THE SCANDINAVIAN BROWN BEAR RESEARCH PROJECTPublications to 23 February 2016

Summary:A) Publications in peer-reviewed journals 201 in press 5B) Books and book chapters 11C) Papers in nonrefereed publications 22D) Completed student theses 109E) Rapports to management agencies, etc. 134F) Popular publications 86entries in total 563

A) Publications in peer-reviewed journalsA1 Bjärvall, A. and F. Sandegren. 1987. Early experiences with the first radio-marked

bears in Sweden. Proceedings of the International Conference on Bear Research and Management 7:9-12.

A2 Bjärvall, A. 1990. The brown bear in Sweden. Aquilo Ser. Zool. 27:17-19A3 Bjärvall, A., F. Sandegren, and P. Wabakken. 1990. Large home ranges and possible

early sexual maturity in Scandinavian bears. Proceedings of the International Conference on Bear Research and Management 8:237-241.

A4 Swenson, J. E., F. Sandegren, A. Bjärvall, A. Söderberg, P. Wabakken, and R. Franzén. 1994. Size, trend, distribution and conservation of the brown bear Ursus arctos population in Sweden. Biological Conservation 70:9-17.

A5 Swenson, J. E., P. Wabakken, F. Sandegren, A. Bjärvall, R. Franzén, and A. Söderberg. 1995. The near extinction and recovery of brown bears in Scandinavia in relation to the bear management policies of Norway and Sweden. Wildlife Biology 1:11-25.

A6 Taberlet, P., J. E. Swenson, F. Sandegren, and A. Bjärvall. 1995. Localization of a contact zone between two highly divrgent mitochondrial DNA lineages of the brown bear Ursus arctos in Scandinavia. Conservation Biology 9:1255-1261.

A7 Swenson, J. E., F. Sandegren, A. Söderberg, and R. Franzén. 1995. Estimating the total weight of Scandinavian brown bears from field-dressed and slaughter weights. Wildlife Biology 1:177-179.

A8 Swenson, J. E. and F. Sandegren. 1996. Sustainable brown bear harvest in Sweden estimated from hunter-provided information. J. Wildl. Res. 1:228-231.

A9 Swenson, J. E. and S. Wikan. 1996. A population estimate for brown bears in Finnmark County, North Norway. Fauna norvegica Ser. A. 17:11-15.

A10 Sagør, J. T., J. E. Swenson, and E. Røskaft. 1997. Compatibility of brown bear Ursus arctos and free-ranging sheep in Norway. Biological Conservation 81:91-95.

A11 Swenson, J. E., F. Sandegren, A. Söderberg, A. Bjärvall, R. Franzén, and P. Wabakken. 1997. Infanticide caused by hunting of male bears. Nature 386:450-451.

A12 Swenson, J. E., F. Sandegren, S. Brunberg, and P. Wabakken. 1997. Winter den sites abandonment by brown bears Ursus arctos: causes and consequences. Wildlife Biology 3:35-38.

A13 Linnell, J. D. C., R. Aanes, J. E. Swenson, J. Odden, and M. E. Smith. 1997. Translocation of carnivores as a method for managing problem animals: a review. Biodiversity and Conserv. 6:1245-1257.

A14 Swenson, J. E., R. Franzén, P. Segerström, and F. Sandegren. 1998. The age of self-sufficiency in Scandinavian brown bears. Acta Theriologica 43:213-218.

A15 Swenson, J. E., F. Sandegren, A. Bjärvall, and P. Wabakken. 1998. Living with success: research needs for an expanding brown bear population. Ursus 10:17-24.

A16 Swenson, J. E., F. Sandegren, and A. Söderberg. 1998. Geographic expansion of an increasing brown bear population: evidence for presaturation dispersal. Journal of Animal Ecology 67:819-826.

A17 Dahle, B., O. J. Sørensen, E. H. Wedul, J. E. Swenson and F. Sandegren. 1998. The diet of brown bears Ursus arctos in central Scandinavia: effect of access to free-ranging domestic sheep Ovis aries. Wildlife Biology 4:147-158.

A18 Sæther, B. E., S. Engen, J. E. Swenson, Ø. Bakke, and F. Sandegren. 1998. Assessing the viability of Scandinavian brown bear, Ursus arctos, populations: the effects of uncertain parameter estimates. Oikos 83:403-416.

A19 Tryland, M, T. Sandvik, J. M. Arnemo, G. Stuve, Ø. Olsvik, and T. Traavik. 1998. Antibodies against orthopoxviruses in wild carnivores from Fennoscandia. Journal of Wildlife Diseases 34:443-450.

A20 Swenson, J. E., K. Wallin, G. Ericsson, G. Cederlund, and F. Sandegren. 1999. Effects of ear tagging on survival of moose calves. Journal of Wildlife Management 63:354-358.

A21 Swenson, J. E., F. Sandegren, A. Söderberg, M. Heim, O. J. Sørensen, A. Bjärvall, R. Franzén, S. Wikan, and P. Wabakken. 1999. Interactions between brown bears and humans in Scandinavia. Biosphere Conservation 2:1-9.

A22 Tufto, J., B.-E. Sæther, S. Engen, J. E. Swenson & F. Sandegren. 1999. Harvesting strategies for conserving minimum viable populations based on World Conservation Union criteria: brown bears in Norway. Proceedings of the Royal Society of London, Series B 266:961-967.

A23 Swenson, J. E., A. Jansson., R. Riig, and F. Sandegren. 1999. Bears and ants: myrmecophagy by brown bears in central Scandinavia. Canadian Journal of Zoology 77:551-561.

A24 Linnell, J. C. D., J. Odden, M. E. Smith, R. Aanes, and J. E. Swenson. 1999. Large carnivores that kill livestock: do «problem animals» really exist? Wildlife Society Bulletin 27:698-705.

A25 Matson, G. M., H. E. Casquilho-Gray, J. D. Paynich, V. G. Barnes, Jr., H. V. Reynolds III, and J. E. Swenson. 1999. Cementum annuli are unreliable reproductive indicators in female brown bears. Ursus 11:275-280.

A26 Swenson, J. E. 1999. Does hunting affect the behavior of brown bears in Eurasia? Ursus 11:157-162.

A27 Linnell, J. D. C., B. Barnes, J. E. Swenson, and R. Andersen. 2000. How vulnerable are denning bears to disturbance? Wildlife Society Bulletin 28:400-413

A28 Waits, L., P. Taberlet, J. E. Swenson, F. Sandegren, and R. Franzén. 2000. Nuclear DNA microsatellite analysis of genetic diversity and gene flow in the Scandinavian brown bear (Ursus arctos). Molecular Ecology 9:421-431.

A29 Linnell, J. D. C., J. E. Swenson, and R. Andersen. 2000. Conservation of biodiversity in Scandinavian boreal forests: large carnivores as flagships, umbrellas, indicators, or keystones? Biodiversity and Conservation 9: 857-868

A30 Smith, M. E., J. D. C. Linnell, J. Odden, and J. E. Swenson. 2000. Review of Methods to reduce livestock depredation: I. Guardian animals. Acta Agriculturæ Scandinavica Section A, Animal Science 50: 279-290.

A31 Smith, M. E., J. D. C. Linnell, J. Odden, and J. E. Swenson. 2000. Review of Methods to reduce livestock depredation: II. Aversive conditioning, deterrents and repellents. Acta Agriculturæ Scandinavica Section A, Animal Science 50: 304-315.

A32 Berger, J., J. E. Swenson, and I.-L. Persson. 2001. Recolonizing carnivores and naïve prey: conservation lessons from Pleistocene extinctions. Science 291: 1036-1039.

A33 Persson, I.-L., S. Wikan, J. E. Swenson, and I. Mysterud. 2001. The diet of the brown bear in the Pasvik, Valley, northeastern Norway. Wildlife Biology 7: 27-37.

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A34 Linnell, J. D. C., J. E. Swenson, and R. Andersen. 2001. Predators and people: conservation of large carnivores is possible at high human densities if management policy is favourable. Animal Conservation 4: 345-349.

A35 Friebe, A., J. E. Swenson, and F. Sandegren. 2001. Denning chronology of female brown bears (Ursus arctos) in central Sweden. Ursus 12: 37-46.

A36 Swenson, J. E., F. Sandegren, S. Brunberg, and P. Segerström. 2001. Factors associated with loss of brown bear cubs in Sweden. Ursus 12: 69-80.

A37 Zedrosser, A., B. Dahle, J. E. Swenson, and N. Gerstl. 2001. Status and management of the brown bear in Europe. Ursus 12: 9-20.

A38 Swenson, J. E., B. Dahle, and F. Sandegren. 2001. Intraspecific predation in Scandinavian brown bears older than cubs-of-the-year. Ursus 12: 81-92.

A39 Waits, L., G. Luikart, and P. Taberlet. 2001. Estimating the probability of identity among genotypes in natural populations: cautions and guidelines. Molecular Ecology 10:249-256.

A40 Cercueil, A., Bellemain, E., and Manel, S. 2002. PARENTE: A software package for parentage analysis. Journal of Heredity 93: 458-459.

A41 Dahle, B. and J. E. Swenson. 2003. Family break-up in brown bears: are young forced to leave? Journal of Mammalogy 84:536-540.

A42 Dahle B, and J. E. Swenson. 2003. Seasonal range size in relation to reproductive strategies in brown bears Ursus arctos. Journal of Animal Ecology 72:660-667.

A43 Dahle, B., and J. E. Swenson. 2003. Home ranges in adult Scandinavian brown bears Ursus arctos: effect of mass, sex, reproductive category, population density and habitat type. Journal of Zoology, London 260:329-335.

A44 Dahle, B., and J. E. Swenson. 2003. Factors influencing length of maternal care in brown bears (Ursus arctos) and its effect on offspring. Behavioural Ecology and Sociobiology 54:352-358.

A45 Schwartz, C. C., K. A. Keating, H. V. Reynolds, III, V. G. Barnes, Jr., R. Sellers, J. E. Swenson, S. D. Miller, B. N. McLellan, J. Keay, R. McCann, M. Gibeau, W. Wakkinen, R. D. Mace, W. Kasworm, R. Smith and S. Herrero. 2003. Reproductive maturation and senescence in the brown/grizzly bear. Ursus 14:109-119.

