 1801): OShr; [EIO-WP]Polydactylus siamensis Motomura, Iwatsuki & Yoshino,

2001: OShr; [IM (presently known from west coast ofThailand and Gulf of Thailand)]; PMBC

SciaenidaeDendrophysa russelli (Cuvier, 1830): Mg, Sg, OShr; [EIO-

IM]; PMBCJohnius amblycephalus (Bleeker, 1855): OShr; [NIO-IA];

PMBCJohnius belangerii (Cuvier, 1830): Mg, OShr; [NIO-IM]Johnius borneensis (Bleeker, 1851): OShr; [NIO-IA]Johnius carutta Bloch, 1793: OShr; [NIO]; PMBCJohnius coitor (Hamilton, 1822): OShr: [EIO-IM]Johnius dussumieri (Cuvier, 1830): OShr; [NIO]Johnius latifrons Sasaki, 1992: OShr; [IM]Macrospinosa cuja (Hamilton, 1822): OShr; [EIO]Nibea soldado (Lacepède, 1802): Mg, OShr; [EIO-IA]Otolithes cuvieri Trewavas, 1974: Mg, OShr; [IO]Otolithes ruber (Bloch & Schneider, 1801): OShr; [IWP]Otolithoides biauritus (Cantor, 1850): OShr; [EIO-IM]Panna microdon (Bleeker, 1849): OShr; [IM]Pennahia anea (Bloch, 1793): Mg, Sg, OShr; [NIO-IM];

PMBCProtonibea diacanthus (Lacepède, 1802): OShr; [NIO-IA];

PMBCMullidae

Mulloidichthys flavolineatus (Lacepède, 1801): Cr, Sg, OShr;[IP]

Mulloidichthys vanicolensis (Valenciennes, 1831): Cr, OShr;[IP]; PMBC

Parupeneus barberinus (Lacepède, 1801): Cr, OShr; [IP];PMBC

Parupeneus cyclostomus (Lacepède, 1801): Cr, OShr; [IP];PMBC

Parupeneus heptacanthus (Lacepède, 1801): Cr, Sg, OShr;[IWP]; PMBC

Parupeneus indicus (Shaw, 1803): Cr, Sg, OShr; [IWP];PMBC

Parupeneus macronema (Lacepède, 1801): Cr, OShr; [IWP];PMBC

Parupeneus pleurostigma (Bennett, 1830): Cr, OShr; [IP];PMBC

Parupeneus trifasciatus Lacepède, 1801: Cr, OShr; [IO]Upeneus asymmetricus Lachner, 1954: OShr; [IWP];

PMBCUpeneus guttatus (Day, 1868): OShr; [IWP]Upeneus moluccensis (Bleeker, 1855): OShr; [IWP]Upeneus sulphureus Cuvier, 1829: Mg, Sg, OShr; [IWP];

PMBCUpeneus sundaicus (Bleeker, 1855): Mg, OShr; [NIO-IA];

PMBCUpeneus tragula Richardson, 1846: Cr, Mg, Sg, OShr;

[IWP]; PMBCUpeneus vittatus (Forsskål, 1775): OShr; [IP]; PMBCPempheridaeParapriacanthus ransonneti Steindachner, 1870: Cr; [WP];

PMBCPempheris mangula Cuvier, 1829: Cr; [IWP]; PMBCPempheris otaitensis Cuvier, 1831: Cr; [IWP]Pempheris vanicolensis Cuvier, 1831: Cr; [IM]; PMBCMonodactylidaeMonodactylus argenteus (Linnaeus, 1758): Cr, Mg, OShr;

[IWP]; PMBCToxotidaeToxotes chatareus (Hamiton, 1822): Mg; [EIO-IA]; PMBCToxotes jaculatrix (Pallas, 1766): Mg; [EIO-WP]; PMBCKyphosidaeKyphosus cinerascens (Forsskål, 1775): Cr; [IP]; PMBCKyphosus vaigiensis (Quoy & Gaimard, 1825): Cr; [IP];

PMBCDrepaneidaeDrepane longimana (Bloch & Schneider, 1801): Mg, Sg,

OShr; [IWP]; PMBCDrepane punctata (Linnaeus, 1758): Mg, Sg, OShr; [IWP];

PMBCChaetodontidaeChaetodon andamanensis Kuiter & Debelius, 1999: Cr;

[EIO]; PMBCChaetodon auriga Forsskål, 1775: Cr; [IP]; PMBCChaetodon bennetti Cuvier, 1831: Cr; [IP]Chaetodon citrinellus Cuvier, 1831: Cr; [IP]; PMBCChaetodon collare Bloch, 1787: Cr; [NIO]; PMBC

61The fishes of southwestern Thailand

Chaetodon decussatus Cuvier, 1831: Cr; [EIO]; PMBCChaetodon ephippium Cuvier, 1831: Cr; [IP]; PMBCChaetodon falcula Bloch, 1793: Cr; [IO]; PMBCChaetodon gardineri Norman, 1939: Cr; [IO]; PMBCChaetodon guttatissimus Bennett, 1832: Cr; [IO]; PMBCChaetodon interruptus Ahl, 1922: Cr; [IO]Chaetodon kleinii Bloch, 1790: Cr; [IP]; PMBCChaetodon lineolatus Cuvier, 1831: Cr; [IP]; PMBCChaetodon lunula (Lacepède, 1803): Cr; [IP]; PMBCChaetodon madagaskariensis Ahl, 1923: Cr; [IO]; PMBCChaetodon melannotus Bloch & Schneider, 1801: Cr; [IP];

PMBCChaetodon meyeri Bloch & Schneider, 1801: Cr; [IP];

PMBCChaetodon octofasciatus Bloch, 1787: Cr; [EIO-WP];

PMBCChaetodon oxycephalus Bleeker, 1853: Cr; [EIO-WP]Chaetodon rafflesi Bennett, 1830: Cr; [EIO-WCP]; PMBCChaetodon semeion Bleeker, 1855: Cr; [EIO-WCP]Chaetodon triangulum Cuvier, 1831: Cr; [IO]; PMBCChaetodon trifascialis Quoy & Gaimard, 1825: Cr; [IP];

PMBCChaetodon trifasciatus Park, 1797: Cr; [IO]; PMBCChaetodon vagabundus Linnaeus, 1758: Cr; [IP]; PMBCChaetodon xanthocephalus Bennett, 1832: Cr; [IO]; PMBCChelmon rostratus (Linnaeus, 1758): Cr; [WP]; PMBCCoradion altivelis Mcculloch, 1916: Cr; [WP]Coradion chrysozonus (Cuvier, 1831): Cr, OShr; [WP];

PMBCForcipiger flavissimus Jordan & Mcgregor, 1898: Cr; [IP];

PMBCForcipiger longirostris (Broussonet, 1782): Cr; [IP]Hemitaurichthys zoster (Bennett, 1831): Cr; [IO]; PMBCHeniochus acuminatus (Linnaues, 1758): Cr, Mg, Sg, OShr;

[IP]; PMBCHeniochus diphreutes Jordan, 1903: Cr, OShr; [IP]Heniochus pleurotaenia Ahl, 1923: Cr; [EIO]; PMBCHeniochus singularius Smith & Radcliffe, 1911: Cr; [EIO-

WCP]; PMBCParachaetodon ocellatus (Cuvier, 1831): Cr, Sg: [EIO-WP];

PMBCPomacanthidaeApolemichthys trimaculatus (Lacapède, 1831): Cr; [IWP];

PMBCApolemichthys xanthurus (Bennett, 1832): Cr; [IO]; PMBCCentropyge eibli Klausewitz, 1963: Cr; [EIO]; PMBCCentropyge flavipectoralis Randall & Klausewitz, 1977:

Cr; [EIO]Centropyge multispinis (Playfair, 1866): Cr; [IO]; PMBCChaetodontoplus mesoleucus (Bloch, 1787): Cr; [WP]Genicanthus caudovittatus (Günther, 1860): Cr, OShr; [IO];

PMBCPomacanthus annularis (Bloch, 1787): Cr, OShr; [IWP];

PMBCPomacanthus imperator (Bloch, 1787): Cr; [IP]; PMBCPomacanthus navarchus (Cuvier, 1831): Cr; [IA]

Pomacanthus semicirculatus (Cuvier, 1831): Cr; [IP]Pomacanthus sexstriatus (Cuvier, 1831): Cr; [WP]Pomacanthus xanthometopon (Bleeker, 1853): Cr; [EIO-

WCP]Pygoplites diacanthus (Boddaert, 1772): Cr; [IP]; PMBCTerapontidaePelates quadrilineatus (Bloch, 1790): Mg, Sg; [IWP];

PMBCPseudoterapon theraps (Cuvier, 1829): Mg, Sg, OShr;

[IWP]; PMBCTerapon jarbua (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP];

PMBCTerapon puta (Cuvier, 1829): Mg, Sg, OShr; [IWP]; PMBCKuhliidaeKuhlia mugil (Bloch & Schneider, 1801): Cr; [IP]; PMBCCirrhitidaeCirrhitichthys falco Randall, 1963: Cr; [EIO-WP]Cirrhitichthys oxycephalus (Bleeker, 1855): Cr; [IP];

PMBCOxycirrhites typus Bleeker, 1857: Cr; [IP]Paracirrhites arcatus (Cuvier, 1829): Cr; [IP]Paracirrhites fosteri (Schneider, 1801): Cr; [IP]; PMBCCepolidaeAcanthocepola abbreviata (Valenciennes, 1835): OShr;

[NIO-IM]; PMBCAcanthocepola indica (Day, 1888): OShr; [IWP]PomacentridaeAbudefduf bengalensis (Bloch, 1787): Cr; [EIO-WP];

PMBCAbudefduf notatus (Day, 1869): Cr; [IWP]; PMBCAbudefduf septemfasciatus (Cuvier, 1830): Cr; [IP]; PMBCAbudefduf sordidus (Forsskål, 1775): Cr; [IP]; PMBCAbudefduf vaigiensis (Quoy & Gaimard, 1825): Cr; [IP];

PMBCAmblyglyphidodon aureus (Cuvier, 1830): Cr; [WCP];

PMBCAmblyglyphidodon indicus Allen & Randall, 2002: Cr; [IO]Amblyglyphidodon leucogaster (Bleeker, 1847): Cr; [WP];

PMBCAmphiprion akallopisos Bleeekr, 1853: Cr; [IO]; PMBCAmphiprion clarkii (Bennett, 1830): Cr, Sg; [IWP]; PMBCAmphiprion ephippium (Bloch, 1790): Cr; [EIO]; PMBCAmphiprion ocellaris Cuvier, 1830: Cr; [WP]; PMBCAmphiprion sebae Bleeekr, 1853: Cr, Sg; [EIO]; PMBCCheiloprion labiatus (Day, 1877): Cr; [IA]; PMBCChromis atripectoralis Welander & Schultz, 1951: Cr;

[IWP]; PMBCChromis cinerascens (Cuvier, 1830): Cr; [EIO-IA]; PMBCChromis delta Randall, 1987: Cr; [EIO-WP]Chromis dimidiata (Kluzinger, 1871): Cr; [IO]; PMBCChromis elerae Fowler & Bean, 1928: Cr; [EIO-WP]Chromis flavipectoralis Randall, 1990: Cr; [EIO]; PMBCChromis lepidolepis Bleeker, 1877: Cr; [IP]Chromis opercularis (Günther, 1866): Cr; [IO]; PMBCChromis ternatensis (Bleeker, 1856): Cr; [IP]; PMBCChromis viridis (Cuvier, 1830): Cr; [IP]; PMBC

Phuket mar. biol. Cent. Res. Bull.62

Chromis weberi Fowler & Bean, 1928: Cr; [IP]; PMBCChromis xouthos Allen & Erdmann, 2005: Cr; [AS]Chrysiptera biocellata (Quoy & Gaimard, 1825): Cr; [IP];

PMBCChrysiptera brownriggii (Bennett, 1828): Cr; [IP]; PMBCChrysiptera glauca (Cuvier, 1830): Cr; [IP]Chrysiptera rollandi (Whitley, 1961): Cr; [IA]; PMBCChrysiptera unimaculata (Cuvier, 1830): Cr; [IWP]; PMBCDascyllus aruanus (Linnaeus, 1758): Cr; [IP]; PMBCDascyllus carneus Fischer, 1885: Cr; [IO]; PMBCDascyllus trimaculatus (Rüppell, 1828): Cr; [IP]; PMBCDischistodus perspicillatus (Cuvier, 1830): Cr; [WP];

PMBCHemiglyphidodon plagiometopon (Bleeker, 1852): Cr; [IA];

PMBCLepidozygus tapeinosoma (Bleeker, 1856): Cr; [IP]; PMBCNeoglyphidodon melas (Cuvier, 1830): Cr; [IWP]; PMBCNeoglyphidodon nigroris (Cuvier, 1830): Cr; [WP]; PMBCNeopomacentrus anabatoides (Bleeker, 1847): Cr; [IA];

PMBCNeopomacentrus bankieri (Richardson, 1846): Cr; [WP]Neopomacentrus cyanomos (Bleeker, 1856): Cr; [IWP];