A46 Schwartz, C. C., J. E. Swenson, and S. M. Miller. 2003. Large carnivores, moose and humans: a changing paradigm of predator management in the 21st century. Alces. 39:41-63. (Invited paper)

A47 Manel, S., E. Bellemain, J. E. Swenson and O. François. 2004. Assumed and inferred spatial structure of populations: the Scandinavian brown bears revisited. Molecular Ecology 13:1327-1331.

A48 Piggott, M. P., E. Bellemain, P. Taberlet and A. C. Taylor. 2004. A multiplex pre-amplification method that significantly improves microsatellite amplification and error rates for faecal DNA in limiting conditions. Conservation Genetics 5:417-420.

A49 Zedrosser, A., Rauer, G., and Kruckenhauser, L. 2004. Early primiparity in brown bears. Acta Theriologica 49:427-432.

A50 Bonin, A., E. Bellemain, P. Bronken Eidesen, F. Pompanon, C. Brochmann, and P. Taberlet. 2004. How to track and assess genotyping errors in population genetics studies. Molecular Ecology 13:3261-3273.

A51 Bellemain, E. and P. Taberlet. 2004. Improved noninvasive genotyping method: application to brown bear (Ursus arctos) faeces. Molecular Ecology Notes 4:519-522.

A52 Tallmon, D. A., E. Bellemain, J. E. Swenson, and P. Taberlet. 2004. Genetic monitoring of Scandianvian brown bear: effective population size and immigration. Journal of Wildlife Management 68:960-965.

A53 Bellemain, E., J. E. Swenson, D. Tallmon, S. Brunberg and P. Taberlet. 2005. Estimating population size of elusive animals using DNA from hunter-collected feces: comparing four methods for brown bears. Conservation Biology 19:150-161.

A54 Manchi, S. and J. E. Swenson. 2005. Denning behaviour of male Scandinavian brown bears (Ursus arctos). Wildlife Biology 11:123-132.

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A55 Støen, O.-G., E. Bellemain, S. Sæbø, and J. E. Swenson. 2005. Kin-related spatial structure in brown bears Ursus arctos. Behavioural Ecology and Sociobiology 59:191-197.

A56 Ågren, E., A. Söderberg and T. Mörner. 2005. Fallot’s tetralogy in a European brown bear (Ursus arctos). Journal of Wildlife Diseases 41:825-828.

A57 Zedrosser, A. and J. E. Swenson. 2005. Do brown bear litter sizes reported by the public reflect litter sizes obtained by scientific methods? Wildlife Society Bulletin. 33:1352-1356.

A58 Solberg, K. H., E. Bellemain, O.-M. Drageset, P. Taberlet and J. E. Swenson. 2006. An evaluation of field and non-invasive genetic methods to estimate brown bear (Ursus arctos) population size. Biological Conservation 128:158-168.

A59 Bellemain, E., Zedrosser, A., S. Manel, L. P. Waits, and Swenson, J. E. 2006. The dilemma of female mate selection in the brown bear, a species with sexually selected infanticide. Proceedings of the Royal Society London, Series B. 273 283 – 291.

A60 Bellemain, E., J. E. Swenson and P. Taberlet. 2006. Mating strategies in relation to sexually selected infanticide in a non-social carnivore: the brown bear. Ethology 112:238-246.

A61 Arnemo, J. M., P. Ahlqvist, R. Andersen, F. Berntsen, G. Ericsson, J. Odden, S. Brunberg, P. Segerström, and J. E. Swenson. 2006. Risk of capture-related mortality in large free-ranging mammals: experiences from Scandinavia. Wildlife Biology 12:109-113.

A62 Zedrosser, A., Dahle, B., and Swenson, J.E. 2006. Population density and food conditions determine adult female size in brown bears. Journal of Mammalogy 87: 510-518.

A63 Katajisto, J. & Moilanen, A. 2006. Kernel-based home range method for data with irregular sampling interval. Ecological Modelling 194:405-413.

A64 Støen O.-G., A. Zedrosser, S. Sæbø, and J. E. Swenson. 2006. Inversely density-dependent natal dispersal in brown bears Ursus arctos.. Oecologia 148:356-364.

A65 Dahle, B., Zedrosser, A., Swenson, J. E. 2006. Correlates with body size and mass in yearling brown bears. Journal of Zoology 269:273-283.

A66 Dahle, B., O.-G. Støen, and J. E. Swenson. 2006. Factors influencing home-range size in subadult brown bears. Journal of Mammalogy 87:859-865.

A67 Støen, O.-G., A. Zedrosser, P. Wegge and J. E. Swenson. 2006. Socially induced delayed primiparity in brown bears Ursus arctos. Behavioural Ecology and Sociobiology 61:1-8.

A68 Miquel, C, Bellemain, E., Poillot, C, Bessière, J., Durand, A. and Taberlet, P. 2006. Quality indexes to assess the reliability of genotypes in studies using non-invasive sampling and multiple-tube approach. Molecular Ecology Notes 6:985-988.

A69 Gervasi, V., S. Brunberg, and J. E. Swenson. 2006. An individual approach to measure activity levels: a test on brown bears. Wildlife Society Bulletin 34:1314-1319.

A70 Bellemain, E., Nawaz, M. A., Valentini, A. Swenson, J. E., Taberlet. 2007. Genetic tracking of the brown bear in northern Pakistan and implications for conservation. Biological Conservation 134:537-547.

A71 Zedrosser, A., Bellemain, E., Taberlet, P., and Swenson, J. E. 2007. Genetic estimates of annual reproductive success in male brown bears: the effects of body size, age internal relatedness and population density. Journal of Animal Ecology 76:368-375.

A72 Manel, S, F. Berthoud, E. Bellemain, M. Gaudeul, G. Luikart, J. E. Swenson, L. P. Waits, P. Taberlet and Intrabiodiv Consortium. 2007. A new individual-based spatial approach for identifying genetic discontinuities in natural populations. Molecular Ecology 16:2031-2043.

A73 Nellemann, C., O.-G. Støen, J. Kindberg, J. E. Swenson, I. Vistnes, G. Ericsson, J. Katajisto, B. P. Kaltenborn, J. Martin, and A. Ordiz. 2007. Terrain use by an expanding brown bear population in relation to age, recreational resorts and human settlements. Biological Conservation 138:157-165.

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A74 Swenson, J. E., M. Adamič, D. Huber, and S. Stokke. 2007. Brown bear body mass and growth in northern and southern Europe. Oecologia 153:37-47.

A75 Moe, T. F., Kindberg, J., and J. E. Swenson. 2007. The importance of diel behaviour when studying habitat selection; examples from female Scandinavian brown bears (Ursus arctos). Canadian Journal of Zoology 85:518-525.

A76 Swenson, J. E., B. Dahle, H. Busk, O. Opseth, T. Johansen, A. Söderberg, K. Wallin and G. Cederlund. 2007. Predation on moose calves by European brown bears. Journal of Wildlife Management 71:1993-1997.

A77 Nawaz, M. A. 2007. Status of the brown bear in Pakistan. Ursus 18:89-100. A78 Zedrosser, A., Støen, O.-G., Sæbø, S., and Swenson, J. E. 2007. Should I stay or should

I go? Natal dispersal in the brown bear. Animal Behaviour 74:369-376. (Featured article)

A79 Ordiz, A., C. Rodríguez, J. Naves, A. Fernández, D. Huber, P. Kazcensky, A. Mertens, Y. Mertzanis, A. Mustoni, S. Palazón, P. Y. Quenette, G. Rauer, C. Rodríguez, and J. E. Swenson. 2007. Distance-based criteria to identify minimum number of brown bear females with cubs in Europe. Ursus 18 :158-167.

A80 Fredriksson, G. M., Danielsen, L. S., and Swenson, J. E. 2007. Impacts of El Niño related drought and forest fires on sun bear fruit resources in lowland dipterocarp forest of East Borneo. Biodiversity and Conservation 16:1823-1838.

A81 Bischof, R., Swenson, J.E., Fujita, R., Zedrosser, A., and Söderberg, A. 2008. Hunting patterns, the ban on baiting, and harvest demographics of brown bears in Sweden. Journal of Wildlife Management 72:79-88. (Featured article)

A82 Bostedet, G., Ericsson, G. & Kindberg, J. 2008. Contingent values as implicit contracts – estimating minimum legal willingness to pay for conservation of large carnivores in Sweden. Environmental and Resource Economics 39:189-198.

A83 Ericsson, G., Bostedet, G. & Kindberg, J. 2008. Wolves as a symbol for people’s willingness to pay for large carnivore conservation. Society and Natural Resources 21: 21:294-309.

A84 Bischof, R., A. Mysterud, and J. E. Swenson. 2008. Should hunting mortality mimic the patterns of natural mortality? Biology Letters 4: 307-310. (Featured article)

A85 Elfström, M., Swenson, J. E., and Ball, J. P. 2008. Selection of denning habitats by Scandinavian brown bears. Wildlife Biology 14:176-187.

A86 Swenson, J. E. and Haroldson, M. A. 2008. Observations of mixed-age litters in brown bears. Ursus 19: 73-79.

A87 Jombart, T., S. Devillard, A.-B. Dufour and D. Pontier. 2008. Revealing cryptic spatial patterns in genetic variability by a new multivariate method. Heredity 101:92-103.

A88 Nawaz M.A., Swenson J.E. and Zakaria V. 2008. Pragmatic management increases a flagship species, the Himalayan brown bears, in Pakistan’s Deosai National Park. Biological Conservation 141:2230-2241.

A89 Martin, J., C. Calenge, P.-Y. Quenette, D. Allainé. 2008. Importance of movement constraints in habitat selection studies, Ecological Modeling 213:257-262.

A90 Ordiz, A., O.-G. Støen, J. E. Swenson, I.Kojola, and R. Bischof. 2008. Distance-dependent effect of the nearest neighbour: spatio-temporal patterns in brown bear reproduction. Ecology 89:3327-3335.

A91 May, R., van Dijk, J., Wabakken, P., Linnell, J. D. C., Swenson, J. E., Zimmerman, B., Odden, J., Pedersen, H. C., Andersen, R. and Landa, A. 2008. Habitat differentiation within the large-carnivore community of Norway’s multiple-use landscapes. Journal of Applied Ecology 45:1382-1391.

A92 Valentini, A, C. Miquel, M. A. Nawaz, E. Bellemain, E. Coissac, F. Pompanon, L. Gielly, C. Cruaud, G. Nascetti, P. Winker, J. E. Swenson, and P. Taberlet. 2009. New perspectives in diet analysis based on DNA barcoding and parallel pyrosequencing: the trnL approach. Molecular Ecology Resources 9:51-60.

A93 Kindberg, J., G. Ericsson, and J.E. Swenson. 2009. Monitoring rare or elusive large mammals using effort- corrected voluntary observers. Biological Conservation 142:159-165.

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A94 Strömquist, A., Å. Fahlman, J. M. Arnemo, and A. Pettersson. 2009. Dental and periodontal health in free-ranging Swedish brown bears (Ursus arctos). Journal of Comparative Pathology 141:170-176.

A95 Bischof, R., J. E. Swenson, N. G. Yoccoz, A. Mysterud, and O. Gimenez. 2009. The magnitude and demographic selectivity of natural and multiple anthropogenic mortality causes in hunted brown bears. Journal of Animal Ecology 78:656-665.

A96 Pagès, M., C. Maudet, E. Bellemain, P. Taberlet, and S. Hughes. 2009. A system for sex determination from degraded DNA: a useful tool for palaeogenetics and conservation genetics of ursids. Conservation Genetics 10:897-907.

A97 Bischof, R., A. Zedrosser, S. Brunberg, and J. E. Swenson. 2009. A note on opportunism and parsimony in data collection. Journal of Wildlife Management 73:1021-1024.

A98 Bischof, R., A. Mysterud, and J. E. Swenson. 2009. With or without equations: What are the dos and don’ts of hunting? Biology Letters 5:213.

A99 Zedrosser, A., B. Dahle, O.-G. Støen, and J. E. Swenson. 2009. The effects of primiparity on reproductive performance in the brown bear. Oecologia 160:847-854.

A100 Kübber-Heiss, A., Zedrosser, A., Rauer, G., Zenker, W., Schmidt, P. and Arnemo, J.M. 2009. Internal hydrocephalus in a wild-born brown bear cub. European Journal of Wildlife Research 55:539–542.

A101 Bischof, R and Zedrosser, A. 2009. The educated prey. Consequences for exploitation and control. Behavioral Ecology 20:1228–1235.

A102 Elfström, M. and Swenson, J. E. 2009. Effects of sex and age on den site use by Scandinavian brown bears. Ursus 20:85-93.

A103 Ordiz, A., O-G. Støen, L. G. Langebro, S. Brunberg, and J. E. Swenson. 2009. A practical method for measuring wildlife horizontal cover. Ursus 20:109-113.

A104 Nawaz, M. A., J. E. Swenson, and V. Zakaria. 2009. Recovery of brown bears in northern Pakistan. Current Conservation 3:12-13.

A105 Mysterud, A and Bischof, R. 2010. Can compensatory culling offset undesirable evolutionary consequences of trophy hunting? Journal of Animal Ecology 79:148-160.

A106 Fahlman, Å., Pringle, J., Arnemo, J. M., Swenson, J. E., Brunberg, S., and Nyman, G. 2010. Treatment of hypoxemia during anesthesia of brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 41:161-164.

A107 Fernández-Gil, A., A. Ordiz and J. Naves. 2010. Are Cantabrian brown bears recovering? Ursus 21:121-124.

A108 Fröbert, O., K. Christensen, Å. Fahlman, S. Brunberg, J. Josefsson, E. Särndahl, J. E. Swenson, J. M. Arnemo. 2010. Platelet function in brown bear (Ursus arctos) compared to man. Thrombosis Journal 8:10

A109 Martin, J., M. Basille, B. Van Moorter, J. Kindberg, D. Allainé, and J. E. Swenson. 2010. Coping with human disturbance: spatial and temporal tactics of the brown bear (Ursus arctos). Canadian Journal of Zoology 88:875-883.

A110 Karamanlidis, A. A., E. Drosopoulou, M. de Gabriel Hernando, L. Georgiadis, L. Krambokoukis, S. Pllaha, A. Zedrosser and Z. Scouras. 2010. Noninvasive genetic studies of brown bears using power poles. European Journal of Wildlife Research 56:693-702.

A111 Gandolf, A. R., Å. Fahlman, J. M. Arnemo, J. L. Dooley, and R. Hamlin. 2010. Baseline normal values and phylogenitic class of the electrocardiogram of anesthetized free-ranging brown bears (Ursus arctos). Journal of Wildlife Diseases 46:724-730.

A112 Støen, O.-G., W. Neumann, G. Ericsson, J. E. Swenson, H. Dettki, J. Kindberg, and C. Nellemann. 2010. Behavioural response of moose Alces alces and brown bears Ursus arctos to direct helicopter approach by researchers. Wildlife Biology 16: 292-300.

A113 Rosell, F., Jojola, S. M., Ingdal, K., Lassen, B.A., Swenson, J. E., Arnemo, J. M, & Zedrosser, A. 2011. Brown bears possess anal sacs and secretions may code for sex. Journal of Zoology 283:143-152.

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A114 Swenson, J. E., P. Taberlet, and E. Bellemain. 2011. Genetics and conservation of European brown bears. Mammal Review 41:87-98.

A115 Fahlman, Å., Arnemo, J. M., Swenson, J. E., Brunberg, S., Pringle, J., and Nyman, G. 2011. Physiologic evaluation of capture and anesthesia with medetomidine-zolazepam-tiletamine in brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 42:1-11.

A116 Ordiz, A., O-G. Støen, M. Delibes, and J. E. Swenson. 2011. Predators or prey? Spatio-temporal discrimination of human-derived risk by brown bears. Oecologia 166:59–67.

A117 Vestergaard, P., O.-G. Støen, J. E. Swenson, L. Mosekilde, L. Heickendorff, and O. Fröbert. 2011. Vitamin D status, bone and connective tissue turnover in brown bears (Ursus arctos) during hibernation and the active state. PLOS One 6 (6): e21483.

A118 Zedrosser, A., Steyaert, S. M.J.G., Swenson, J. E. and Gossow, H. 2011. Brown bear conservation and the ghost of persecution past. Biological Conservation 144:2163-2170.

A119_118 Sahlén, E., O.-G. Støen, and J. E. Swenson. 2011. Brown bear den site concealment in relation to human activity in Sweden. Ursus 22:152-158.

A120 Fink, T., J. G. Rasmussen, J. Emmersen, L. Pilgaard, Å. Fahlman, S. Brunberg, J. Josefsson, J. M. Arnemo, V. Zachar, J. E. Swenson, and O. Fröbert. 2011. Adipose-derived stem cells from the brown bear (Ursus arctos) spontaneously undergo chondrogenic and osteogenic differentiation in vitro. Stem Cell Research 7:89–95.

A121_120 Kindberg, J., J. E. Swenson, G. Ericsson, E. Bellemain, C. Miquel, P. Taberlet. 2011. Estimating population size and trends of the Swedish brown bear (Ursus arctos) population. Wildlife Biology 17: 114-123.

A122 Karamanlidis, A. A., A. Sanopoulos, L. Georgiadis, A. Zedrosser. 2011. Structural and economic aspects of human – bear conflicts in Greece. Ursus 22:141-151.

A123_122 Steyaert S. M. J. G, Støen O-G, Elfström M, Karlsson J, Van Lammeren R, Bokdam J, Zedrosser, A, Brunberg, S., and Swenson J. E. 2011. Resource selection by sympatric free-ranging dairy cattle brown bears Ursus arctos. Wildlife Biology 17: 389-403.

A124_123 Bischof, R. and Swenson, J. E. 2012. Combining noninvasive genetic sampling and traditional monitoring to aid management of a trans-border carnivore population. Ecological Applications 22:361-373.

A125 Steyaert, S. M.J.G., A. Endrestøl, K. Hackländer, J. E. Swenson, and A. Zedrosser. 2012. The mating system of the brown bear Ursus arctos. Mammalian Review 42:12-34.

A126 Gervasi, V., E. B. Nilsen, H. Sand, M. Panzacchi, G. R. Rauset, H. C. Pedersen, J. Kindberg, P. Wabakken, B. Zimmermann, J. Odden, O. Liberg, J. E. Swenson, and J. D. C. Linnell. 2012. Predicting the potential demographic impact of predators on their prey: a comparative analysis of two carnivore–ungulate systems in Scandinavia. Journal of Animal Ecology 81:443-454.

A127 Arinell, K., B. Sahdo, A. L. Evans, J. M. Arnemo, U. Baandrup, and O. Fröbert. 2012. Brown bears (Ursus arctos) seem resistant to atherosclerosis despite highly elevated plasma lipids during hibernation and active state. Clinical and Translational Science 5:269-272.

A128 Fahlman, Å., N. Caulkett, J. M. Arnemo, P. Neuhaus, and K. E. Ruckstuhl. 2012. Efficacy of a portable oxygen concentrator with pulsed delivery for treatment of hypoxemia during anesthesia of wildlife. Journal of Zoo and Wildlife Medicine 43:67-76.

A129_128 Moen, G. K., O.-G. Støen, V. Sahlén, and J. E. Swenson. 2012. Behaviour of solitary adult Scandinavian brown bears (Ursus arctos) when approached by humans on foot. PLOS One 7(2): e31699.

A130_129 Gonzalez, O., A. Zedrosser, F. Pelletier, J. E. Swenson, and M. Festa-Bianchet. 2012. Litter reductions reveal a trade-off between offspring size and number in brown bears. Behavioral Ecology and Sociobiology 66:1025-1032.

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A131_131 Rauset, G. R., J. Kindberg, and J. E. Swenson. 2012. Modeling female brown bear kill rates on moose calves using Global Positioning Satellite data. Journal of Wildlife Management 76:1597-1606.

A132 Schregel, J., A. Kopatz, S. B. Hagen, H. Brøseth, M. E. Smith, S. Wikan, I. Wartiainen, P. E. Aspholm, O. Makarova, N. Polikarpova, M. Schneider, P. M. Knappskog, J. E. Swenson, I. Kojola, M. Ruokonen, J. Aspi, K. F. Tirronen, P. I. Danilov, and H. G. Eiken. 2012. Limited gene flow among brown bear populations in far Northern Europe? Genetic analysis of the east-west border population in the Pasvik Valley. Molecular Ecology 21:3474-3488.