PMBCNeopomacentrus filamentosus (Macleay, 1883): Cr; [IA];

PMBCNeopomacentrus sororius Randall & Allen, 2005: Cr; [IO];

PMBCNeopomacentrus taeniurus (Bleeker, 1856): Cr; [IWP];

PMBCNeopomacentrus violascens (Bleeker, 1848): Cr; [IA]Plectroglyphidodon dickii (Leinard, 1839): Cr; [IP]; PMBCPlectroglyphidodon johnstonianus Fowler & Ball, 1924:

Cr; [IP]; PMBCPlectroglyphidodon lacrymatus (Quoy & Gaimard, 1825):

Cr; [IP]; PMBCPlectroglyphidodon leucozona (Bleeker, 1859): Cr; [IP];

PMBCPomacentrus adelus Allen, 1991: Cr; [IA]; PMBCPomacentrus alleni Burgess, 1981: Cr; [EEIO]; PMBCPomacentrus amboinensis Bleeker, 1868: Cr; [WCP];

PMBCPomacentrus azuremaculatus Allen, 1991: Cr; [EEIO];

PMBCPomacentrus chrysurus Cuvier, 1830: Cr; [EIO-WP];

PMBCPomacentrus lepidogenys Fowler & Bean, 1928: Cr; [WP];

PMBCPomacentrus moluccensis Bleeker, 1853: Cr; [WP]; PMBCPomacentrus nagasakiensis Tanaka, 1917: Cr; [EIO-WP];

PMBCPomacentrus pavo (Bloch, 1787): Cr; [IP]; PMBCPomacentrus philippinus Evermann & Seale, 1907: Cr;

[EIO-WP]; PMBCPomacentrus polyspinus Allen, 1991: Cr; [AS]; PMBCPomacentrus similis Allen, 1991: Cr; [EIO]; PMBCPomacentrus tripunctatus Cuvier, 1830: Cr; [EIO-WP];

PMBC

Pomacentrus xanthosternus Allen, 1991: Cr; [EEIO];PMBC

Premnas biaculeatus (Bloch, 1790): Cr; [IA]Pristotis obtusirostris (Günther, 1862): Cr, OShr; [IWP];

PMBCStegastes fasciolatus (de Vis, 1884): Cr; [IP]; PMBCStegastes nigricans (Lacepède, 1803): Cr; [IP]; PMBCStegastes obreptus (Whitley, 1948): Cr; [EIO-WP]; PMBCStegastes punctatus (Quoy & Gaimard, 1825): Cr; [IWP];

PMBCTeixeirichthys jordani (Rutter, 1897): OShr; [IWP]LabridaeAnampses caeruleopunctatus Rüppell, 1829: Cr; [IP];

PMBCAnampses lineatus Randall, 1972: Cr; [IO]; PMBCAnampses meleagrides Valenciennes, 1840: Cr; [IWP];

PMBCAnampses twistii Bleeker, 1856: Cr; [IP]; PMBCBodianus axillaris (Bennett, 1832): Cr; [IP]Bodianus diana (Lacepède, 1801): Cr; [IO]; PMBCBodianus mesothorax (Bloch & Schneider, 1801): Cr; [WP];

PMBCBodianus neilli (Day, 1867): Cr; [EIO]; PMBCCheilinus chlorourus (Bloch, 1791): Cr, OShr; [IP]; PMBCCheilinus fasciatus (Bloch, 1791): Cr; [IP]; PMBCCheilinus oxycephalus Bleeker, 1853: Cr; [IP]Cheilinus trilobatus Lacepède, 1801: Cr; [IP]; PMBCCheilinus undulatus Rüppell, 1835: Cr; [IP]Choerodon robustus (Günther, 1862): OShr; [IWP];

PMBCCirrhilabrus cyanopleura (Bleeker, 1851): Cr; [EEIO];

PMBCCirrhilabrus exquisitus Smith, 1957: Cr; [IP]; PMBCCirrhilabrus joanallenae Allen, 2000: Cr; [AS]Coris batuensis (Bleeker, 1856): Cr; [EIO-WCP]; PMBCCoris cuvieri (Bennett, 1831): Cr; [IO]; PMBCDiproctacanthus xanthurus (Bleeker, 1856): Cr; [IA]Epibulus insidiator (Pallas, 1770): Cr; [IP]; PMBCGomphosus caeruleus Lacepède, 1801: Cr; [IO]; PMBCHalichoeres argus (Bloch, 1791): Cr; [WP]; PMBCHalichoeres bicolor (Bloch & Schneider, 1801): Cr, Mg,

Sg; [EIO-IM]; PMBCHalichoeres chloropterus (Bloch, 1791): Cr; [IA]; PMBCHalichoeres cosmetus Randall & Smith, 1982: Cr; [IO]Halichoeres hortulanus (Lacepède, 1801): Cr; [IP]; PMBCHalichoeres kallochroma (Bleeker, 1835): Cr; [AS];

PMBCHalichoeres leucoxanthus Randall & Smith, 1982: Cr;

[EIO]; PMBCHalichoeres margaritaceus (Valenciennes, 1839): Cr;

[WCP]Halichoeres marginatus Rüppell, 1835: Cr; [IP]; PMBCHalichoeres nebulosus (Valenciennes, 1839): Cr; [IWP];

PMBCHalichoeres nigrescens (Bloch & Schneider, 1801): Cr, Sg;

[IWP]; PMBCHalichoeres scapularis (Bennett, 1832): Cr; [IWP]; PMBC

63The fishes of southwestern Thailand

Halichoeres timorensis (Bleeker, 1852): Cr; [EIO-IM];PMBC

Halichoeres vrolikii (Bleeker, 1855): Cr; [EIO]; PMBCHalichoeres zeylonicus (Bennett, 1832): Cr, OShr; [IO];

PMBCHemigymnus fasciatus (Bloch, 1792): Cr; [IP]; PMBCHemigymnus melapterus (Bloch, 1791): Cr; [IP]; PMBCHologymnosus annulatus (Lacepède, 1801): Cr; [IP];

PMBCHologymnosus doliatus (Lacepède, 1801): Cr; [IP]Iniistius baldwini (Jordan & Evermann, 1903): OShr; [IP];

PMBCIniistius bimaculatus (Rüppell, 1829): OShr; [IO]; PMBCIniistius pavo (Valenciennes, 1840): Cr, OShr; [IP]; PMBCLabrichthys unilineatus (Guichenot, 1847): Cr; [IP]; PMBCLabroides bicolor Fowler & Bean, 1928: Cr; [IP]Labroides dimidiatus (Valenciennes, 1839): Cr; [IP];

PMBCLabropsis xanthonota Randall, 1981: Cr; [IWP]Leptojulis chrysotaenia Randall & Ferraris, 1981: Cr; [EIO]Leptojulis cyanopleura (Bleeker, 1853): Cr, OShr; [NIO-

WP]; PMBCLeptojulis urostigma Randall, 1996: OShr; [IA]Macropharyngodon ornatus Randall, 1978: Cr; [IWP];

PMBCNovaculichthys taeniourus (Lacepède, 1801): Cr; [IP]Oxycheilinus areanatus (Valenciennes, 1840): Cr; [IP]Oxycheilinus bimaculatus (Valenciennes, 1840): Cr; [IP]Oxycheilinus celebicus (Bleeker, 1853): Cr; [WP]; PMBCOxycheilinus digramma (Lacepède, 1801): Cr, OShr; [IP];

PMBCOxycheilinus orientalis (Günther, 1862): Cr; [IWP]Paracheilinus mccoskeri Randall & Harmelin-Vivien, 1977:

Cr; [IO]; PMBCPseudocheilinus evanidus Jordan & Evermann, 1903: Cr;

[IP]Pseudocheilinus hexataenia (Bleeker, 1857): Cr; [IP];

PMBCPseudodax moluccanus (Valenciennes, 1840): Cr; [IP];

PMBCPseudojuloides kaleidos Kuiter & Randall, 1995: Cr; [EIO];

PMBCPteragogus cryptus Randall, 1981: Cr; [IWP]; PMBCStethojulis albovittata (Bonnaterre, 1788): Cr; [IO]; PMBCStethojulis bandanensis (Bleeker, 1851): Cr; [WCP]Stethojulis interrupta (Bleeker, 1851): Cr; [IWP]; PMBCStethojulis strigiventer (Bennett, 1832): Cr, Sg; [IP]Stethojulis trilineata (Bloch & Schneider, 1801): Cr; [EIO-

WCP]; PMBCSuezichthys caudovittatus (Steindachner, 1898): OShr; [IO];

PMBCThalassoma amblycephalum (Bleeker, 1856): Cr; [IP];

PMBCThalassoma hardwicke (Bennett, 1830): Cr; [IP]; PMBCThalassoma jansenii (Bleeker, 1856): Cr; [EIO-WP];

PMBCThalassoma lunare (Linnaeus, 1758): Cr; [IP]; PMBC

Thalassoma purpureum (Forsskål, 1775): Cr; [IP]Thalassoma trilobatum (Lacepède, 1801): Cr; [IP]; PMBCWetmorella nigropinnata (Seale, 1901): Cr; [IP]; PMBCXiphocheilus quadrimaculatus (Günther, 1880): OShr;

[EIO-WP]ScaridaeBolbometopon muricatum (Valenciennes, 1840): Cr; [IP];

PMBCCalotomus carolinus (Valenciennes, 1840): Cr; [IP]; PMBCCetoscarus bicolor (Rüppell, 1829): Cr; [IP]Chlorurus capistratoides (Bleeker, 1847): Cr; [IO]; PMBCChlorurus rhakoura Randall & Anderson, 1997: Cr; [EIO];

PMBCChlorurus sordidus (Forsskål, 1775): Cr; [IP]; PMBCChlorurus strongylocephalus (Bleeker, 1854): Cr; [IO];Chlorurus troschelii (Bleeker, 1853): Cr; [EEIO]Hipposcarus harid (Forsskål, 1775): Cr; [IO]Scarus frenatus Lacepède, 1802: Cr; [IP]; PMBCScarus ghobban Forsskål, 1775: Cr, Sg; [IP]; PMBCScarus maculipinna Westneat, Satapoomin & Randall,

2007: Cr; [EEIO]; PMBCScarus niger Forsskål, 1775: Cr; [IP]; PMBCScarus prasiognathos Valenciennes, 1840: Cr; [EIO-WP];

PMBCScarus quoyi Valenciennes, 1840: Cr; [EIO-WP]; PMBCScarus rubroviolaceus Bleeker, 1847: Cr; [IP]; PMBCScarus russelii Valenciennes, 1840: Cr; [IO]; PMBCScarus scaber Valenciennes, 1840: Cr; [IO]; PMBCScarus tricolor Bleeker, 1847: Cr; [IWP]; PMBCScarus viridifucatus (Smith, 1956): Cr; [IO]; PMBCChampsodontidaeChampsodon nudivittis (Ogilby, 1895): OShr; [IWP]Champsodon vorax Günther, 1867: OShr; [EIO-WP]PinguipedidaeParapercis alboguttata (Günther, 1872): OShr; [NIO-IA];

PMBCParapercis clathrata Ogilby, 1910: Cr, OShr; [WCP];

PMBCParapercis cylindrica (Bloch, 1792): Cr: [EIO-WP], PMBCParapercis hexophthalma (Cuvier, 1829): Cr, OShr; [IWP];

PMBCParapercis maculata (Bloch & Schneider, 1801): OShr;

[IO]Parapercis millepunctata (Günther, 1860): Cr, OShr; [EIO-

WCP]; PMBCParapercis punctata (Cuvier, 1829): Mg, OShr; [IO];

PMBCParapercis quadrispinosa (Weber, 1913): OShr; [EIO];

PMBCParapercis schauinslandi (Steindachner, 1900): Cr; [IP]Parapercis snyderi Jordan & Starks, 1905: Cr, OShr; [WP]Parapercis tetracantha (Lacepède, 1800): Cr, OShr; [IWP];

PMBCParapercis xanthozona (Bleeker, 1849): Cr, OShr; [IWP];

PMBCTrichonotidaeTrichonotus setiger Bloch & Schneider, 1801: OShr; [WP]

Phuket mar. biol. Cent. Res. Bull.64

Trichonotus sp.1: Cr, OShrPercophidaeBembrops caudimaculata Steindachner, 1876: OShr; [IWP]Bembrops platyrhynchus (Alcock, 1894): OShr; [IWP];

PMBCAmmodytidaeBleekeria kallolepis Günther, 1862: OShr; [EIO]UranoscopidaeIchthyoscopus lebeck (Schneider, 1801): OShr; [EIO];

PMBCUranoscopus affinis Cuvier, 1829: OShr; [IWP]; PMBCUranoscopus bicinctus Temminck & Schlegel, 1843: OShr;

[EIO-WP]Uranoscopus cognatus Cantor, 1849: OShr; [EIO-WP]Uranoscopus oligolepis Bleeker, 1878: OShr; [IO]TripterygiidaeEnnaepterygius fasciatus (Weber, 1909): Cr; [IP]Ennaepterygius nanus (Schultz, 1960): Cr; [WP]Ennaepterygius philippinus (Peters, 1868): Cr; [IWP]Enneapterygius sp.1:Enneapterygius tutuilae Jordan & Seale, 1906: Cr; [IP];