A133_133 Martin, J., E. Revilla, P.-Y. Quenette, J. Naves, D. Allaine, and J. E. Swenson. 2012. Assessing brown bear habitat suitability in the Pyrenees: transferability across sites and linking scales to make the most of scarce data. Journal of Applied Ecology 49:621-631.

A134_134 Painer, J., A. Zedrosser, J. M. Arnemo, Å. Fahlman, S. Brunberg, P. Segerström, and J. E. Swenson. 2012. Effects of different doses of medetomidine and tiletamine-zolazepam on the duration of induction and immobilization in free-ranging yearling brown bears (Ursus arctos). Canadian Journal of Zoology 90:753-757.

A135_135 Ordiz, A., O.-G. Støen, S. Sæbø, J. Kindberg, M. Delibes, and J. E. Swenson. 2012. Do bears know they are being hunted? Biological Conservation 152:21–28.

A136_136 Jojola, S. M, F. Rosell, I. Warrington, J. E. Swenson & A. Zedrosser. 2012. Subadult brown bears (Ursus arctos) discriminate between unfamiliar adult male and female anal gland secretion. Mammalian Biology 77:363-368.

A137_137(a) Evans, A. V. Salén, O.-G. Støen, Å. Fahlman, S. Brunberg, K. Madslien, O. Fröbert, J. E. Swenson, and J. M. Arnemo. 2012. Capture, anesthesia, and disturbance of free-ranging brown bears (Ursus arctos) during hibernation. PLOS One 7(7): e40520.

A138_137(b) Steyaert. M. J. G., F. J. Hütter, M. Elfström, A. Zedrosser, K. Hackländer, M. H. Lê, W. M. Windisch, J. E. Swenson, and T. Isaksson. 2012. Faecal spectroscopy: a practical tool to assess diet quality in an opportunistic omnivore. Wildlife Biology 18:431-438.

A139_140 Kuduk, K., W. Babik, K. Bojarska, E. Śliwińska, J. Kindberg, P. Taberlet, J. E. Swenson, & J. Radwan. 2012. Evolution of Major Histocompatibility Complex class I and class II genes in the brown bear. BMC Evolutionary Biology 12:197.

A140 Koren, L., D. Whiteside, Å. Fahlman, K. Ruckstuhl, S. Kutz, S. Checkley, M. Dumond, and K. Wynne-Edwards. 2012. Cortisol and corticosterone independence in cortisol-dominant wildlife. General and Comparative Endocrinology 177:113-119.

A141 Koren L, E. S. M. Ng, K. K. Soma, K. E. Wynne-Edwards. 2012. Sample preparation and liquid chromatography-tandem mass spectrometry for multiple steroids in mammalian and avian circulation. PLoS ONE 7(2): e32496.

A142_145 Stewart P., L. Campbell, S. Skogtvedt, K. A. Griffin, J. M. Arnemo, M. Tryland, S. Girling, M. W. Miller, M. A. Tranulis, and W. Goldmann. 2012. Genetic predictions of prion disease susceptibility in carnivore species based on variability of the prion gene coding region. PLoS ONE 7(12): e50623.

A143 Bischof, R., E. B. Nilsen, H. Brøseth, P. Männil, J. Ozolinš, and J. D. C. Linnell. 2012. Implementation uncertainty when using recreational hunting to manage carnivores. Journal of Applied Ecology 49:824-832.

A144_130 Zedrosser, A., F. Pelletier, R. Bishof, M. Festa-Bianchet, and J. E. Swenson. 2013. Determinants of lifetime reproduction in female brown bears: early body mass, longevity, and hunting regulations. Ecology 94:231-240.

A145_139 Martin, J., B. Van Moorter, E. Revilla, P. Blanchard, S. Dray, P.-Y. Quenette, D. Allainé, and J.E. Swenson. 2013. Reciprocal modulation of internal and external factors determines individual movements. Journal of Animal Ecology 82:290-300.

A146_143 Dahle, B., K. Wallin, G. Cederlund, I.-L. Persson, L. S. Selvaag, and J. E. Swenson. 2013. Predation on adult moose Alces alces by European brown bears Ursus arctos. Wildlife Biology 19:165-169.

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A147_144 Ordiz, A., O.-G. Støen, S. Sæbø, V. Sahlén, B. E. Pedersen, J. Kindberg, and Jon E. Swenson. 2013. Lasting behavioural responses of brown bears to experimental encounters with human. Journal of Applied Ecology 50:306-314.

A148_147 Steyaert, S.M.J.G., J. Kindberg, J.E. Swenson, and A. Zedrosser. 2013. Male reproductive strategy explains spatiotemporal segregation in brown bears. Journal of Animal Ecology 82:836-845.

A149 Revsbech, I.G.; H. Malte, O. Frobert, A. Evans, S. Blanc, J. Josefsson, A. Fago. 2013. Decrease in the red cell cofactor 2,3-diphosphoglycerate increases hemoglobin oxygen affinity in the hibernating brown bear Ursus arctos. American Journal of Physiology—Regulatory, Integrative, and Comparative Physiology 304: R43-R49.

A150 Sahdo, B., A. L. Evans, J. M. Arnemo, O. Fröbert, E. Särndahl, and S. Blanc. 2013. Body temperature during hibernation is highly correlated with a decrease in circulating innate immune cells in the brown bear (Ursus arctos): a common feature among hibernators? International Journal of Medical Sciences 10:508-514.

A151_150 Elfström, M., O.-G. Støen, A. Zedrosser, I. Warrington, and J. E. Swenson. 2013. Gut retention time in captive Scandinavian brown bears Ursus arctos. Wildlife Biology 19:317-324.

A152_151 Friebe, A., A. Zedrosser, and J. E. Swenson. 2013. Detection of pregnancy in a hibernator based on activity data. European Journal of Wildlife Research 59:731-741.

A153_153 Stenvinkel, P., O. Fröbert, B. Anderstam, F. Palm, M. Ericson, A.-C. Bragfors-Helin, A. R. Qureshi, T. Larsson, L. Bankir, A. Friebe, A. Zedrosser, J. Josefsson, M. Svensson, B. Sahdo, L. Bankir, and R. J. Johnson. 2013. Metabolic changes in summer active and anuric hibernating free-ranging brown bears (Ursus arctos) PLoS One 8(9): e7234.

A154 Jonzén, N., H. Sand, P. Wabakken, J. E. Swenson, J. Kindberg and O. Liberg, and , G. Chapron. 2013. Sharing the bounty—adjusting harvest to predator return in the Scandinavian human-wolf-bear-moose system. Ecological Modelling 265:140-148.

A155_155 López-Alfaro, C., C. T. Robbins, A. Zedrosser, and S. E. Nielsen. 2013. Energetics of hibernation and reproductive trade-offs in brown bears. Ecological Modelling 270:1-10.

A156_156 Steyaert S.M.J.G., C. Reusch, S. Brunberg, J. E. Swenson, K. Hackländer, and A. Zedrosser. 2013. Infanticide as a male reproductive strategy has a nutritive risk effect in brown bears. Biology Letters 9:20130624.

A157_157 Ordiz, A., R. Bischof, and J. E. Swenson. 2013. Saving large carnivores, but losing the apex predator? Biological Conservation 168:128-133.

A158_138 Elfström, M., A. Zedrosser, O.-G. Støen, and J. E. Swenson. 2014. Ultimate and proximate causes behind the occurrence of bears close to settlements and its management consequences: a review. Mammal Review 44:5-18.

A159 Karamanlidis, A.A., Stojanov, A., de Gabriel Hernando, M., Georgiadis, L., Ivanov, G., Karmbokoukis, L., Melovski, D., and Zedrosser, A. 2014. Distribution and genetic status of brown bears in FYR Macedonia: implications for conservation. Acta theriologica 59:119-128.

A160_158 Steyaert, S.M.J.G., J. Kindberg, J.E. Swenson, and A. Zedrosser. 2014. Litter loss triggers estrus in a nonsocial seasonal breeder. Ecology and Evolution ece3.935.

A161 Nawaz, M. A., J. Martin, and J. E. Swenson. 2014. Identifying key habitats to conserve the threatened brown bear in the Himalaya. Biological Conservation 170:198-206.

A162 Odden, M., R. A. Ims, O.-G. Støen, J. E. Swenson, and H. P. Andreassen. 2014. Bears are simply voles writ large: social structure determines the mechanisms of intrinsic population regulation in mammals. Oecologia 175:1-10.

A163_161 Ordiz, A., J. Kindberg, S. Sæbø, J. E. Swenson, and O.-G. Støen. 2014. Brown bear circadian behavior reveals human environmental encroachment. Biological Conservation 173:1-9.

A164_162 Elfström, M, A. Zedrosser, K. Jerina, O.-G. Støen, J. Kindberg, L. Budic, M. Jonozovič, and J. E. Swenson. 2014. Does despotic behavior or food search explain

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the occurrence of problem brown bears in Europe? Journal of Wildlife Management 78:881-893.

A165 Karamanlidis, A.A., S. Pllaha, L. Krambokoukis, K. Shore, and A. Zedrosser. 2014. Preliminary brown bear survey in southeastern Albania. Ursus 25:1-7.

A166_165 Revsbech, I. G., X. Shen, R. Chakravarti, F. B. Jensen, B. Thiel, A. L. Evans, J. Kindberg, O. Fröbert, D. J. Stuehr, C. G. Kevil, and A. Fago. 2014. Hydrogen sulfide and nitric oxide metabolites in the blood of free-ranging brown bears and their potential roles in hibernation. Free Radical Biology & Medicine 73:349–357.

A167 Cattet, M., G. Stenhouse, B. Macbeth, D. Janz, A. Zedrosser, J. E. Swenson, and M. Dumond. 2014. Quantifying long-term stress in brown bears with the hair cortisol concentration: a biomarker that may be confounded by rapid changes in response to capture and handling. Conservation Physiology 2: doi:10.1093/conphys/cou026.

A168 Sergiel, A., R. Maślak, A. Zedrosser, Ł. Paśko, D. L. Garshelis, S. Reljić, and D. Huber. 2014. Fellatio in captive brown bears: evidence of long-term effects of suckling deprivation? Zoo Biology 33:349-352.

A169_168 Friebe, A., A. L. Evans, J. M. Arnemo, S. Blanc, S. Brunberg, G. Fleissner, J. E. Swenson, and A. Zedrosser. 2014. Factors affecting date of implantation, parturition, and den entry estimated from activity and body temperature in free-ranging brown bears. PLoS ONE 9(7): e101410.