PMBCHelcogramma chica Rosenblatt, 1960: Cr; [EIO-WP]Helcogramma lacuna Williams & Howe, 2003: Cr; [AS]Helcogramma obtusirostre Klunzinger, 1871: Cr; [IWP]Helcogramma rosea Holleman, 2006: Cr; [EIO]Helcogramma sp.1: CrHelcogramma springeri Hansen, 1986: Cr; [WP]; PMBCHelcogramma striata Hansen, 1986: Cr; [EIO-WP]; PMBCNorfolkia brachylepis (Schultz, 1960): Cr; [IP]Ucla xenogrammus Holleman, 1993: Cr; [IP]; PMBCBlenniidaeAlticus saliens (Lacepède, 1800): Cr; [IP]; PMBCAndamia reyi Sauvage, 1880: Cr; [IM]; PMBCAspidontus dussumieri (Valenciennes, 1836): Cr; [IP]Aspidontus tractus Fowler, 1903: Cr; [IO]; PMBCAtrosalarias fuscus (Rüppell, 1835): Cr; [IO]; PMBCBlenniella chrysospilos (Bleeker, 1857): Cr; [IP]Blenniella cyanostigma (Bleeker, 1849): Cr; [IO]Blenniella leopardus (Fowler, 1904): Cr; [AS]; PMBCBlenniella periophthalmus (Valenciennes, 1836): Cr; [IWP]Cirripectes auritus Carlson, 1980: Cr; [IP]Cirripectes castaneus (Valenciennes, 1836): Cr; [IWP]Cirripectes filamentosus (Alleyne & Macleay, 1877): Cr;

[IWP]; PMBCCirripectes stigmaticus Strasburg & Schultz, 1953: Cr;

[IWP]Crossosalarias macrospilus Smith-Vaniz & Springer,

1971: Cr; [WP]Ecsenius bicolor (Day, 1888): Cr; [EIO-WP]; PMBCEcsenius lubbocki Springer, 1988: Cr; [EEIO]; PMBCEcsenius midas Starck, 1969: Cr; [IP]; PMBCEcsenius nalolo Smith, 1959: Cr; [IO]; PMBCEcsenius paroculus Springer, 1988: Cr; [EEIO]; PMBCEnchelyurus kraussi (Klunzinger, 1871): Cr; [IWP]; PMBCEntomacrodus striatus (Quoy & Gaimard, 1836): Cr; [IP];

PMBC

Entomacrodus vermiculatus (Valenciennes, 1836): Cr; [IO];PMBC

Exallias brevis (Kner, 1968): Cr; [IP]Glyptoparus delicatulus Smith, 1959: Cr; [IWP]Istiblennius bellus (Günther, 1861): Cr; [IP]; PMBCIstiblennius dussumieri (Valenciennes, 1836): Cr; [IWP];

PMBCIstiblennius edentulus (Schneider, 1801): Cr; [IP]; PMBCIstiblennius lineatus (Valenciennes, 1836): Cr; [EIO-WCP]Laiphognathus multimaculatus Smith, 1955: Cr; [IWP];

PMBCMeiacanthus smithi Klausewitz, 1916: Cr; [EIO]; PMBCMeiacanthus urostigma Smith-Vaniz, Satapoomin & Allen,

2001: Cr; [AS]; PMBCOmobranchus elongatus (Peters, 1855): Cr; [IWP]; PMBCOmobranchus fasciolatus (Valenciennes, 1836): Mg; [IO];

PMBCOmobranchus ferox (Herre, 1927): Mg; [IP]; PMBCOmobranchus obliquus (Garman, 1903): Cr; [WCP];

PMBCOmobranchus punctatus (Valenciennes, 1836): Mg; [IWP];

PMBCOmobranchus smithi (Rao, 1973): Mg; [EIO-IM]; PMBCOmobranchus zebra (Bleeker, 1868): Mg; [EIO-IM]Petroscirtes breviceps (Valenciennes, 1836): Cr, Mg, Sg;

[IWP]; PMBCPetroscirtes mitratus Rüppell, 1830: Cr, Sg; [IP]; PMBCPetroscirtes thepassi Bleeker, 1853: Sg; [WP]Petroscirtes variabilis Cantor, 1850: Sg; [EIO-IA]; PMBCPlagiotremus phenax Smith-Vaniz, 1976: Cr; [EIO]; PMBCPlagiotremus rhinorhynchos (Bleeker, 1852): Cr; [IP];

PMBCPlagiotremus tapeinosoma (Bleeker, 1857): Cr; [IP];

PMBCPlagiotremus townsendi (Regan, 1905): Cr; [IO]Praealticus oortii (Bleeker, 1851): Cr; [IM]Praealticus triangulus (Chapman, 1951): Cr; [EEIO]Salarias fasciatus (Bloch, 1786): Cr; [IWP]; PMBCSalarias guttatus Valenciennes, 1836: Cr; [IA]; PMBCSalarias sinuosus Snyder, 1908: Cr; [WP]; PMBCXiphasia setifer Swainson, 1839: OShr; [IWP]GobiesocidaeDiademichthys lineatus (Sauvage, 1883): Cr; [IWP]Discotrema crinophilum Briggs, 1976: Cr; [WP]Lepadichthys lineatus Briggs, 1966: Cr; [IWP]CallionymidaeBathycallionymus kaianus (Günther, 1880): OShr; [IWP]Calliurichthys japonicus (Houttuyn, 1782): OShr; [WP];

PMBCDactylopus dactylopus (Valenciennes, 1837): Mg, Sg, OShr;

[WP]; PMBCRepomucenus belcheri recurvispinnis (Li, 1966): OShr;

[EIO-IM]Repomucenus doryssus (Jordan & Fowler, 1903): OShr;

[WP]Repomucenus filamentosus (Valenciennes, 1837): OShr; [IP]

65The fishes of southwestern Thailand

Repomucenus hindsi (Richardson, 1844): Sg, OShr; [NIO-IM]

Repomucenus octostigmatus (Fricke, 1981): OShr; [IM]Repomucenus sagitta (Pallas, 1770): OShr; [EIO-IM]Repomucenus schaapii (Bleeker, 1852): Mg, Sg, OShr; [EIO-

IM]; PMBCParadiplogrammus enneactis (Bleeker, 1879): Mg, Sg;

[WP]; PMBCSynchiropus lineolatus (Valenciennes, 1837): Cr; [IWP]Synchiropus stellatus Smith, 1963: Cr; [IO]EleotridaeBostrychus sinensis (Lacepède, 1801): Mg; [EIO-WP]Butis butis (Hamilton, 1822): Mg, Sg; [IWP]; PMBCButis humeralis (Valenciennes, 1837): Mg; [EIO-WP];

PMBCButis koilomatodon (Bleeker, 1849): Mg, Sg; [IWP]; PMBCCalumia godeffroyi (Günther, 1877): Cr; [IP]Eleotris melanosoma Bleeker, 1852): Mg; [IWP]; PMBCOdonteleotris macrodon (Bleeker, 1853): Mg; [EIO-IM];

PMBCOphiocara aporos (Bleeker, 1854): Mg; [IWP]; PMBCOphiocara porocephala (Valenciennes, 1837): Mg; [IWP];

PMBCXenisthmidaeXenisthmus africanus Smith, 1958: Cr; [IO]Xenisthmus polyzonatus (Klunzinger, 1817): Cr; [IP]GobiidaeAcentrogobius audax Smith, 1959: Sg; [IWP]; PMBCAcentrogobius caninus (Valenciennes, 1837): Mg, Sg; [IWP];

PMBCAcentrogobius chlorostigmatoides (Bleeker, 1849): OShr;

[WP]; PMBCAcentrogobius cyanomos (Bleeker, 1849): Mg, Sg; [EIO-

IM]; PMBCAcentrogobius janthinopterus (Bleeker, 1852): Mg; [WP];

PMBCAcentrogobius madraspatensis (Day, 1868): Mg, Sg; [EIO-

WP]; PMBCAcentrogobius multifasciatus (Herre, 1927): Cr, Sg; [WP];

PMBCAcentrogobius sp.1: SgAcentrogobius viridipunctatus (Valenciennes, 1837): Mg;

[IWP]; PMBCAmblyeleotris aurora (Polunin & Lubbock, 1977): Cr; [IO]Amblyeleotris diagonalis Polunin & Lubbock, 1979: Cr;

[IP]; PMBCAmblyeleotris downingi Randall, 1994: Cr; OShr; [NIO];

PMBCAmblyeleotris fontanesii (Bleeker, 1852): Cr, OShr; [WP];

PMBCAmblyeleotris gymnocephala (Bleeker, 1853): Cr; [IA];

PMBCAmblyeleotris latifasciata Polunin & Lubbock, 1979: Cr;

[IA]; PMBCAmblyeleotris periophthalma (Bleeker, 1853): Cr; [IWP];

PMBC

Amblyeleotris steinitzi (Klausewits, 1974): Cr; [IWP];PMBC

Amblyeleotris triguttatus Randall, 1994: Cr; [IO]Amblyeleotris wheeleri (Polunin & Lubbock, 1977): Cr;

[IWP]Amblygobius hectori (Smith, 1957): Cr; [IWP]; PMBCAmblygobius nocturnus (Herre, 1945): Cr; [NIO-WCP];

PMBCAmblygobius semicinctus (Bennett, 1833): Cr; [IO]; PMBCAmblygobius stethophthalmus (Bleeker, 1851): Cr; [IM]Amoya gracilis (Bleeker, 1875): Mg; [IA]; PMBCAmoya moloanus (Herre, 1927): Mg, Sg; [WP]; PMBCApocryptodon madurensis (Bleeker, 1849): Mg, Sg; [EIO-

WP]; PMBCArcygobius baliurus (Valenciennes, 1837): Sg; [IWP];

PMBCAsterropteryx dfh sp.3: CrAsterropteryx ensiferus (Bleeker, 1874): Cr; [IP]; PMBCAsterropteryx semipunctatus Rüppell, 1828: Cr; [IP];

PMBCBarbuligobius boehlkei Lachner & McKinney, 1974: Cr;

[IWP]Bathygobius cocosensis (Bleeker, 1854): Cr; [IP]; PMBCBathygobius crassiceps (Jordan & Seale, 1906): Cr; [IP];

PMBCBathygobius cyclopterus (Valenciennes, 1837): Cr; [IWP];

PMBCBathygobius fuscus (Rüppell, 1830): Cr, Sg; [IWP]; PMBCBathygobius karachiensis Hoda & Goren, 1990: Cr; [EIO]Bathygobius laddi (Fowler, 1931): Cr; [IP]; PMBCBathygobius meggitti (Hora & Mukuji, 1936): Cr; [WP]Boleophthalmus boddarti (Pallas, 1770): Mg; [NIO-WP];

PMBCBrachygobius kabiliensis Inger, 1958: Mg; [IM]Bryaninops amplus Larson, 1985: Cr; [IP]Bryaninops erythops (Jordan & Seale, 1906): Cr; [EIO-

WP]Bryaninops loki Larson, 1985: Cr; [IWP]Bryaninops natans Larson, 1985: Cr; [IWP]; PMBCBryaninops ridens Smith, 1959: Cr; [IWP]; PMBCBryaninops youngei (Davis & Cohen, 1969): Cr; [IP]Cabillus tongarevae (Fowler, 1927): Cr; [EIO-WP]; PMBCCallogobius andamanensis Menon & Chatterjee, 1974: Mg;

[AS]Callogobius dfh sp.3: CrCallogobius dfh sp.6: CrCallogobius dfh sp.11: CrCallogobius dfh sp.13: CrCallogobius dfh sp.15: CrCallogobius dfh sp.16: CrCallogobius flavobrunneus (Smith, 1958): Cr; [IO]; PMBCCallogobius maculipinnis (Fowler, 1918): Cr; [IWP]Callogobius plumatus (Smith, 1959): Cr; [IO]Callogobius rw-th sp.2: MgCallogobius sclateri (Steindachner, 1880): Cr; [IP]; PMBCCristatogobius nonatoae (Ablan, 1940): Mg; [WP]; PMBC

Phuket mar. biol. Cent. Res. Bull.66

Cryptocentrus caeruleomaculatus (Herre, 1933): Cr, Sg;[IP]; PMBC

Cryptocentrus cyanotaenia (Bleeker, 1853): Cr, Mg, OShr;[IA]; PMBC

Cryptocentrus fasciatus (Playfair & Gunther, 1867): Cr;[IWP]; PMBC

Cryptocentrus leucostictus (Günther, 1871): Cr; [WCP]Cryptocentrus maudae Fowler, 1937: Cr; [IA]; PMBCCryptocentrus melanopus (Bleeker, 1860): Cr, Sg, OShr;

[WP]; PMBCCryptocentrus pavoninoides (Bleeker, 1849): Cr, OShr;

[IA]; PMBCCryptocentrus sp.1 (dfh sp.27): Cr, Sg; [WP]; PMBCCryptocentrus strigilliceps (Jordan & Seale, 1906): Cr, Sg;

[IP]; PMBCCtenogobiops aurocingulus (Herre, 1935): Cr; [WCP]Ctenogobiops crocineus Smith, 1959: Cr; [IWP]; PMBCCtenogobiops feroculus Lubbock & Polunin, 1977: Cr;

[IWP]Ctenogobiops pomastictus Lubbock & Polunin, 1977: Cr;

[WP]; PMBCDrombus globiceps (Hora, 1923): Mg; [EIO-WP]; PMBCDrombus key (Smith, 1947): Cr, Mg, Sg; [IO]; PMBCDrombus ocyurus (Jordan & Seale, 1907): Mg, Sg; [WP];

PMBCDrombus triangularis (Weber, 1911): Mg, Sg; [IWP];

PMBCEugnathogobius illotus (Larson, 1999): Mg; [IM]; PMBCEugnathogobius microps Smith, 1931: Mg; [IM (presently

known from west coast of Thailand, Gulf of Thailandand southern Vietnam)]

Eugnathogobius mindora (Herre, 1945): Mg; WPEugnathogobius polylepis (Wu & Ni, 1985): Mg; [IA]Eugnathogobius variegatus (Peters, 1869): Mg; [IM];

PMBCEviota afelei Jordan & Seale, 1906: Cr; [WCP]Eviota albolineata Jewett & Lachner, 1983: Cr; [IP]; PMBCEviota herrei Jordan & Seale, 1906: Cr; [WCP]Eviota indica Lachner & Karnella, 1980: Cr; [IO]; PMBCEviota infulata (Smith, 1956): Cr; [IP]Eviota mikiae Allen, 2001: Cr; [EIO]Eviota nigriventris Giltay, 1933: Cr; [IA]; PMBCEviota pellucida Larson, 1976: Cr; [WCP]Eviota prasina (Klunzinger, 1871): Cr; [IWP]; PMBCEviota prasites Jordan & Seale, 1906: Cr; [IP]Eviota punctulata Jewett & Lachner, 1983: Cr; [WCP];

PMBCEviota queenslandica Whitley, 1932: Cr; [WP]; PMBCEviota rw-th sp.1: CrEviota rw-th sp.2: CrEviota rw-th sp.4: CrEviota sebreei Jordan & Seale, 1906: Cr; [IP]; PMBCEviota sigillata Jewett & Lachner, 1983: Cr; [IWP]; PMBCEviota spilota Lachner & Karnella, 1980: Cr; [WP]; PMBCEviota zebrina Lachner & Karnella, 1978: Cr: [IWP];

PMBC

Eviota zonura Jordan & Seale, 1906: Cr; [WCP]Exyrias bellissimus (Smith, 1959): Cr; [IWP]; PMBCExyrias puntang (Bleeker, 1851): Mg; [EIO-WP]Favonigobius melanobranchus (Fowler, 1934): Cr, Mg,

Sg; [IWP]; PMBCFavonigobius reichei (Bleeker, 1853): Cr, Mg, Sg; [IWP];

PMBCFlabelligobius latruncularius (Klausewitz, 1974): Cr; [IO]Flabelligobius russus (Cantor, 1849): OShr; [IM]; PMBCFusigobius dfh sp.4: CrFusigobius duospilus Hoese & Reader, 1985: Cr; [IP];

PMBCFusigobius humeralis (Randall, 2001): Cr; [IP]Fusigobius inflamaculatus Randall, 1994: Cr; [IWP]Fusigobius maximus (Randall, 2001): Cr; [IWP]Fusigobius melacron (Randall, 2001): Cr; [WP]Fusigobius neophytus (Günther, 1877): Cr; [IP]; PMBCFusigobius pallidus (Randall, 2001): Cr; [IWP]Fusigobius signipinnis Hoese & Obika, 1988: Cr; [WP]Gladiogobius rex Shibukawa & Allen, 2007: Cr; [IO];

PMBCGlossogobius bicirrhosus (Weber, 1894): Mg; [WP];

PMBCGlossogobius circumspectus (Macleay, 1883): Mg; [WP];

PMBCGlossogobius guiris (Hamilton, 1822): Mg; [IWP]Glossogobius sparsipapillus Akihito & Meguro, 1976: Mg;

[WP]; PMBCGnatholepis anjerensis (Bleeker, 1850): Cr; [IP]; PMBCGnatholepis cauerensis (Bleeker, 1853): Cr; [IP]Gobiodon citrinus (Rüppell, 1853): Cr; [IWP]Gobiodon dfh sp.11: CrGobiodon erythrospilus Bleeker, 1857: Cr; [WP]Gobiodon histrio (Valenciennes, 1837): Cr; [IWP]; PMBCGobiodon prolixus Winterbottom & Harold, 2005: Cr;

[EIO-WP]Gobiodon quinquestrigatus (Cuvier & Valenciennes, 1837):

Cr; [WCP]Gobiodon rivulatus (Rüppell, 1828): Cr; [IWP]; PMBCGobiodon rw-th sp.1: CrGobiodon rw-th sp.2: CrGobiodon rw-th sp.3: CrGobiopsis aporia Lachner & McKinney, 1978: Cr; [EIO-

WP]; PMBCGobiopsis macrostoma Steindachner, 1861: Mg; [NIO-

IM]; PMBCGobiopsis quinquecincta (Smith, 1931): Cr; [EIO-WP];

PMBCGobiopterus brachypterus (Bleeker, 1855): Mg; [IA];

PMBCGobiopterus panayensis (Herre, 1944): Mg; [WP]; PMBCGobiopterus rw-th sp.1: MgHemigobius hoevenii (Bleeker, 1851): Mg; [IA]Hemigobius mingi Herre, 1936: Mg; [IM]Istigobius decoratus (Herre, 1927): Cr; [IWP]; PMBCIstigobius diadema (Steindachner, 1877): Cr, Mg; [EIO-

IA]; PMBC

67The fishes of southwestern Thailand

Istigobius goldmanni (Bleeker, 1852): Cr; [WP]; PMBCIstigobius ornatus (Rüppell, 1830): Cr; [IP]; PMBCIstigobius rigilius (Herre, 1953): Cr; [WCP]Lophogobius bleekeri Popta, 1921: Mg; [IM]; PMBCMahidolia mystacina (Valenciennes, 1837): Cr, Sg, OShr;

[IWP]; PMBCMangarinus seshaiyai (Jacop & Rangarajan, 1960): Mg;

[EIO]Mangarinus waterousi Herre, 1943: Mg; [WP]; PMBCMugilogobius chulae (Smith, 1932): Mg; [IM]; PMBCMugilogobius fasciatus Larson, 2001: Mg; [IM (presently

known from west coast of Thailand and Singapore)];PMBC

Mugilogobius mertoni (Weber, 1911): Mg; [IWP]Mugilogobius tigrinus Larson, 2001: Mg; [IM (presently

known from west coast of Thailand, Singapore and Gulfof Thailand)]; PMBC

Myersina adonis Shibukawa & Satapoomin, 2006; Mg, Sg;[IM (presently known from west coast of Thailand andSingapore]; PMBC

Myersina crocatus (Wongratana, 1975): Cr, OShr; [IM];PMBC

Myersina filifer (Cuvier & Valenciennes, 1837): OShr; [IWP];PMBC

Myersina yangii (Chen, 1960): Cr, OShr; [WP]; PMBCOligolepis acutipennis (Valenciennes, 1837): Mg; [IWP]Oplopomus caninoides (Bleeker, 1852): Cr, OShr; [EIO-

WP]; PMBCOplopomus oplopomus (Valenciennes, 1837): Cr, OShr;

[IP]; PMBCOxyurichthys microlepis (Bleeker, 1849): Mg, Sg; [IWP];

PMBCOxyurichthys ophthalmonema (Bleeker, 1856): Mg, Sg;

[EIO-WP]Oxyurichthys papuensis (Valenciennes, 1837): Cr, OShr;

[IWP]; PMBCOxyurichthys tentacularis (Valenciennes, 1837): Mg, Sg;

[WP]; PMBCPalutrus scapulopunctatus (De Beaufort, 1912): Cr; [IWP];

PMBCPandaka lidwilli (McCulloch, 1917): Mg; [WP]; PMBCPandaka pygmaea Herre, 1927: Mg; [IM]Parachaeturichthys polynema (Bleeker, 1853): Mg; [IWP];

PMBCParagobiodon modestus (Regan, 1908): Cr; [IP]; PMBCParatrypauchen microcephalus (Bleeker, 1860): Mg, Sg;

[IWP]Periophthalmus argentilineatus Valenciennes, 1837: Mg;

[IWP]; PMBCPeriophthalmus chrysospilos Bleeker, 1852: Mg; [EIO-

IM]; PMBCPeriophthalmus kalolo Lesson, 1830: Mg; [IWP]; PMBCPeriophthalmus minutus Eggert, 1935: Mg; [IA]; PMBCPeriophthalmus novemradiatus (Hamilton, 1822): Mg;

[EIO-IA]; PMBCPeriophthalmus septemradiatus (Hamilton, 1822): Mg;

[EIO-IM]

Periophthalmus walailakae Darumas & Tantochodok,2002: Mg; [IM (presently known from west coast ofThailand and Singapore)]; PMBC

Pleurosicya bilobata (Koumans, 1941): Sg; [IWP]Pleurosicya coerulea Larson, 1990: Cr; [IP]Pleurosicya fringilla Larson, 1990: Cr; [IWP]; PMBCPleurosicya labiata (Weber, 1913): Cr; [EIO-IA]; PMBCPleurosicya micheli Fourmanoir, 1971: Cr; [IP]Pleurosicya mossambica Smith, 1959: Cr; [IP]; PMBCPleurosicya plicata Larson, 1990: Cr; [IWP]Pleurosicya prognatha Goren, 1984: Cr; [IWP]Priolepis cinctus (Regan, 1908): Cr; [IWP]; PMBCPriolepis inhaca (Smith, 1949): Cr; [IWP]Priolepis nuchifasciatus (Günther, 1873): Cr; [IA]Priolepis semidoliatus (Valenciennes, 1837): Cr; [IP];

PMBCPsammogobius biocellatus (Valenciennes, 1837): Mg, Sg;

[IWP]; PMBCPseudapocryptes lanceolatus (Bloch & Schneider, 1801):

Mg; [EIO-IM]Pseudogobius javanicus (Bleeker, 1856): Mg; [IM]; PMBCRedigobius bikolanus Herre, 1927: Mg; [WP]; PMBCRedigobius chrysosoma (Bleeker, 1857): Mg; [IA]; PMBCScartelaos cantoris (Day, 1871): Mg, Sg; [AS]; PMBCScartelaos histophorus (Valenciennes, 1837): Mg, Sg; [NIO-

WP]; PMBCSilhouettea sp.1: Cr, Sg; PMBCStigmatogobius pleurostigma (Bleeker, 1849): Mg; [IM]Stigmatogobius sandanundio (Hamilton, 1822): Mg; [EIO];

PMBCStonogobiops nematodes Hoese & Randall, 1982: Cr; [IA]Sueviota lachneri Winterbottom & Hoese, 1988: Cr; [IWP];

PMBCTaenioides cirratus (Blyth, 1860): Mg, Sg, OShr; [IWP];

PMBCTaenioides gracilis (Valenciennes, 1837): OShr; [IWP];

PMBCTrimma emeryi Winterbottom, 1985: Cr; [IO]Trimma flammeum (Smith, 1959): Cr; [IO]; PMBCTrimma fucatum Winterbottom & Southcott, 2007: Cr;

[AS]; PMBCTrimma naudei Smith, 1956: Cr; [IWP]; PMBCTrimma okinawae (Aoyaki, 1949): Cr; [WCP]; PMBCTrimma sanguinellus Winterbottom & Southcott, 2007:

Cr; [AS]; PMBCTrimma striata (Herre, 1945): Cr; [EIO-WP]; PMBCTrimma talori Lobel, 1979: Cr; [IP]Trimma winterbottomi Randall & Downing, 1994: Cr; [IO];

PMBCTrypauchen vagina (Bloch & Schneider, 1801): OShr; [NIO-

IM]; PMBCValenciennea immaculata (Ni, 1981): Sg; [WP]Valenciennea limicola Hoese & Larson, 1994: Cr; [WP];

PMBCValenciennea mularis (Valenciennes, 1837): Cr, Sg, OShr;

[IA]Valenciennea parva Hoese & Larson, 1994: Cr; [IWP]

Phuket mar. biol. Cent. Res. Bull.68

Valenciennea puellaris (Tomiyama, 1956): Cr, OShr; [IP];PMBC

Valenciennea randalli Hoese & Larson, 1994: Cr, OShr;[WP]; PMBC

Valenciennea sexguttata (Valenciennes, 1837): Cr, OShr;[IP]; PMBC

Valenciennea strigata (Broussonet, 1782): Cr; [IP]; PMBCValenciennea wardii (Playfair, 1867): Cr, OShr; [IWP];

PMBCVanderhorstia ambanoro (Fourmanoir, 1957): Cr, OShr;