A170_169 Ozeki, L. M., Å. Fahlman, G. Stenhouse, J. M. Arnemo, N. Caulkett. 2014. Evaluation of the accuracy of different methods of monitoring body temperature in anesthetized brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 45: 819-824.

A171_170 Græsli, A. R., Å. Fahlman, A. L. Evans, M. F. Bertelsen, J. M. Arnemo, and S. S. Nielsen. 2014. Haematological and biochemical reference intervals for free-ranging brown bears (Ursus arctos) in Sweden. BMC Veterinary Research 10:183.

A172_171 Kuduk, K. W. Babik, E. Bellemain, A. Valentini, A. Zedrosser, P. Taberlet, J. Kindberg, Jon E. Swenson, and J. Radwan. 2014. No evidence for the effect of MHC on male mating success in the brown bear. PLoS ONE 9(12): e113414.

A173_172 Elfström, M., M. L. Davey, A. Zedrosser, M. Müller, M. De Barba, O.-G. Støen, C. Miquel, P. Taberlet, K. Hackländer, and J. E. Swenson. 2014. Do Scandinavian brown bears approach settlements to obtain high-quality food? Biological Conservation 178:128–135.

A174 McKay, T., E. Sahlén, O.-G. Støen, J. E. Swenson, and G. B. Stenhouse. 2014. Wellsite selection by grizzly bears Ursus arctos in west-central Alberta. Wildlife Biology 5:310-319.

A175_174 Jørgensen, P. G., J. Arnemo, J. E. Swenson, J. S. Jensen, S. Galatius, and O. Frøbert. 2014. Low cardiac output as physiological phenomenon in hibernating free-ranging Scandinavian brown bears (Ursus arctos) - an observational study. Cardiovascular Ultrasound 12:36.

A176 Hodder, D.P., Johnson C.J., Rea, R.V., and Zedrosser, A. 2014. Application of a species distribution model to identify and manage American black bear denning habitat in central British Columbia, Canada. Wildlife Biology 20, 238-245.

A177 Fahlman, Å, J. M. Arnemo, J. Pringle, and G. Nyman. 2014. Oxygen supplementation in anesthetized brown bears (Ursus arctos)—how low can you go? Journal of Wildlife Diseases 50:574–581.

A178_177 Steyaert, S. M. J. G., J. Kindberg, K. Jerina, M. Krofel, M. Stergar, J. E. Swenson, and A. Zedrosser. 2014. Behavioral correlates of supplementary feeding of wildlife: can general conclusions be drawn? Basic and Applied Ecology 15:669–676.

A179 Chapron, G., P. Kaczensky, J. D. C. Linnell, M. von Arx, D. Huber, H. Andrén, J. V. López-Bao, M. Adamec, F. Álvares, O. Anders, L. Balciauskas, V. Balys, P. Bedo, F. Bego, J. C. Blanco, U. Breitenmoser, H. Brøseth, L. Bufka, R. Bunikyte, P. Ciucci, A. Dutsov, T. Engleder, C. Fuxjäger, C. Groff, M. Heltai, K. Holmala, B. Hoxha, Y. Iliopoulos, O. Ionescu, G. Ivanov, J. Jeremić, K. Jerina, F. Knauer, I. Kojola, I. Kos, M. Krofel, J. Kubala, S. Kunovac, J. Kusak, M. Kutal, P. Mannil, R. Manz, E. Marboutin, F. Marucco, D. Melovski, K. Mersini, Y. Mertzanis, R. W. Mysłajek, S.

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Nowak, J. Odden, J. Ozolins, G. Palomero, M. Paunovic, J. Persson, H. Potočnik, P.-Y. Quenette, G. Rauer, I. Reinhardt, R. Rigg, A. Ryser, V. Salvatori, T. Skrbinšek, A. Skrbinšek-Majić, A. Stojanov, J. E. Swenson, A. Trajçe, E. Tzingarska-Sedefcheva, M. Váňa, R. Veeroja, M. Wölfl, S. Wölfl, F. Zimmermann, D. Zlatanova & Luigi Boitani. 2014. Successful recovery of large carnivores in Europe’s human-dominated landscapes. Science 346:1517-1519.

A180_179 Sahlén, V., A. Friebe, S. Sæbo, J. E. Swenson, and O.-G. Støen. 2015. Den entry behavior in Scandinavian brown bears: implications for preventing human injuries. Journal of Wildlife Management. 79:274–287.

A181_176 Gosselin, J., A. Zedrosser, Jon E. Swenson, and F. Pelletier. 2015. The relative importance of direct and indirect effects of hunting mortality on the population dynamics of brown bears. Proceedings of the Royal Society of London, B 282: 20141840.

A182_181 Sahlén, V., A. Ordiz, J. E Swenson, and O.-G. Støen. 2015. Behavioural differences between single Scandinavian brown bears (Ursus arctos) and females with dependent young when experimentally approached by humans. PLoS ONE 10(4): e0121576.

A183_182 Ozeki, L. M., N. Caulkett, G. Stenhouse, J. M. Arnemo, Å. Fahlman. 2015. Effect of active cooling and α2-adrenoceptor antagonism on core temperature in anesthetized brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 46:279-285.

A184 Gilroy, J. J., A. Ordiz, and R. Bischof. 2015. Carnivore coexistence: value the wilderness. Science 347:382.

A185 Stenvinkel, P. J. Kindberg, & O. Fröbert. 2015. Biomimetik – att efterlikna naturen för att förebygga sjukdom. Läkartidningen 112:586-590.

A186_185 Frank, S., S. M. J. G. Steyaert, J. E. Swenson, I. Storch, J. Kindberg, H. Barck, A. Zedrosser. 2015. A clear-cut case? Brown bear foraging for carpenter ants on clear-cuts across spatial scales in central Sweden. Forest Ecology and Management 348:164–173.

A187_186a Græsli, A. R., A. L. Evans, Å. Fahlman, M. F. Bertelsen, S. Blanc, and J. M. Arnemo. 2015. Seasonal variation in haematological and biochemical variables in free-ranging subadult brown bears (Ursus arctos) in Sweden. BMC Veterinary Research 11:301.

A188_187 Bischof, R., Brøseth, H. and Gimenez, O. 2015, Wildlife in a Politically Divided World: Insularism Inflates Estimates of Brown Bear Abundance. Conservation Letters. 2015

A189 Xenikoudakis, G., E. Ersmark, J.-L. Tison, L. Waits, J. Kindberg, J. E. Swenson, and L. Dalén. 2015. Consequences of a demographic bottleneck on genetic structure and variation in the Scandinavian brown bear. Molecular Biology 24:3441-3454.

A190_189 Paillard, L, K Jones, A. Evans, J. Berret, M. Jacquet, R. Lienhard, M. Bouzelboudjen, J. M. Arnemo, J. E. Swenson, and Maarten J. Voordouw. 2015. Serological signature of tick-borne pathogens in Scandinavian brown bears over two decades. Parasites & Vectors 8:398.

A191 Mills, J. A., C. Teplitsky, B. A. Lopez, A. Charmantier, P. H. Becker, T. R. Birkhead, P. Bize, D. T. Blumstein, C. Bonenfant, S. Boutin, A. Bushuev, E. Cam, A. Cockburn, S. D. Côté, F. Daunt, N. Dingemanse, B. Doligez, H. Drummond, R. H. M. Espie, M. Festa-Bianchet, F. Frentiu, J. W. Fitzpatrick, R. W. Furness, D. Garant, G. Gauthier, P. R. Grant, M. Griesser, L. Gustafsson, B. Hansson, M. P. Harris, F. Jiguet, P. Kjellander, E. Korpimäki, C. J. Krebs, L. Lens, J. D.C. Linnell, M. Low, A. McAdam, A. Margalida, J. Merilä, A. P. Møller, S. Nakagawa, J.-Å. Nilsson, I. C. T. Nisbet, A. van Noordwijk, D. Oro, T. Pärt, F. Pelletier, J. Potti, B. Pujol, D. Réale, R. F. Rockwell, Y. Ropert-Coudert, A. Roulin, J. S. Sedinger, Jon E. Swenson, M. E. Visser, S. Wanless, D. F. Westneat, A. Wilson, and A. Zedrosser. 2015. Archiving primary data: solutions for long-term studies. Trends in Ecology and Evolution 30:581-589.

A192_191 Ordiz, A., C. Milleret, J. Kindberg, J. Månsson, P. Wabakken, J. E. Swenson, and H. Sand. 2015. Wolves, people, and brown bears influence the expansion of the recolonizing wolf population in Scandinavia. Ecosphere 6(12): article 284.

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A193 Berg von Linde, M., L. Arevström, and O. Fröbert. 2015. Insights from the den: how hibernating bears may help us understand and treat human disease. Clinical and Translational Science 8:601-605.

A194 Schregel, J., Eiken, H. G., Grøndahl, F. A., Hailer, F., Aspi, J., Kojola, I., Tirronen, K., Danilov, P., Rykov, A., Poroshin, E., Janke, A., Swenson, J. E. and Hagen, S. B. 2015. Y chromosome haplotype distribution of brown bears (Ursus arctos) in Northern Europe provides insight into population history and recovery. Molecular Ecology, 24: 6041–6060.

A195_194 Leclerc, M., E. Vander Wal, A. Zedrosser, J. E. Swenson, J. Kindberg, and F. Pelletier. 2015. Quantifying consistent individual differences in habitat selection. Oecologia pp1-9.

A196_195 Støen, O.-G., A. Ordiz, A. L. Evans, T. Laske, J. Kindberg, O. Fröbert, J. E. Swenson, and J. M. Arnemo. 2015. Physiological evidence for a human-induced landscape of fear in brown bears (Ursus arctos). Physiology and Behavior 152A:244-248.