[IWP]; PMBCVanderhorstia ornatissima Smith, 1959: Cr; [IP]Yongeichthys nebulosus (Forsskål, 1775): Cr, Mg, Sg, OShr;

[IWP]; PMBCMicrodesmidaeGunnellichthys curiosus Dawson, 1968: Cr; [IP]Gunnellichthys viridescens Dawson, 1968: Cr; [IP]PtereleotridaeNemateleotris decora Randall & Allen, 1973: Cr; [IWP]Nemateleotris magnifica Fowler, 1928: Cr; [IP]Oxymetopon compressus Chan, 1966: Cr, OShr; [WP];

PMBCParioglossus formosus (Smith, 1931): Cr; [WP]; PMBCParioglossus palustris (Herre, 1945): Mg; [WP]Parioglossus philippinus (Herre, 1940): Cr; [IWP]; PMBCPtereleotris arabica Randall & Hoese, 1985: Cr, OShr;

[IO]; PMBCPtereleotris evides (Jordan & Hubbs, 1925): Cr; [IP]Ptereleotris heteroptera (Bleeker, 1855): Cr; [IP]; PMBCPtereleotris microlepis (Bleeker, 1856): Cr; [IP]; PMBCPtereleotris monoptera (Bleeker, 1855): Cr; [IP]Ptereleotris zebra (Fowler, 1938): Cr; [IP]KurtidaeKurtus indicus Bloch, 1786: OShr; [EIO-IM]; PMBCEphippidaeEphippus orbis (Bloch, 1787): Mg, Sg, OShr; [IWP];

PMBCPlatax boersi Bleeker, 1852: Cr; [IP]Platax orbicularis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP];

PMBCPlatax pinnatus (Linnaeus, 1758): Cr, OShr; [WP]; PMBCPlatax teira (Forsskål, 1775): Cr, OShr; [IWP]; PMBCScatophagidaeScatophagus argus (Linnaeus, 1758): Mg, Sg; [NIO-WP];

PMBCSiganidaeSiganus argenteus (Quoy & Gaimard, 1825): Cr; [IP];

PMBCSiganus canaliculatus (Park, 1797): Cr, Mg, Sg, OShr; [NIO-

IA]; PMBCSiganus corallinus (Valenciennes, 1835): Cr; [IWP]; PMBCSiganus fuscescens (Houttuyn, 1782): Cr; [WP]; PMBCSiganus guttatus (Bloch, 1787): Cr, Mg, Sg, OShr; [IM]Siganus javus (Linnaeus, 1766): Cr, Mg, Sg, OShr; [NIO-

WP]Siganus lineatus (Valenciennes, 1835): Cr; [EIO-WP]

Siganus magnificus (Burgess, 1977): Cr; [EEIO]; PMBCSiganus puelloides Woodland & Randall, 1979: Cr; [EIO];

PMBCSiganus stellatus laqueus Von Bonde, 1934: Cr, OShr; [IO];

PMBCSiganus vermiculatus (Valenciennes, 1835): Cr, Mg, Sg,

OShr; [EIO-WP]; PMBCSiganus virgatus (Valenciennes, 1835): Cr, Sg, OShr; [EIO-

WP]; PMBCZanclidaeZanclus cornutus (Linnaeus, 1758): Cr; [IP]; PMBCAcanthuridaeAcanthurus bariene Lesson, 1830: Cr; [WP-EIO]; PMBCAcanthurus blochii Valenciennes, 1835: Cr; [IP]Acanthurus guttatus Forster, 1801: Cr; [IP]Acanthurus leucocheilus Herre, 1927: Cr; [IWP]Acanthurus leucosternon Bennett, 1832: Cr; [IO]; PMBCAcanthurus lineatus (Linnaeus, 1758): Cr; [IP]; PMBCAcanthurus maculiceps (Ahl, 1923): Cr; [EIO-WCP]Acanthurus mata (Cuvier, 1829): Cr; [IP]; PMBCAcanthurus nigricauda Duncker & Mohr, 1929: Cr; [IP];

PMBCAcanthurus nigrofuscus (Forsskål, 1775): Cr; [IP]Acanthurus olivaceus Forster, 1801: Cr; [WCP]Acanthurus tennenti Günther, 1861: Cr; [IO]Acanthurus thompsoni (Fowler, 1923): Cr; [IP]; PMBCAcanthurus triostegus (Linnaeus, 1758): Cr; [IP]; PMBCAcanthurus tristis Tickell In Day, 1888: Cr; [IO]; PMBCAcanthurus xanthopterus Valenciennes, 1835: Cr; [IP];

PMBCCtenochaetus binotatus Randall, 1955: Cr; [IP]; PMBCCtenochaetus striatus (Quoy & Gaimard, 1825): Cr; [IP];

PMBCCtenochaetus truncatus Randall & Clements, 2001: Cr;

[IO]; PMBCNaso brachycentron (Valenciennes, 1835): Cr; [IP]Naso brevirostris (Valenciennes, 1835): Cr; [IP]; PMBCNaso caeruleocauda Randall, 1994: Cr; [IA]Naso elegans (Rüppell, 1829): Cr; [IO]; PMBCNaso hexacanthus (Bleeker, 1855): Cr; [IP]; PMBCNaso lopezi Herre, 1927: Cr; [WP]Naso mcdadei Johnson, 2002: Cr; [IWP]Naso thynnoides (Valenciennes, 1835): Cr; [IP]Naso unicornis (Forsskål, 1775): Cr; [IP]; PMBCNaso vlamingii (Valenciennes, 1835): Cr; [IP]; PMBCParacanthurus hepatus (Linnaeus, 1766): Cr; [IP]Zebrasoma desjardinii (Bennett, 1835): Cr; [IO]; PMBCZebrasoma scopas (Cuvier, 1829): Cr; [IP]; PMBCSphyraenidaeSphyraena baracuda (Walbaum, 1792): Cr, Sg, OShr;

[COS]; PMBCSphyraena forsteri Cuvier, 1829: Cr, OShr; [IP]; PMBCSphyraena jello Cuvier, 1829: Cr, Mg, Sg, OShr; [IP];

PMBCSphyraena obtusata Cuvier, 1829: Cr, Sg, OShr; [IWP];

PMBC

69The fishes of southwestern Thailand

Sphyraena pinguis Günther, 1874: Cr; [IWP]Sphyraena putnamae Jordan & Seale, 1905: Mg, Sg, OShr;

[IWP]Sphyraena qenie Klunzinger, 1870: Cr; [IP]GempylidaeGempylus serpens Cuvier, 1829: OShr; [COS]Neoepinnula orientalis (Gilchrist & von Bonde, 1924): OShr;

[IWP]; PMBCNesiarchus nasutus Johnson, 1862: OShr; [COS]; PMBCPromethichthys prometheus (Cuvier, 1832): OShr; [COS]Rexea bengalensis (Alcock, 1884): OShr; [IWP]Thyrsitoides marleyi Fowler, 1929: OShr; [IWP]TrichiuridaeEupleurogrammus muticus (Gray, 1831): OShr; [NIO-IM]Tentoriceps cristatus (Klunzinger, 1884): OShr; [IWP];

PMBCTrichiurus gangeticus Gupta, 1966: OShr; [EIO]; PMBCTrichiurus lepturus Linnaeus, 1758: Mg, Sg, OShr; [COS];

PMBCScombridaeAcanthocybium solandi (Cuvier, 1831): Cr, OShr; [COS]Auxis rochei rochei (Risso, 1810): OShr; [COS]; PMBCAuxis thazard thazard (Lacepède, 1800): OShr; [COS];

PMBCEuthynnus affinis (Cantor, 1849): Cr, OShr; [IP]; PMBCGrammatorcynos bilineatus (Rüppell, 1836): Cr, OShr;

[IP]; PMBCGymnosarda unicolor (Rüppell, 1838): Cr, OShr; [IP];

PMBCKatsuwonus pelamis (Linnaeus, 1758): OShr; [COS];

PMBCRastrelliger brachysoma (Bleeker, 1851): OShr; [WP];

PMBCRastrelliger faughni Matsui, 1967: OShr; [EIO-WP];

PMBCRastrelliger kanagurta (Cuvier, 1817): Cr, Mg, Sg, OShr;

[IP]; PMBCScomberomorus commerson (Lacepède, 1800): Cr, Mg,

OShr; [IP]; PMBCScomberomorus guttatus (Bloch & Schneider, 1801): OShr;

[NIO-WP]; PMBCScomberomorus lineolatus (Cuvier, 1831): OShr; [NIO-

IM]Thunnus alalunga (Bonnaterre, 1788): OShr; [COS]Thunnus albacares (Bonnaterre, 1788): OShr; [COS];

PMBCThunnus obesus (Lowe, 1839): OShr; [COS]Thunnus tonggol (Bleeker, 1851): OShr; [IWP]; PMBCXiphiidaeXiphias gladius Linnaeus, 1758: OShr; [COS]IstiophoridaeIstiophorus platypterus (Shaw & Nodder, 1792): OShr;

[IP]; PMBCMakaira indica (Cuvier, 1832): OShr; [IP]Tetrapturus audax (Philippi, 1887): OShr; [IP]Centrolophidae

Psenopsis obscura Headrich, 1967: OShr; [IM]NomeidaeCubiceps pauciradiatus Günther, 1872: OShr; [CIR];

PMBCCubiceps whiteleggii (Waite, 1894): OShr; [IWP]AriommatidaeAriomma indicum (Day, 1870): OShr; [IWP]; PMBCStromateidaePampus argenteus (Euphrasen, 1788): OShr; [NIO-WP];

PMBCPampus chinensis (Euphrasen, 1788): OShr; [NIO-WP]CaproidaeAntigonia rubescens (Günther, 1860): OShr; [WP]Order PleuronectiformesPsettodidaePsettodes erumei (Bloch & Schneider, 1801): Mg, OShr;

[IWP]; PMBCCitharidaeBrachypleura novaezeelandiae Günther, 1862: OShr;

[NIO-IA]; PMBCParalichthyidaePseudorhombus argus Weber, 1913: OShr; [IA]Pseudorhombus arsius (Hamilton, 1822): Mg, Sg, OShr;

[IWP]; PMBCPseudorhombus dupliciocellatus Regan, 1905: OShr; [IA];

PMBCPseudorhombus elevatus Ogilby, 1912: Mg, OShr; [IWP];

PMBCPseudorhombus malayanus Bleeker, 1866: Mg, OShr;

[NIO-IM]Pseudorhombus quinquocellatus Weber & de Beaufort,

1929: OShr; [EIO-IA]; PMBCPseudorhombus triocellatus (Schneider, 1801): OShr; [EIO-

IA]; PMBCBothidaeArnoglossus aspilos (Bleeker, 1851): Cr; [NIO-IA]; PMBCArnoglossus macrolophus Alcock, 1889: OShr; [IWP]Arnoglossus tapeinosoma (Bleeker, 1865): OShr; [NIO-

WP]Arnoglossus tenuis Günther, 1880: OShr; [WP]; PMBCAsterorhombus intermedius (Bleeker, 1866): OShr; [IWP]Bothus pantherinus (Rüppell, 1830): OShr; [IP]Chascanopsetta lugubris Alcock, 1894: OShr; [IWP]Engyprosopon grandisquama (Temminck & Schlegel,

1846): OShr; [IWP]; PMBCEngyprosopon latifrons (Regan, 1908): OShr; [IWP];

PMBCEngyprosopon maldivensis (Regan, 1908): OShr; [EIO-WP]Grammatobothus polyophthalmus (Bleeker, 1866): OShr;

[EIO-WP]; PMBCLaeops guentheri Alcock, 1890: OShr; [NIO-IM]Laeops kitaharae (Smith & Pope, 1906): OShr; [IWP]PoecilopsettidaePoecilopsetta colorata Günther, 1880: OShr; [EIO-IM]SamaridaeSamaris cristatus Gray, 1831: OShr; [IWP]; PMBC

Phuket mar. biol. Cent. Res. Bull.70

SoleidaeAesopia cornuta Kaup, 1858: OShr; [IWP]; PMBCAseraggodes dubius Weber, 1913: OShr; [WP]Aseraggodes satapoomini Randall & Desoutter-Meniger,

2007: Cr; [AS]Brachirus pan (Hamilton, 1822): OShr; [EIO-IM]Heteromycteris oculus (Alcock, 1889): OShr; [IWP];

PMBCLiachirus melanospilos (Bleeker, 1854): OShr; [EIO-WP]Pardachirus pavoninus (Lacepède, 1802): Cr, Sg, OShr;

[EIO-WP]; PMBCSolea ovata Richardson, 1846: Sg, OShr; [EIO-IM]; PMBCSynaptura commersonii (Lacepède, 1802): Sg, OShr; [NIO-

IM]; PMBCZebrias altipinnis (Alcock, 1890): OShr; [EIO]Zebrias quagga Kaup, 1858: Mg, Sg, OShr; [IWP]; PMBCZebrias synapturoides (Jenkins, 1910): OShr; [NIO-WP]CynoglossidaeCynoglossus arel (Schneider, 1801): Mg, OShr; [NIO-IM]Cynoglossus bilineatus (Lacepède, 1802): Mg, OShr; [NIO-