A197_186b Esteruelas, N. F., N. Huber, A. L. Evans, A. Zedrosser, M. Cattet, F. Palomares, M. Angel, J. E. Swenson, J. M. Arnemo. 2016. Leukocyte coping capacity as a tool to assess capture and handling-induced stress in Scandinavian brown bears (Ursus arctos). Journal of Wildlife Diseases 52:S40-S53

A198_197 Stenset, N. E., P. N. Lutnæs, V. Bjarnadóttir, B. Dahle, K. H. Fossum, P. Jigsved, T. Johansen, W. Neumann, O. Opseth, O. Rønning, S. M. J. G. Steyaert, A. Zedrosser, S. Brunberg, and J. E. Swenson. In press. Seasonal and annual variation in the diet of brown bears (Ursus arctos) in the boreal forest of southcentral Sweden. Wildlife Biology

A199 Rauset, G. R., H. Andrén, J. E. Swenson, G. Samelius, P. Segerström, A. Zedrosser, and J. Persson. In press. National parks in northern Sweden as refuges for illegal killing of large carnivores. Conservation Letters

A200_199 Sommer, F., M. Ståhlman, O. Ilkayeva, J. M. Arnemo, J. Kindberg, J. Josefsson, C. Newgard, O. Fröbert, and F. Bäckhed. 2016. The gut microbiota modulates energy metabolism in the hibernating brown bear (Ursus arctos). Cell Reports 14:1-7.

A201_200. Steyaert, S. M. J. G., A. Zedrosser, M. Elfström, A. Ordiz, M. Leclerc, S. C. Frank, J. Kindberg, O.-G. Støen, S. Brunberg; and J. E. Swenson. In press. Ecological implications from spatial patterns in human-caused brown bear mortality. Wildlife Biology

A202_201. Evans, A. L., N. J. Singh, A. Friebe, J. M. Arnemo, T. G. Laske, O. Fröbert, J. E. Swenson, and S. Blanc. 2016. Ecophysiological drivers of den entry and den exit in the brown bear. Frontiers in Zoology 13:7.

A203 Penteriani, V., M. del Mar Delgado, F. Pinchera, J. Naves, A. Fernández-Gil, I. Kojola, S. Härkönen, H. Norberg, J. Frank, J. M. Fedriani, V. Sahlén, O.-G. Støen, J. E. Swenson, P. Wabakken, M. Pellegrini, S. Herrero, and J. V. López-Bao. 2016 Human behaviour may trigger large carnivore attacks in developed countries. Scientific Reports 6:20552

A204 Mills, J. A., C. Teplitsky, B. A. Lopez, A. Charmantier, P. H. Becker, T. R. Birkhead, P. Bize, D. T. Blumstein, C. Bonenfant, S. Boutin, A. Bushuev, E. Cam, A. Cockburn, S. D. Côté, F. Daunt, N. Dingemanse, B. Doligez, H. Drummond, R. H. M. Espie, M. Festa-Bianchet, F. Frentiu, J. W. Fitzpatrick, R. W. Furness, D. Garant, G. Gauthier, P. R. Grant, M. Griesser, L. Gustafsson, B. Hansson, M. P. Harris, F. Jiguet, P. Kjellander, E. Korpimäki, C. J. Krebs, L. Lens, J. D.C. Linnell, M. Low, A. McAdam, A. Margalida, J. Merilä, A. P. Møller, S. Nakagawa, J.-Å. Nilsson, I. C. T. Nisbet, A. van Noordwijk, D. Oro, T. Pärt, F. Pelletier, J. Potti, B. Pujol, D. Réale, R. F. Rockwell, Y. Ropert-Coudert, A. Roulin, J. S. Sedinger, J. E. Swenson, M. E. Visser, S. Wanless, D. F. Westneat, A. Wilson, and A. Zedrosser. 2016 Solutions for archiving data in long-term studies - a reply to Whitlock et al. Trends in Ecology and Evolution 31:85–87.

A205_204 Johansson, M., O.-G. Støen, and A. Flykt. In press? Exposure as an intervention to address human fear of bears. Human Dimensions of Wildlife

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A206_206 Esteruelas, N. F., J. Malmsten, C. Bröjer, G. Grandi, A. Linström, P. Brown, J. E. Swenson, A. L. Evans, J. M. Arnemo. In press. Chewing lice Trichodectes pinguis pinguis in Scandinavian brown bears (Ursus arctos). Journal of Parasitology: Parasites and Wildlife

B) Books and book chaptersB1 Bjärvall, A., R. Franzén, M. Nordkvist and G. Åhman. 1990. Renar och rovdjur—

rovdjurens effekter på rennäringen. Naturvårdsverket förlag, Solna. (Reindeer and predators—the effects of predators on the reindeer industry; in Swedish)

B2 Sandegren, F. and J. Swenson. 1997. Björnen—viltet, ekologin och människan. Svenska Jägareförbundet, Stockholm, Sweden. (The brown bear—the animal, ecology, and man; in Swedish). 70 pp.

B3 Sørensen, O. J., J. E. Swenson and T. Kvam. 1999. Status and management of the brown bear in Norway. Pages 86-89 in C. Servheen, S. Herrero and B. Peyton, compliers. Bears, status survey and conservation action plan. IUCN/SSC bear and polar bear specialist groups, IUCN, Gland, Switzerland and Cambridge, UK. 309 pp. (Invited chapter)

B4 Swenson, J. E., F. Sandegren, A. Bjärvall, R. Franzén, A. Söderberg and P. Wabakken. 1999. Status and management of the brown bear in Sweden. Pages 111-113 in C. Servheen, S. Herrero and B. Peyton, compliers. Bears, status survey and conservation action plan. IUCN/SSC bear and polar bear specialist groups, IUCN, Gland, Switzerland and Cambridge, UK. 309 pp. (Invited chapter)

B5 Swenson, J. 2000. Der Braunbär (Ursus arctos) in Eurasien. Pages 89-108 in U. Gansloßer, editor. Die Bären. Filander Verlag, Fürth. 314 pp. (Invited chapter)

B6 Swenson, J. E., N. Gerstl, B. Dahle & A. Zedrosser. 2000. Action plan for the conservation of the brown bear (Ursus arctos) in Europe. Council of Europe, Report T-PVS (2000) 24. 79 pp.

B7 Kreeger, T. J., J. M. Arnemo, and J. P. Raath. 2002. Handbook of Wildlife Chemical Immobilization. International Edition. Wildlife Pharmaceuticals Inc., Ft. Collins, Colorado, USA: 412 pp.

B8 Swenson, J. E. 2003. Implications of sexually selected infanticide for hunting of large carnivores. Pages 171-189 in M. Festa-Bianchet and M. Apollonio, eds. Animal Behavior and Wildlife Management. Island Press, Washington, D. C. 371 pp.

B9. Swenson, J. E. and H. Andrén. 2005. A tale of two countries: large carnivore depredations and compensation schemes in Sweden and Norway. Pages 323-339 in R. Woodroffe, S. Thirgood, and A. Rabinowitz, eds. People and Wildlife: Conflict or Co-existence? Cambridge University Press. (Invited chapter)

B10 Linnell, J. D. C., C. Promberger, L. Boitani, J. E. Swenson, U. Breitenmoser and R. Andersen. 2005. The linkage between conservation strategies for large carnivores and biodiversity: the view from the “half-full” forests of Europe. Pages 381-399 in J. C. Ray, K. H. Redford, R. S. Steneck and J. Berger . Large carnivores and the conservation of biodiversity. Island Press, Washington, D. C. 562 pp.

B11 Martin, J., V. Tolon, B. van Moorter, M. Basille and C. Calenge. 2009. On the use of telemetry in habitat selection studies. Pp 37-55 in D. Barculo and J. Daniels, eds. Telemetry: research, technology and applications, Nova Science Publishers Inc, New York, USA.

B12. Swenson, J. E. 2015. Conservation research and animal activism: in search of a better balance. Pp 207-215 in L. Kemmerer (ed). Animals and the Environment: Advocacy, Activism, and the Quest for Common Ground, Routledge, London and New York.

C) Papers in nonrefereed publications

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C1 Swenson, J. 1994. Sweden and Norway: historic and present status of the brown bear in Scandinavia. Intern. Bear News 3(3):5-6.

C2 Dalen, L., Johansen, T. Dahle, B. 1996. Yet another bryophyte consuming beast. Bryolog. Times 87 (2):1.

C3 Arnemo, J. M. & P. Dypsund. 1997. Kirurgisk implantation av radiosändare på vilda rovdjur. Svensk Veterinärtidning 49(1):17-18. (Surgical implantation of radio transmitters in wild carnivores. In Swedish)

C4 Arnemo, J. M. & P. Dypsund. 1997. Ny merkemetodikk på brunbjørn. Norsk Veterinærtidsskrift 109:440. (A new method for marking brown bears. In Norwegian)

C5 Sandegren, F. & J. Swenson. 1997. Det skandinaviska björnprojektet. Pp. 76-81 in Rovviltforvaltning: problemer og utfordringer. Nordisk Jägersamvirke, Rapport Nr. 4-1997. (The Scandinavian Brown Bear Research Project. In Swedish)

C6 Gossow, H. and J. Swenson. 1997. Large predator meeting in Austria. Intern. Bear News 6(3):12-13.

C7 Swenson, J. and C. Servheen. 1997. Does bear conservation without hunting produce problem bears? Intern. Bear News 6(4):11.

C8 Swenson, J. E. 1998. Coordination of large-carnivore monitoring, management, and research in Scandinavia. Pages 85-88 in C. Breitenmoser-Würster, C. Rohner, and U. Breitenmoser, editors. The re-introduction of the lynx into the Alps, Council of Europe Publishing, Environmental Encounters No. 38.

C9 Arnemo, J. M., P. Dypsund, F. Berntsen, S. J. Wedul, B. Ranheim & L. Lundstein. 1998. Bruk av implanterbare radiosendere på store rovdyr. Norsk Veterinærtidsskrift 110:799-803. (Use of implantable radio transmitters in large carnivores. In Norwegian)

C10 Andersen, R. & J. Swenson. 1999. Wildlife and nature conservation in Scandinavia with special regard to large predators. Pages 59-67 in Natura 2000 Eine Chance für den Naturschutz Europas/A chance for nature conservation in Europe. Schriftenreihe des Bundesministerium für Umwelt, Jugend und Familie, Band 14/1999. Vienna.

C11 Swenson, J. & F. Sandegren. 1999. Den svenske bjørnebestandens levedyktighet. Pages 85-90 in T. Ebenhard & M. Höggren, eds. Livskraftiga rovdjurstammar. Centrum för biologisk mångfold, Skriftserie 1. Uppsala, Sweden. (The viability of the Swedish brown bear population. In Norwegian)

C12 Swenson, J. E. and F. Sandegren. 2000. Conservation of European brown bear populations: experiences from Scandinavia. Pages 111-116 in J. F. Layna, B. Heredia, G. Palomero and I. Doadrio, editors. La conservación del oso pardo en Europa: un reto de cara al siglo XXI. Serie encuentros I. Fundación Biodiversidad, Ministerio de medio ambiente, Madrid, Spain.

C13 Dahle, B. 2000. Extensive illegal killing of brown bears in Sweden. Intern. Bear News 9(2):7.

C14 Dahle, B. 2000. Large-carnivore white paper presented in Sweden. Intern. Bear News 9(2):8.

C15 Arnemo, J. M. 2001. Reversible anaesthesia in brown bears. DDA News 2:8.C16 Swenson, J. E. 2001. Bjørnens tilbakekomst. Det norske videnskaps-akademi,

Årbok 2000: 463-470. (In Norwegian: The return of the brown bear)C17 Arnemo, J. M., S. Brunberg, P. Ahlqvist, R. Franzén, A. Friebe, P. Segerström, A.

Söderberg, and J. E. Swenson. 2001. Reversible immobilization and anesthesia of free-ranging brown bears (Ursus arctos) with medetomidine-Tiletamine-Zolazepam and Atipamezole: a review of 575 captures. Proceedings of the American Association of Zoo Veterinarians, American Association of Wildlife Veterinarians, Association of Reptilian and Amphibian Veterinarians, and National Association of Zoo and Wildlife Veterinarians Joint Conference. 2001:234-236.

C18 Zedrosser, A. & B. Dahle. 2002. Brown bear attack in central Sweden. Int. Bear News 11:9.

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C19 Bjärvall, A. 2003. Stora rovdjur—samspel mellan forskning och förvaltning med björnen som exempel. Pages 13-20 in L. Terenius, Chefsredaktör. Icke-traditionella försöksdjur I forskningen. Centrala försöksdjursnämnden, Skriftserie Nr. 48. Stockholm.

C20 Swenson, J. And F. Sandegren. 2003. A summary of results from the Scandinavian brown bear research project. Pages 186-189 in P. I. Danilov and V. B. Zimin, editors. Dynamics of game animals populations in Northern Europe—Proceedings of the third international symposium. Karel’skii Nauchnyi Tsentr RAN, Redaktsionno-izdatel’skii Otdel, Petrozavodsk, Russia.

C21 Karamanlidis, A., Georgiadis, L., and Zedrosser, A. 2009. The "Southwestern Balkans Bear Register": a tool in the conservation of brown bears in the southwestern Balkans. International Bear News 18: 13-14.

C22 Swenson, J. E., A. Zedrosser, and R. Bischof. 2013. Potential evolutionary consequences of hunting brown bears. Pages 1-4 in M. Beuković, Editor-in-Chief. Modern aspects of sustainable management of game populations. The second international symposium on hunting. Faculty of Agriculture, University of Novi Sad, Novi Sad, Serbia. 268 pp.

D) Completed student theses(* indicates theses that have resulted in a publication, (*) indicates that work on a manuscript is

underway or the manuscript has been submitted)

Doctorate level theses D1.1 2003 * Bjørn Dahle, Norwegian University of Science and Technology, Trondheim.

Dr. scient. (PhD) thesis: Reproductive strategies in Scandinavian brown bears.D1.2 2004 * Jon E. Swenson, Institut für Wildbiologie und Jagdwirtschaft, Universität für

Bodenkultur Wien, Vienna, Austria. Habilitation (Dr.habil.) thesis: The ecology of an increasing brown bear population: managing a successful recovery.

D1.3 2004* Alain Cercueil, Laboratoire des Techniques de l’Imagerie, de la Modélisation et de la Cognition & Laboratoire d’Ecologie Alpine, Université Joseph Fourier, Grenoble, France. Thèse d’université (PhD thesis): Contributions statistiques en génétique des populations.

D1.4 2004 * Eva Bellemain, Agricultural University of Norway, Ås (PhD) and Université Joseph Fourier, Grenoble (Docteur); joint (co-tutelle) degree program. PhD thesis/thèse d’université: Genetics of the Scandinavian brown bear: implications biology and conservation.

D1.5 2006 * Ole-Gunnar Støen, Norwegian University of Life Sciences, Ås. PhD thesis: Natal dispersal and social organization in brown bears.

D1.6 2006* Andreas Zedrosser, Norwegian University of Life Sciences, Ås and University of Natural Resources and Applied Life Sciences, Vienna. PhD thesis: Life-history strategies of brown bears.

D1.7 2006* Jonna Katajisto, University of Helsinki. Dr. scient. (Doctor of Science) thesis: Habitat use and spatial population dynamics of brown bears (Ursus arctos) in Scandinavia.

D1.8 2008* Alice Valentini, Università delgi studi della Tuscia di Viterbo (Dottorato di Recerca) and Université Joseph Fourier, Grenoble (Docteur); joint (co-tutelle) degree program. Doctoral thesis: Non-invasive diet analysis based on DNA barcoding: the Himalayan brown bears (Ursus arctos isabellinus) as a case study.

D1.9 2008* Mohammad Ali Nawaz, Norwegian University of Life Sciences, Ås. PhD thesis: Ecology, genetics and conservation of Himalayan brown bears.

D1.10 2008* Åsa Fahlman, Swedish University of Agricultural Sciences, Uppsala. PhD thesis: Advances in wildlife immobilisation and anaesthesia; clinical and physiological evaluation in selected species.

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D1.11 2009* Jodie Martin, Norwegian University of Life Sciences, Ås and Université Claude Bernard – Lyon 1 / Université de Lyon, Lyon. PhD thesis: Habitat selection and movement by brown bears in multiple-use landscapes.

D1.12 2009* Richard Bischof, Norwegian University of Life Sciences, Ås. PhD thesis: Selective harvesting: short- and long-term consequences for wild populations and their management.

D1.13 2010* Andrés Ordiz, Norwegian University of Life Sciences, Ås and Universidad de León, León. PhD thesis: Reproductive patterns and human-influenced behavior in brown bears. Implications for the conservation of large carnivores.

D1.14 2010* Jonas Kindberg, Swedish University of agricultural Sciences, Umeå, PhD thesis: Monitoring and management of the Swedish brown bear (Ursus arctos) population.

D1.15 2011* Susan M. Jojola, Norwegian University of Life Sciences, Ås. PhD thesis: Social organization and olfactory communication in brown bears, Eurasian beavers, and yellow-bellied marmots.

D1.16 2012* Sam Steyaert, Norwegian University of Life Sciences, Ås and University of Natural Resources and Applied Life Sciences, Vienna. PhD thesis: The mating system of the brown bear in relation to the sexually selected infanticide theory.

D1.17 2013* Veronica Sahlén, Norwegian University of Life Sciences, Ås. PhD thesis: Encounters between brown bears and humans in Scandinavia—contributing factors, bear behavior and management perspectives.

D1.18 2013* Marcus Elfström, Norwegian University of Life Sciences, Ås. PhD thesis: Patterns and mechanisms behind the occurrence of bears close to human settlements.

D1.19 2015* Andrea Friebe, Goethe-Universität in Frankfurt am Main. PhD thesis. Winter ecology of free-ranging brown bears (Ursus arctos) in central Sweden. Dissertation.

D1.20 2015* Julia Schregel, Norwegian University of Life Sciences, Ås. PhD thesis: Genetic structure and gene flow in a continuously distributed large terrestrial carnivore – the brown bear (Ursus arctos) in Northern Europe.

D1.21 2015 Marine Plumel, Institut pluridisciplinaire Hubert Curien, Strasbourg, France. PhD thesis: Optimization of quantitative proteomics analytical strategies: application to studying metabolic adaptive responses in various organisms.

Master of Science level theses (74) (bold = accepted to a doctoral program, 20 = 27%)D2.01 1990 Marie Dahlström, Stockholm University. Undergraduate thesis (20 points):

Licensjakten på björn 1981-89—en sammanställning. (License hunting of bears, 1981-89—a summary)

D2.02 1993 Caroline Paulson, Veterinary College, Swedish University of Agricultural Sciences, Uppsala. Thesis. Immobilisering av brunbjörn. (Immobilization of brown bears)

D2.03 1995* Jens Thomas Sagør, University of Trondheim. Cand. scient. (Master of Science) thesis: Et studie av konflikten mellom bjørn og sau i perioden 1981-1993. (A study of the conflict between bears and sheep in the period 1981-1993).

D2.04 1996* Bjørn Dahle, Norwegian University of Science and Technology, Trondheim. Cand. scient. (Master of Science) thesis: Nutritional ecology of brown bears (Ursus arctos) in Scandinavia with special reference to moose (Alces alces)

D2.05 1996* Raili Ytterberg, Stockholm University. Undergraduate thesis (10 points): Do ants support the high reproductive rate in the Scandinavian brown bear (Ursus arctos) population?

D2.06 1997* Anna Jansson, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Can a high protein availablility explain the high reproductive rate in the Scandinavian brown bear (Ursus arctos)?

D2.07 1997* Thomas Johansen, Norwegian University of Science and Technology, Trondheim. Cand. scient. (Master of Science) thesis: The diet of the brown bear (Ursus arctos) in central Sweden.

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D2.08 1998* Inga-Lill Persson, University of Oslo. Cand. scient. (Master of Science) thesis: Brown bear Ursus arctos predator upon adult moose in Scandinavia: a study at two levels of scale.

D2.09 1998* Ole Opseth, Norwegian University of Science and Technology, Trondheim. Cand. scient. (Master of Science) thesis: Brown bear (Ursus arctos) diet and predation on moose (Alces alces) calves in the southern tiaga zone in Sweden.

D2.10 1998* Helena Busk, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Brown bear (Ursus arctos) predation on moose (Alces alces) calves in a Swedish boreal forest.

D2.11 1999* Line Stabell, University of Oslo. Cand. scient. (Master of Science) thesis: Use of ungulates by brown brown bears Ursus arctos in Scandinavia: effects of area, season, sex, age, and individual.

D2.12 2000 Torleiv Yli Myre, Agricultural University of Norway, Ås. Cand. agric. (Master of Science) thesis: Strategies for female brown bear (Ursus arctos) to avoid infanticide: activity patterns.

D2.13 2000* Andrea Friebe, Johann-Wolfgang Goethe Universität, Frankfurt am Main, Diplomarbeit (Master of Science) thesis: Das Winterverhalten und die täglichen Wanderungen von weiblichen Braunbären (Ursus arctos) in Zentralschweden.