WP]; PMBCCynoglossus cynoglossus (Hamilton, 1822): OShr; [NIO-

IM]; PMBCCynoglossus kopsi (Bleeker, 1851): OShr; [NIO-WP]Cynoglossus lida (Bleeker, 1851): Mg, Sg, OShr; [IWP];

PMBCCynoglossus lingua Hamilton, 1822: Mg, OShr; [IWP]Cynoglossus macrolepidotus (Bleeker, 1851): OShr; [WP];

PMBCCynoglossus monopus (Bleeker, 1849): OShr; [EIO-IM]Cynoglossus oligolepis (Bleeker, 1854): OShr; [WP];

PMBCCynoglossus puncticeps (Richardson, 1846): Sg, OShr;

[NIO-IA]; PMBCCynoglossus semifasciatus Day 1877: Mg, Sg, OShr; [EIO];

PMBCCynoglossus sibogae Weber, 1913: OShr; [IM]; PMBCParaplagusia bilineata (Bloch, 1787): OShr; [IWP]Paraplagusia blochii (Bleeker, 1851): OShr; [NIO-WP]Symphurus septemstriatus (Alcock, 1891): OShr; [EIO-

IM]Order TetraodontiformesTriacanthodidaeHalimochirurgus alcocki Weber, 1913: OShr; [IWP]Tydemania navigatoris Weber, 1913: OShr; [IWP]; PMBCTriacanthidaePseudotriacanthus strigilifer (Cantor, 1849): OShr; [NIO-

IA]; PMBCTriacanthus biaculeatus (Bloch, 1786): Mg, Sg, OShr;

[NIO-WP]; PMBCTriacanthus nieuhofi Bleeker, 1852: Sg, OShr; [EIO-IA]Trixiphichthys weberi (Chaudhuri, 1910): OShr; [EIO-IA];

PMBCBalistidaeAbalistes stellatus (Anonymous, 1798): Cr, OShr; [IWP];

PMBC

Balistapus undulatus (Park, 1797): Cr, OShr; [IP]; PMBCBalistoides conspicillum (Bloch & Schneider, 1801): Cr,

OShr; [IP]; PMBCBalistoides viridescens (Bloch & Schneider, 1801): Cr,

OShr; [IP]; PMBCCanthidermis maculatus (Bloch, 1786): OShr; [COS];

PMBCCanthidermis rotundatus (Marion de Procé, 1822): OShr;

[IWP]; PMBCMelichthys indicus Randall & Klausewitz, 1973: Cr, OShr;

[IO]; PMBCMelichthys vidua (Solander, 1844): Cr; [IP]Odonus niger (Rüppell, 1837): Cr, OShr; [IP]; PMBCPseudobalistes flavimarginatus (Rüppell, 1829): Cr, OShr;

[IP]Pseudobalistes fuscus (Bloch & Schneider, 1801): Cr, OShr;

[IP]; PMBCRhinecanthus aculeatus (Linnaeus, 1758): Cr, OShr; [IP]Rhinecanthus rectangulus (Bloch & Schneider, 1801): Cr,

OShr; [IP]Rhinecanthus verrucosus (Linnaeus, 1758): Cr, OShr; [IWP]Sufflamen bursa (Bloch & Schneider, 1801): Cr; [IP]Sufflamen chrysopterum (Bloch & Schneider, 1801): Cr,

OShr; [IWP]; PMBCSufflamen fraenatum (Latreille, 1804): Cr, OShr; [IP];

PMBCXanthichthys auromarginatus (Bennett, 1831): Cr, OShr;

[IP]MonacanthidaeAcreichthys tomentosus (Linnaeus, 1758): Sg; [IA]Aluterus monoceros (Linnaeus, 1758): OShr; [CIR]; PMBCAluterus scriptus (Osbeck, 1765): Cr, OShr; [CIR]; PMBCAmanses scopas (Cuvier, 1829): Cr; [IP]; PMBCAnacanthus barbatus Gray, 1831: Sg, OShr; [NIO-IA]Cantherhines dumerilli (Hollard, 1854): Cr, OShr; [IP]Cantherhines multilineatus (Tanaka, 1918): OShr; [IWP;

PMBCCantherhines pardalis (Rüppell, 1837): Cr, OShr; [IP];

PMBCMonacanthus chinensis Osbeck, 1765: Cr, Mg, Sg, OShr;

[WP]; PMBCOxymonacanthus longirostris (Bloch & Schneider, 1801):

Cr; [IP]; PMBCParaluteres sp.1: Cr; [AS]Paramonacanthus japonicus (Tilesius, 1810): OShr; [EIO-

WP]Paramonacanthus tricuspis (Hollard, 1854): Mg, Sg, OShr;

[EIO]; PMBCPseudomonacanthus macrurus (Bleeker, 1857): Mg, Sg,

OShr; [IWP]; PMBCPseudomonacanthus peroni (Hollard, 1854): OShr; [IA]OstraciidaeLactoria cornuta (Linnaeus, 1758): Cr, Mg, Sg, OShr; [IP];

PMBCLactoria diaphana (Bloch & Schneider, 1801): Mg, OShr;

[IP]

71The fishes of southwestern Thailand

Ostracion cubicus Linnaeus, 1758: Cr, Mg, Sg, OShr; [IP];PMBC

Ostracion meleagris Shaw, 1796: Cr; [IP]; PMBCOstracion nasus Bloch, 1785: OShr; [EIO-WP]; PMBCOstracion rhinorhynchus Bleeker, 1852: Cr, OShr; [IWP]Ostracion solorensis Bleeker, 1853: Cr; [IWP]Tetrosomus gibbosus (Linnaeus, 1758): Cr, OShr; [IWP];

PMBCTetrosomus reipublicae (Whitley, 1930): OShr; [IWP]TetraodontidaeAmblyrhynchotes honckenii (Bloch, 1795): OShr; [IWP]Arothron caeruleopunctatus Matsuura, 1994: Cr, Mg, Sg;

[EIO-WP]Arothron hispidus (Linnaeus, 1758): Cr, Mg, Sg, OShr;

[IP]; PMBCArothron immaculatus (Bloch & Schneider, 1801): Cr, OShr;

[IWP]; PMBCArothron manilensis (Marion de Porcé, 1822): Cr, Sg, OShr;

[WCP]Arothron mappa (Lesson, 1830): Cr, OShr; [IP]; PMBCArothron nigropunctatus (Bloch & Schneider, 1801): Cr,

OShr; [IP]; PMBCArothron reticularis (Bloch & Schneider, 1801): Mg, OShr;

[IWP]; PMBCArothron stellatus (Bloch & Schneider, 1801): Cr, OShr;

[IP]; PMBCCanthigaster smithae Allen & Randall, 1977: Cr; [IO]Canthigaster solandri (Richardson, 1844): Cr, OShr; [IP];

PMBC

Chelonodon patoca (Hamilton, 1822): Mg, Sg, OShr; [IWP];PMBC

Lagocephalus gloveri Abe & Tabeta, 1983: OShr; [IWP]Lagocephalus inermis (Temminck & Schlegel, 1844): OShr;

[IWP]; PMBCLagocephalus lunaris (Bloch & Schneider, 1801): Mg, Sg,

OShr; [IWP]; PMBCLagocephalus scleratus (Gmelin, 1789): OShr; [IWP];

PMBCLagocephalus spadiceus (Richardson, 1844): OShr; [IWP]Takifugu oblongus (Bloch, 1786): Mg, Sg, OShr; [IWP];

PMBCTetraodon fluviatilis Hamilton, 1822: OShr; [EIO-IM];

PMBCTetraodon nigroviridis Procé, 1822: Mg, Sg, OShr; [EIO-

WP]; PMBCTorquigener hypselogeneion (Bleeker, 1852): Cr; [IWP];

PMBCDiodontidaeCyclichthys orbicularis (Bloch, 1785): Mg, Sg, OShr;

[IWP]; PMBCDiodon eydouxii Brisout de Barneville, 1846: OShr; [IWP]Diodon holocanthus Linnaeus, 1758: OShr; [CIR]; PMBCDiodon hystrix Linnaeus, 1758: Cr, OShr; [CIR]; PMBCDiodon liturosus Shaw, 1804: Cr, Sg, OShr; [IP]; PMBCMolidaeMasturus lanceolatus (Lienard, 1840): OShr; [COS]Mola mola (Linnaeus, 1758): OShr; [COS]; PMBC

Phuket mar. biol. Cent. Res. Bull.72

Appendix B

Selected taxonomic literature pertaining to the records of fishes from southwestern Thailand,the Andaman Sea

Class MyxiniWongratana, T. 1983. Eptatretus indrambaryai, a new species of hagfish (Myxinidae) from the Andaman Sea. Nat Hist.

Bull. Siam Soc. 31(2): 139–150.Class ChondrichthyesCompagno, L.J.V. and P.R. Last. 2008. A new species of wedgefish, Rhynchobatus palpebratus sp. nov. (Rhynchobatoidei:

Rhynchobatidae), from the Indo-West Pacific. In: P.R. Last, W.T. White and J.J. Pogonoski (eds.). Descriptionsof New Australian Chondrichthyans. CISRO Marine and Atmospheric Research Paper No. 022. pp. 227–240.

Krajangdara, T., S. Puteeka, R. Phoonsawat, S. Vibunpant, R. Sojittosakul and R. Chayakul. 2006. Sharks found in Thaiwaters and adjacent areas. Marine Fisheries Research and Development Bureau, Department of Fisheries.Technical Paper No. 17/2006. 99 p. (in Thai)

Monkolprasit, S. 1984. The cartilaginous fishes (Class Elasmobranchii) found in Thai waters and adjacent areas.Department of Fishery Biology, Faculty of Fisheries, Kasetsart University, Bangkok, Thailand. 175 p.

Class ActinopterygiiOrder AlbuliformesHidaka, K., Y. Iwatsuki and J.E. Randall. 2008. A review of the Indo-Pacific bonefishes of the Albula argentea complex,

with a description of a new species. Ichthyol. Res. 55(1): 53–64.Order AnguilliformesMcCosker, J.E. 1972. Two new genera and two new species of western Pacific snake-eels (Apodes: Ophichthidae).

Proc. Cal. Acad. Sci. 39(10): 111–119.McCosker, J.E. and J.E. Randall. 2001. Revision of the snake-eel genus Brachysomophis (Anguilliformes: Ophichthidae),

with descriptions of two new species and comments on the species of Mystriophis. Indo-Pacific Fishes 33: 1–32, 2 pls.

Böhlke, E.B. 2000. Notes on the identity of small, brown, unpatterned Indo-Pacific moray eels, with descriptions ofthree new species (Anguilliformes: Muraenidae). Pac. Sci. 54(4): 395–416.

Böhlke, E.B. and J.E. Randall. 2000. A review of the moray eels (Anguilliformes; Muraenidae) of the Hawaiian Islands,with descriptions of two new species. Proc. Acad. nat. Sci. Philad. 150: 203–278, 9 pls.

Order ClupeiformesWhitehead, P.J.P. 1985. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated

and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies, and wolf-herrings. Part1 – Chirocentridae, Clupeidae and Pristigasteridae. FAO Species Synopsis No. 125, Vol. 7, Pt. 1: 1–303.

Whitehead, P.J.P., G.J. Nelson and T. Wongratana. 1988. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world(suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats,shads, anchovies, and wolf-herrings. Part 2 – Engraulididae. FAO Species Synopsis No. 125, Vol. 7, Pt. 2: 305–579.

Order OphidiiformesMarkle, D.F. and J.E. Olney. 1990. Systematics of the pearlfishes (Pisces: Carapidae). Bull. Mar. Sci. 47(2): 269–410.Møller, P.R. and W. Schwarzhans. 2008. Review of the Dinematichthyini (Teleostei: Bythitidae) of the Indo-west

Pacific. Part IV. Dinematichthys and two new genera with descriptions of nine new species. The Beagle,Records of the Museum of Art Galleries of the Northern Territory 24: 87–146.

Nielsen, J.G. and Y. Machida. 1988. Revision of the Indo-Pacific bathyal fish genus Glyptophidium (Ophidiiformes,Ophidiidae). Jap. J. Ichthyol. 35(3): 289–319.

Order LophiiformesBradbury, M.G. 1988. Rare fishes of the deep-sea genus Halieutopsis: a review with descriptions of four new species

(Lophiiformes: Ogcocephalidae). Fieldiana, Zool. 44: 1–22.Order AtheriniformesKimura, S., D. Golani, Y. Iwatsuki, M. Tabuchi and T. Yoshino. 2007. Redescriptions of the Indo-Pacific atherinid fishes

Atherinomorus forskalii, Atherinomorus lacunosus, and Atherinomorus pinguis. Ichthyol. Res. 54(2): 145–159.

Parenti, L.R. 1989. A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proc. Cal.Acad. Sci. 46(11): 243–277.

Order Beloniformes

73The fishes of southwestern Thailand

Parenti, L.R. 2008. A phylogenetic analysis and taxonomic revision of ricefishes, Oryzias and relatives (Beloniformes,Andrianichthyidae). Zool. J. Linn. Soc. 154: 494–610.

Order BeryciformesRandall, J.E. 1998. Revision of the Indo-Pacific squirrelfishes (Beryciformes: Holocentridae: Holocentrinae) of the

genus Sargocentron, with descriptions of four new species. Indo-Pacific Fishes 27: 1–105, 11 pls.Order GasterosteiformesDawson, C.E. 1985. Indo-Pacific Pipefishes (Red Sea to the Americas). The Gulf Coast Research Laboratory, Ocean

Springs, Mississippi. 230 p.Order ScorpaeniformesImamura, H. 1996. Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes). Species Diversity

1(2): 123–233.Knapp, L.W. and T. Wongratana. 1987. Sorsogona melanoptera, a new flathead fish from the northern Indian Ocean

(Teleostei: Platycephalidae). Proc. Biol. Soc. Wash. 100(2): 381–385.Poss, S.G. and W.N. Eschmeyer. 1980. Xenaploactis, a new genus for Prosopodasys asperrimus Günther (Pisces:

Aploactinidae), with descriptions of two new species. Proc. Cal. Acad. Sci. 42(8): 287–293.Randall, J.E. and W.N. Eschmeyer. 2001. Revision of the Indo-Pacific scorpionfish genus Scorpaenopsis, with descriptions

of eight new species. Indo-Pacific Fishes 34: 1–79, 12 pls.Wongratana, T. 1975. A record of a flathead fish, Thysanophrys papillolabium Schultz, in the Andaman Sea, with a key

to fourteen species of the Platycephalidae from Thai waters (Pisces: Platycephalidae). Phuket mar. biol. Cent.Res. Bull. 7: 1–9.

Order PerciformesFamily AcropomatidaeOkamoto, M. and H. Ida. 2002. Acropoma argentistigma, a new species from the Andaman Sea, off southern Thailand

(Perciformes: Acropomatidae). Ichthyol. Res. 49(3): 281–285.Family SerranidaeRandall, J.E. and C.C. Baldwin. 1997. Revision of the serranid fishes of the subtribe Pseudogrammina, with descriptions

of five new species. Indo-Pacific Fishes 26: 1–56, 1 pl.Randall, J.E. and P.C. Heemstra. 1991. Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with

descriptions of five new species. Indo-Pacific Fishes 20: 1–332, 41 pls.Randall, J.E. and P.C. Heemstra. 2007. Meganthias filiferus, a new species of anthiine fish (Perciformes: Serranidae),

from the Andaman Sea off southwestern Thailand. Phuket mar. biol. Cent. Res. Bull. 68: 5–9.Randall, J.E. and R. Lubbock. 1981. A revision of the serranid fishes of the subgenus Mirolabrichthys (Anthiinae:

Anthias), with descriptions of five new species. Contr. Sci. 333: 1–27.Randall, J.E. and U. Satapoomin. 2000. Cephalopholis polyspila, a new species of grouper (Perciformes: Serranidae:

Epinephelinae) from southwestern Thailand and Sumatra. Phuket mar. biol. Cent. Res. Bull. 63: 1–7.Family PseudochromidaeGill, A.C. 2004. Revision of the Indo-Pacific dottyback fish subfamily Pseudochrominae (Perciformes: Pseudochromidae).

Smithiana Monograph 1. 213 p, 12 pls.Winterbottom, R. and J.E. Randall. 1994. Two new species of congrogadins (Teleostei: Pseudochromidae), with range

extensions for four other species. Can. J. Zool. 72: 750–756.Family PlesiopidaeMooi, R.D. 1995. Revision, phylogeny, and discussion of biology and biogeography of the fish genus Plesiops

(Perciformes: Plesiopidae). Royal Ontario Museum Life Sciences Contributions No. 159. iv + 107 p.Family OpistognathidaeSmith-Vaniz, W.F. 2009. Three new species of Indo-Pacific jawfishes (Opistognathus: Opistognathidae), with the

posterior end of the upper jaw produced as a thin flexible lamina. Aqua, Int. J. Ichthyol. 15(2): 69–108.Family ApogonidaeGon, O. 1996. Revision of the cardinalfish subgenus Jaydia (Perciformes, Apogonidae, Apogon). Trans. Roy. Soc. S. Afr.

51: 147–194.Gon, O. 2000. The taxonomic status of the cardinalfish species Apogon niger, A. nigripinnis, A. pharaonis, A. sialis, and

related species (Perciformes: Apogonidae). J.L.B. Smith Inst. Ichthyol. Spec. Publ. 65: 1–20.Gon, O. and J.E. Randall. 2003. Revision of the Indo-Pacific cardinalfish genus Archamia (Perciformes: Apogonidae),

with description of a new species. Indo-Pacific Fishes 35: 1–49, 3 pls.Fraser, T.H. 2005. A review of the species in the Apogon fasciatus group with a description of a new species of

cardinalfish from the Indo-west Pacific (Perciformes: Apogonidae). Zootaxa 924: 1–30.Fraser, T.H. 2008. Cardinafishes of the genus Nectamia (Apogonidae, Perciformes) from the Indo-Pacific region with

descriptions of four new species. Zootaxa 1691: 1–52.

Phuket mar. biol. Cent. Res. Bull.74

Fraser, T.H. and E.A. Lachner. 1985. A revision of the cardinalfish subgenera Pristiapogon and Zoramia (Genus Apogon)of the Indo-Pacific region (Teleostei: Apogonidae). Smithson. Contr. Zool. 42. 47 p.

Randall, J.E., T.H. Fraser and E.A. Lachner. 1990. On the validity of the Indo-Pacific cardinalfishes Apogon aureus(Lacepède) and A. fleurieu (Lacepède), with description of a related new species from the Red Sea. Proc. Biol.Soc. Wash. 103(1): 39–62.

Family SillaginidaeMcKay, R.J. 1985. A revision of the fishes of the family Sillaginidae. Mem. Qd. Mus. 22(1): 1–73.Family LeiognathidaeKimura, S., T. Ito, T. Peristiwady, Y. Iwatsuki and T. Yoshino. 2005. The Leiognathus splendens complex (Perciformes:

Leiognathidae) with the description of a new species, Leiognathus kupanensis Kimura and Peristiwady.Ichthyol. Res. 52(3): 275–291.

Kimura, S., R. Kimura and K. Ikejima. 2008. Revision of the genus Nuchequula with descriptions of three new species(Perciformes: Leiognathidae). Ichthyol. Res. 55(1): 22–42.

Kimura, S., T. Yamashita and Y. Iwatsuki. 2000. A new species, Gazza rhombea, from the Indo-west Pacific, with aredescription of G. achlamys Jordan & Starks, 1917 (Perciformes: Leiognathidae). Ichthyol. Res. 47(1): 1–12.

Mochizuki, K. and M. Hayashi. 1989. Revision of the leiognathid fishes of the genus Secutor, with two new species. Sci.Rept. Yokosuka City Mus. 37: 83–95.

Yamashita, T., S. Kimura and Y. Iwatsuki. 1998. Validity of the leiognathid fish, Gazza dentex (Valenciennes in Cuvier &Valenciennes, 1835), with designation of a lectotype, and redescription of G. minuta (Bloch, 1795). Ichthyol.Res. 45(3): 271–280.

Family LutjanidaeAllen, G.R. and F.H. Talbot. 1985. Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-

Pacific, with description of a new species. Indo-Pacific Fishes 11: 1–87, 10 pls.Family CaesionidaeCarpenter, K.E. 1987. Revision of the Indo-Pacific fish family Caesionidae (Lutjanoidea), with descriptions of five new

species. Indo-Pacific Fishes 15: 1–56, 7 pls.Family GerreidaeIwatsuki, Y. and P.C. Heemstra. 2001. Gerres phaiya: a new species of gerreid fish (Teleostei: Perciformes: Gerreidae)

from India, with comments on Gerres poieti and the Gerres erythrourus complex. Copeia 2001(4): 1043–1049.

Iwatsuki, Y., S. Kimura, H. Kishimoto and T. Yoshino. 1996. Validity of the gerreid fish, Gerres macracanthus Bleeker,1854, with designation of lectotype, and designation of a neotype for G. filamentosus Cuvier, 1829. Ichthyol.Res. 43(4): 417–429.

Iwatsuki, Y., S. Kimura and T. Yoshino. 1999a. Description of Gerres chrysops sp. nov. from Thailand and redescriptionof Gerres setifer (Hamilton, 1822) and G. decacanthus (Bleeker, 1865) (Perciformes: Gerreidae). Ichthyol.Res. 46(1): 27–41.

Iwatsuki, Y., S. Kimura and T. Yoshino. 1999b. Redescription of Gerres baconensis (Evermann & Seale, 1907), G.equulus Temminck & Schlegel, 1844 and G. oyena (Forsskål, 1775), included in the “G. oyena complex”, withnotes on other related species (Perciformes: Gerreidae). Ichthyol. Res. 46(4): 377–395.

Iwatsuki, Y., S. Kimura and T. Yoshino. 2001. Redescription of Gerres longirostris (Lacepède, 1801) and Gerresoblongus Cuvier in Cuvier and Valenciennes, 1830, included in the Gerres longirostris complex (Perciformes:Gerreidae). Copeia 2001(4): 954–965.

Iwatsuki, Y., S. Kimura and T. Yoshino. 2007. A review of the Gerres subfasciatus complex from the Indo-West Pacific,with three new species (Perciformes: Gerreidae). Ichthyol. Res. 54(2): 168–185.

Family HaemulidaeJohnson, J.W., J.E. Randall and S.F. Chenoweth. 2001. Diagramma melanacrum, new species of haemulid fish from

Indonesia, Borneo and the Philippines with a generic review. Mem. Qd. Mus. 46(2): 657–676.Iwatsuki, Y. and B.C. Russell. 2006. Revision of the genus Hapalogenys (Teleostei: Perciformes) with two new species

from Indo-West Pacific. Mem. Mus. Vict. 63(1): 29–46.Iwatsuki, Y., U. Satapoomin and K. Amaoka. 2000. New species: Hapalogenys merguiensis (Teleostei; Perciformes)

from Andaman Sea. Copeia 2000(1): 129–139.McKay, R.J. and U. Satapoomin. 1994. Pomadasys andamanensis, a new species of haemulid fish from Thailand.

Phuket mar. biol. Cent. Res. Bull. 59: 1–4.Satapoomin, U and J.E. Randall. 2000. Plectorhinchus macrospilus, a new species of sweetlip (Perciformes: Haemulidae)

from the southwestern Thailand, Andaman Sea. Phuket mar. biol. Cent. Res. Bull. 63: 9–16.Family Nemipteridae

75The fishes of southwestern Thailand

Russell, B.C. 1986. Review of the western Indian Ocean species of Nemipterus Swainson 1839, with description of anew species. Senckenberg. biol. 67(1/3): 19–35.

Russell, B.C. 1991. On the validity of Nemipterus furcosus (Valenciennes) (Nemipteridae). Cybium 15(1): 35–41.Russell, B.C. and D. Golani. 1993. A review of the fish genus Parascolopsis (Nemipteridae) of the western Indian Ocean,

with description of a new species from the northern Red Sea. Israel J. Zool. 39: 337–347.Family PolynemidaeFeltes, R.M. 1991. Revision of the polynemid fish genus Filimanus, with description of two new species. Copeia

1991(2): 302–322.Motomura, H. 2002. Revision of the Indo-Pacific threadfin genus Polydactylus (Perciformes, Polynemidae) with a key

to the species. Bull. Natl. Sci. Mus., Ser. A. 28(3): 171–194.Motomura, H., Y. Iwatsuki, S. Kimura and T. Yoshino. 2001. A new species, Polydactylus siamensis, from Thailand and

redescription of P. plebeius (Broussonet, 1782) with designation of a neotype (Perciformes: Polynemidae).Ichthyol. Res. 48(2): 117–126.

Motomura, H., Y. Iwatsuki and T. Yoshino. 2002. Revision of the Indo-Pacific polynemid fish genus Eleutheronema(Teleostei: Perciformes). Ichthyol. Res. 49(1): 47–61.

Motomura, H., U. Satapoomin and Y. Iwatsuki. 2000. A new record of the threadfin, Filimanus perplexa Feltes, 1991(Perciformes: Polynemidae), from the Andaman Sea, Thailand. Phuket mar. biol. Cent. Res. Bull. 63: 17–20.

Family SciaenidaeSasaki, K. 1992. Two new and two resurrected species of sciaenid genus Johnius (Johnius) from the west Pacific. Jap.

J. Ichthyol. 39(3): 191–199.Sasaki, K. 1996. Sciaenid fishes of the Indian Ocean (Teleostei, Perciformes). Mem. Fac. Sci. Kochi Univ. Ser. D (Biol.)

16/17: 83–95.Family MullidaeRandall, J.E. 2004. Revision of the goatfish genus Parupeneus (Perciformes: Mullidae), with descriptions of two new

species. Indo-Pacific Fishes 36: 1–64, 16 pls.Family KuhliidaeRandall, J.E. and H.A. Randall. 2001. Review of the fishes of the genus Kuhlia (Perciformes: Kuhliidae) of the central

Pacific. Pac. Sci. 55(3): 227–256.Family PomacentridaeAllen, G.R. 1991. Damselfishes of the World. Mergus Publishers, Melle, Germany. 271 p.Allen, G.R. and A.R. Emery. 1985. A review of the pomacentrid fishes of the genus Stegastes from the Indo-Pacific, with

descriptions of two new species. Indo-Pacific Fishes 3: 1–31, 3 pls.Allen, G.R. and M.V. Erdmann. 2005. Chromis xouthos, a new species of damselfish (Pomacentridae) from the east

Andaman Sea and central Indian Ocean. Aqua, J. Ichthyol. Aquat. Biol. 10(3): 89–94.Allen, G.R. and J.E. Randall. 2002. A review of the leucogaster species complex of the Indo-Pacific pomacentrid genus

Amblyglyphidodon, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 5(4): 139–152.Randall, J.E. and G.R. Allen. 2005. Neopomacentrus sororius, a new species of damselfish from the Indian Ocean, with

description of a neotype for its sister species, N. azysron (Bleeker). Aqua, J. Ichthyol. Aquat. Biol. 10(2): 73–80.

Family LabridaeGomon, M.F. 2006. A revision of the labrid fish genus Bodianus with descriptions of eight new species. Rec. Aust. Mus.

Suppl. 30: 1–133.Randall, J.E. 1996. Second revision of the labrid fish genus Leptojulis, with descriptions of two new species. Indo-

Pacific Fishes. 24: 1–20, 3 pls.Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Pseudocheilinus, with descriptions of three

new species. Indo-Pacific Fishes. 28: 1–34, 2 pls.Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Coris, with descriptions of five new species.

Indo-Pacific Fishes. 29: 1–74, 22 pls.Randall, J.E. 2000. Revision of the Indo-Pacific labrid fishes of the genus Stethojulis, with descriptions of two new

species. Indo-Pacific Fishes. 31: 1–42, 6 pls.Randall, J.E. and Lubbock, R. 1981. Labrid fishes of the genus Paracheilinus, with descriptions of three new species

from the Philippines. Jap. J. Ichthyol. 28(1): 19–30, 2 pls.Randall, J.E. and M.M. Smith. 1982. A review of the labrid fishes of the genus Halichoeres of the western Indian Ocean,

with descriptions of six new species. Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 45: 1–26, 6 pls.Family ScaridaeSatapoomin, U., R.H. Kuiter and J.E. Randall. 1994. First Record of the parrotfish Scarus viridifucatus from Thailand

(the Andaman Sea) and Indonesia. Phuket mar. biol. Cent. Res. Bull. 59: 5–9.

Phuket mar. biol. Cent. Res. Bull.76

Westneat, M., U. Satapoomin and J.E. Randall. 2007. Scarus maculipinna, a new species of parrotfish (Perciformes,Scaridae) from eastern Indian Ocean. Zootaxa. 1628: 59–68.

Family AmmodytidaeIda, H., P. Sirimontaporn and S. Monkolprasit. 1994. Comparative morphology of the fishes of the family Ammodytidae,

with description of two new genera and two new species. Zool. Stud. 33(4): 251–277.Family TripterygiidaeHolleman, W. 2005. A review of the triple fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the western Indain

Ocean, with descriptions of four new species. Smithiana Bulletin. 5: 1–25, 2 pls.Holleman, W. 2006. Fishes of the Helcogramma steinitzi species group (Blennioidei: Tripterygiidae) from the Indain

Ocean, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 11(3): 89–104.Williams, J.T. and J.C. Howe. 2003. Seven new species of the triplefin genus Helcogramma (Tripterygiidae) from the

Indo-Pacific. Aqua, J. Ichthyol. Aquat. Biol. 7(4): 151–176.Family BlenniidaeBath, H. 1992. Revision der gattung Praealticus Schultz & Chapman 1960 (Pisces: Blenniidae). Senckenberg. biol. 72(4/

6): 237–316.Carlson, B.A. 1980. A new Indo-Pacific fish of the genus Cirripectes (Blenniidae, Salariini). Pac. Sci. 34(4): 407–414.Smith-Vaniz, W.F. 1976. The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). The Academy of Natural

Sciences of Philadelphia. Monograph 19. vii + 196 p.Smith-Vaniz, W.F., U. Satapoomin and G.R. Allen. 2001. Meiacanthus urostigma, a new fangblenny from the northeastern

Indian Ocean, with discussion and examples of mimicry in species of Meiacanthus (Teleostei: Blenniidae:Nemophini). Aqua, J. Ichthyol. Aquat. Biol. 5(1): 25–43.

Springer, V.G. 1988. The Indo-Pacific blenniid fish genus Ecsenius. Smithson. Contr. Zool. 465. 134 p., 14 pls.Springer, V.G. and M.F. Gomon. 1975. Revision of the blenniid fish genus Omobranchus with descriptions of three new

species and notes on other species of the tribe Omobranchini. Smithson. Contr. Zool. 177. 135 p.Springer, V.G. and J.E. Randall. 1999. Ecsenius polystictus, new species of blenniid fish from Mentawai Islands,

Indonesia, with notes on other species of Ecsenius. Revue fr. Aquariol. 26(1–2): 39–48.Springer, V.G. and J.T. Williams. 1994. The Indo-west Pacific blenniid fish genus Istiblennius reappraised: A revision of

Istiblennius, Blenniella, and Paralticus, new genus. Smithson. Contr. Zool. 565. 193 p.Williams, J.T. 1988. Revision and phylogenetic relationships of the blenniid fish genus Cirripectes. Indo-Pacific Fishes.

17: 1–78, 7 pls.Family CallionymidaeFricke, R. 1983. Revision of the Indo-Pacific genera and species of the dragonet family Callionymidae. Theses Zoologicae

Vol. 3. 774 p.Family GobiidaeAkihito and Meguro, K. 2000. Review of the Gobiid genus Cristatogobius found in Japan with description of a new

species. Ichthyol. Res. 47(3): 249–261.Darumas, U. and P. Tantichodok. 2002. A new species of mudskipper (Gobiidae: Oxucercinae) from southern Thailand.

Phuket mar. biol. Cent. Res. Bull. 64: 101–107.Hoese, D.F. and H.K. Larson. 1994. Revision of the Indo-Pacific gobiid fish genus Valenciennea, with descriptions of

seven new species. Indo-Pacific Fishes. 23: 1–71, 6 pls.Lachner, E.A. and S.J. Karnella. 1980. Fishes of the Indo-Pacific fish genus Eviota with descriptions of eight new species

(Teleostei: Gobiidae). Smithson. Contr. Zool. 315. 127 p.Lachner, E.A. and J.F. McKinney. 1978. A revision of the Indo-Pacific fish genus Gobiopsis with descriptions of four

new species (Pisces: Gobiidae). Smithson. Contr. Zool. 262. 52 p.Larson, H.K. 1999. A review of the mangrove goby genus Hemigobius (Gobioidei, Gobiidae, Gobionellinae). The Beagle,

Records of the Museums and Art Galleries of the Northern Territory. 15: 23–42.Larson, H.K. 1999. Allocation to Calamiana and redescription of the fish species Apocryptes variegates and Vaimosa

mindora (Gobioidei: Gobiidae: Gobionellinae), with description of a new species. Raffles Bull. Zool. 47(1):257–281.

Larson, H.K. 2001. A revision of the gobiid fish genus Mugilogobius (Teleostei: Gobioidei). Rec. West. Aust. Mus.Suppl. No. 62: 1–233.

Larson, H.K. 2005. A revision of the gobiid genus Stigmatogobius (Teleostei: Gobiidae), with descriptions of two newspecies. Ichthyol. Explor. Freshwaters. 16(4): 347–370.

Larson, H.K. 2009. Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae:Gobionellinae), with descriptions of three new species. Raffles Bull. Zool. 57(1): 127–181.

Murdy, E.O. 1985. A review of the gobiid fish genera Exyrias and Macrodontogobius, with description of a new speciesof Exyrias. Indo-Pacific Fishes. 10: 1–14, 2 pls.

77The fishes of southwestern Thailand

Murdy, E.O. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Rec.Aust. Mus. Suppl. 11: 1–93.

Murdy, E.O. and D.F. Hoese. 1985. Revision of the gobiid fish genus Istigobius. Indo-Pacific Fishes 4: 1–41, 3 pls.Polunin, N.V.C. and R. Lubbock. 1979. Five new prawn-associated gobies (Teleostei: Gobiidae) of the genus Amblyeleotris.

Bull. Br. Mus. Nat. Hist. (Zool.). 36(4): 239–249.Randall, J.E. 2001. Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zool. Stud. 40(3): 206–225.Randall, J.E. and D.W. Greenfield. 2001. A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis.

Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 69: 1–17, 2 pls.Randall, J.E. and D.F. Hoese. 1985. Revision of the Indo-Pacific dartfishes, genus Ptereleotris (Perciformes: Goioidei)

Indo-Pacific Fishes. 7: 1–36, 4 pls.Shibukawa, K. and G.R. Allen. 2007. Review of the cheek-spine goby genus Gladiogobius (Actinopterigii, Perciformes,

Gobiidae), with descriptions of two new species from the Indo-Pacific. Bull. Natl. Mus. Nat. Sci., Ser. A,33(4): 193–206.

Shibukawa, K. and U. Satapoomin. 2006. Myersina adonis, a new species of shrimp-associated goby (Pisces: Perciformes:Gobiidae) from the Andaman Sea. Bull. Natl. Sci. Mus., Ser. A, 32(1): 29–37.

Shibukawa, K., T. Suzuki, H. Senou and K. Yano. 2005. Records of three shrimp-goby species (Teleostei, Perciformes,Gobiidae) from the Ryukyu Archipelago, Japan. Bull. Natl. Sci. Mus., Ser. A, 31(4): 191–204.

Satapoomin, U. and R. Winterbottom. 2002. Redescription of the gobioid fish Cryptocentrus pavoninoides (Bleeker,1849), with notes on sexual dichromatism in shrimp gobies. Aqua, J. Ichthyol. Aquat. Biol. 5(2): 53–64.

Winterbottom, R. 2002. A redescription of Cryptocentrus crocatus Wongratana, a redefinition of Myersina Herre(Acanthopterygii; Gobiidae), a key to the species, and comments on relationships. Ichthyol. Res. 49(1): 69–75.

Winterbottom, R. and M. Burridge. 1993. Revision of the species of Priolepis possessing a reduced transverse patternof cheek papillae and no predorsal scales (Teleostei: Gobiidae). Can. J. Zool. 71: 494–514.

Winterbottom, R. and A.S. Harold. 2005. Gobiodon prolixus, a new species of gobiid fish (Teleostei: Perciformes:Gobiidae) from the Indo-west Pacific. Proc. Biol. Soc. Wash. 118(3): 582–589.

Winterbottom, R. and L. Southcott. 2007. Two new species of the genus Trimma (Percomorpha: Gobiidae) from westernThailand. Aqua, Int. J. Ichthyol. 13(2): 69–76.

Family SiganidaeWoodland, D.J. 1990. Revision of the fish family Siganidae with descriptions of two new species and comments on

distribution and biology. Indo-Pacific Fishes. 19: 1–136, 11 pls.Family AcanthuridaeRandall, J.E. 1993. Acanthurus tristis, a valid Indian Ocean surgeonfish (Perciformes: Acanthuridae). J.L.B. Smith Inst.

Ichthyol. Spec. Publ. 54: 1–8.Randall, J.E. and K.D. Clements. 2001. Second revision of the surgeonfish genus Ctenochaetus (Perciformes:Acanthuridae),

with descriptions of two new species. Indo-Pacific Fishes. 32: 1–33, 6 pls.Family SphyraenidaeDoiuchi, R. and T. Nakabo. 2005. The Sphyraena obtusata group (Perciformes: Sphyraenidae) with a description of a

new species from southern Japan. Ichthyol. Res. 52(2): 132–151Order PleuronectiformesMenon, A.G.K. 1977. A systematic monograph of the tongue soles of the genus Cynoglossus Hamilton-Buchanan

(Pisces: Cynoglossidae). Smithson. Contr. Zool. 238. 129 p.Randall, J.E. and M. Desoutter-Meniger. 2007. Review of the soles of the genus Aseraggodes (Pleuronectiformes:

Soleidae) from the Indo-Malayan region, with descriptions of nine new species. Cybium. 31(3): 301–331.Randall, J.E. and J.W. Johnson. 2007. Revision of the soleid fish genus Pardachirus. Indo-Pacific Fishes. 39: 1–22, 4 pls.Order TetraodontiformesHutchins, J.B. 1997. Review of the monacanthid fish genus Paramonacanthus, with descriptions of three new species.

Rec. West. Aust. Mus. Suppl. No. 54. 57 p.Leis, J.M. 2006. Nomenclature and distribution of the species of the porcupinefish family Diodontidae. Mem. Mus.

Vict. 63(1): 77–90.Randall, J.E. and W. Klausewitz. 1973. A review of the triggerfish genus Melichthys, with description of a new species

from the Indian Ocean. Senckenberg. biol. 54(1/3): 57–69.

Phuket mar. biol. Cent. Res. Bull.78