D2.14 2000* Håkon Hustad, Agricultural University of Norway, Ås. Cand. agric. (Master of Science) thesis: The issuing of kill permits for brown bears in response to domestic sheep depredation in Norway, 1989-99.

D2.15 2000 Beverly Ann Wilson, University of Stirling, UK. Master of Science thesis: Conservation management of the brown bear , Ursus arctos, in Scandinavia: using GIS and a habitat quality index to explain the temporal and spatial variation in sub-population core areas.

D2.16 2000 Christer Zakrisson, Swedish University of Agricultural Sciences, Umeå. Undergraduate thesis (20 points): Do brown bear (Ursus arctos) females with cubs alter their movement pattern in order to avoid infanticidal males?

D2.17 2000* Rikako Fujita, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Bait-hunting for brown bear in Sweden: temporal and spatial occurrence and potential effects on the population.

D2.18 2001 Wiebke Neumann, Universität Osnabrück, Osnabrück, Diplomarbeit (Master of Science) thesis: The brown bear Ursus arctos and berries in the Swedish boreal forest: exploiting a temporarily and spatially variable resource.

D2.19 2001 Oddmund Rønning, Agricultural University of Norway, Ås. Cand. agric. (Master of Science) thesis: Spatial and temporal supply of berries and their use by brown bears in south-central Sweden.

D2.20 2001 Valgerður Bjarnadottír, Swedish University of Agricultural Sciences, Uppsala. International Master of Science thesis: Brown bear (Ursus arctos) use of three berry species in central Sweden: do bears choose foraging plots based on the densities of berries?

D2.21 2001 Kristin Høivik Fossum, Agricultural University of Norway, Ås. Cand. scient. (Master of Science) thesis: Brown bears (Ursus arctos) and berries: use of a spatially and temporally variable resource by bears of different age and sex.

D2.22 2001 Jonna Katajisto, University of Helsinki, Helsinki. Master of Science thesis: Bed site selection of female brown bears (Ursus arctos) as a counter-strategy to avoid sexually selected infanticide by males.

D2.23 2001 * Eva Bellemain, Université Claude Bernard, Villeurbanne, France. Master of Science thesis: Etude du système d'appariement de l'ours brun de Scandinavie. (Study of the Scandinavian brown bear mating system) (In Franch and English).

D2.24 2001 Ander Larunbe Anderson, University of Stirling, UK. Master of Science thesis: Use of GIS to study brown bear, Ursus arctos, distribution in relation to habitat and traditional reindeer (Rangifer tarandus) herding in Scandinavia.

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D2.25 2002 Solveig Kristoffersen, University of Oslo, Oslo. Cand. scient. (Master of Science) thesis: Restricting daily movements as a counterstrategy against sexually selected infanticide in brown bears (Ursus arctos).

D2.26 2002 Paul Antoni Nilsen, Agricultural University of Norway, Ås. Cand. scient. (Master of Science) thesis: Scandinavian brown bear (Ursus arctos L.) foraging on tempory and spatially variable berry resources in the boreal forest.

D2.27 2002 Jessica Åsbrink, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis: Parasites and metals in the Scandinavian Brown Bear (Ursus arctos). Study project from the Department of Pathology, No. 30.

D2.28 2002 Lars Plahn, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Avoidance of potentially infanticidal males by female brown bears (Ursus arctos): patterns of diel activity.

D2.29 2003 Patrik Jigsved, Umeå University, Umeå. Undergraduate thesis (30 ECTS): Utilisation of berries by brown bears in the Scandinavian boreal forest.

D2.30 2003* Knut Håkon Solberg & Ola-Mattis Drageset, Agricultural University of Norway, Ås. Cand. scient. (Master of Science) thesis: A test of methods to estimate brown bear (Ursus arctos) population size.

D2.31 2004 Insa Bauhaus, Albert-Ludwigs-Universität, Freiburg, Germany. Diplomarbeit thesis: Factors for the use of carpenter ants (Campanotus spp.) on clear-cuts by Scandinavian brown bears (Ursus arctos L.).

D2.32 2004 Eskil Nerheim, Agricultural University of Norway, Ås. Master of Science thesis: Futility of shooting brown bears Ursus arctos to stop sheep loss in Norway is confirmed.

D2.33 2004* Marcus Elfström, Umeå University, Umeå. Undergraduate thesis (30 ECTS): Denning ecology of Scandinavian brown bears.

D2.34 2004 Knut Madslien, Norwegian School of Veterinary Sciences, Oslo. Student thesis: Use of intraperitoneal radiotransmitters in yearling female brown bears; anesthetic and surgical procedures.

D2.35 2005* Ingela Jansson, Swedish University of Agricultural Sciences, Umeå. Undergraduate thesis (20 points): A pilot study of brown bear (Ursus arctos) habitat use in central Sweden using GPS.

D2.36 2005* Hanna Barck, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Factors affecting foraging by brown bears Ursus arctos on carpenter ants, Camponotus herculeanus.

D2.37 2005 Claudio Signer, University of Basel, Switzerland. Master of Science thesis: Multi-level habitat analsis of female brown bears (Ursus arctos) in central Sweden.

D2.38. 2005* Therese Fosholt Moe, Norwegian University of Life Sciences, Ås. Master thesis: Diel variation in habitat selection of female Scandinavian brown bears (Ursus arctos) in relation to resting and foraging behavor.

D2.39 2006 Veronica Sahlén, University College London. Master of Science thesis: Female brown bear Ursus arctos use of an anthropogenic food source: a study from south-central Sweden

D2.40 2006* Geir Rune Rauset, Norwegian University of Life Sciences, Ås. Master thesis: Estimating individual kill rates on moose calves by brown bears based on GPS technology and GIS cluster analysis.

D2.41 2006 Inger Marie Yri, Norwegian University of Life Sciences, Ås. Master thesis: Seasonal and diel variation in road avoidance behaviour of female Scandinavian brown bears.

D2.42 2007* Bjørn Erik Pedersen, Norwegian University of Life Sciences, Ås. Master thesis: Immediate and delayed behaviour of Scandinavian female brown bears when encountered by humans on foot

D2.43 2007 Kent Ove Moren, Norwegian University of Life Sciences, Ås. Master thesis: Brunbjørns Ursus arctos predasjon på elg Alces alces: Økonomiske effekter for grunneiere i sentrale Sør-Sverige (In Norwegian: Predation by brown bears on moose: economic effects for landowners in central southern Sweden).

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D2.44 2007 Jee-Yun Hyun, Seoul National University, Seoul, Republic of Korea. Masters thesis: Habitat use of female brown bears (Ursus arctos) in central Sweden (In Korean).

D2.45 2007 Maria Collett Knagenhjelm, Norwegian University of Life Sciences, Ås. Master thesisi: Bears or tears? A study of people- predator conflicts in the Bolivian Andes.

D2.46 2008* Bård Andreas Lassen, Kristian Ingdal , Telemark University College, Bø. Master of Science thesis: Chemical communication in yellow-bellied marmot (Marmota flaviventris) and brown bear (Ursus arctos).

D2.47 2008 Diala Dathe, Friedrich-Alexander Universität, Erlangen-Nürnberg, Germany. Diplomarbeit: Das Markierungsverhalten von Braunbären in süd-zentral Schweden.

D2.48 2008 Peter Marcus K. Greve, Norwegian University of Life Sciences, Ås. Master thesis: Behavior of the Scandinavian brown bear (Ursus arctos) when approached by people on foot.

D2.49 2008* Lisa Grenlund Langebro, Norwegian University of Life Sciences, Ås. Master thesis: Brown bear (Ursus arctos) bed-site selection and behaviour in relation to human settlements.

D2.50 2009* Gro Kvelprud Moen, Norwegian University of Life Sciences, Ås. Master of Science thesis: Responses by Scandinavian brown bears (Ursus arctos) to human approaches on foot.

D2.51 2009* Silje Vang, Norwegian University of Life Sciences, Ås. Master thesis: Factors affecting the ability and accuracy of dogs (Canis familiaris) to track brown bears (Ursus arctos).

D2.52 2009* Sam M. J. G. Steyaert, Wageningen University, Wageningen, Netherlands, Master of Science thesis, Habitat modelling as a predictive tool in human-wildlife conflicts: Brown bear (Ursus arctos) and free-ranging cattle in central Sweden.

D2.53 2010 Solveig Silset Berg, Norwegian University of Life Sciences, Ås. Master thesis: Small-scale habitat use by male and female brown bears (Ursus arctos) in the mating and the hyperphagy season.

D2.54 2010* Lara Budic, Norwegian University of Life Sciences, Ås. Master thesis: Brown bears in northern and southern Europe: are they seeking food or avoiding dominant bears?

D2.55 2010* Ellinor Sahlén, Swedish University of Agricultural Sciences, Umeå, Master thesis in Biology (30 points): Do grizzly bears use or avoid wellsites in west-central Alberta, Canada?

D2.56 2011* Shane Frank, University of Freiburg, Freiburg, Germany, Master of Science thesis in Forest Ecology and Management: Bears, clear-cuts and ants: predicting bear use and movement among and within clear-cuts in Sweden during summer.

D2.57 2011 David Haberkorn, Wageningen University, Wageningen, Netherlands, Master of Science thesis, Spatio-temporal analysis of brown bear (Ursus arctos) interactions in the mating season.

D2.58 2011* Cyril Milleret, Université Joseph Fourier, Grenoble, France, Master of Science thesis in Biodiversité – Écologie – Environnement. Estimating wolves (Canis lupus) and brown bear (Ursus arctos) interactions in Central Sweden. Does the emergence of brown bears affect wolf predation patterns?

D2.59 2011 Vera Menges, Edinburgh Napier University, Edinburgh. Master of Science thesis, Influence of season, sex, age and behaviour on the defecation rate of brown bears (Ursus arctos) in south-central Sweden.

D2.60 2011* Ofelia Gonzalez, Universitè de Sherbrooke, Sherbrooke, Quebec, Canada, Master thesis. Causes et consequences de la variation en tille de portée chez l’ours brun (Ursus arctos). (The determinants and effects of litter size and cub sex ratio in brown bears).

D2.61 2012* Franziska Johanna Hütter. University for Natural Resources and Applied Life Sciences, Austria, Master thesis. Fecal analysis suggests that female brown bears with dependent offspring avoid adult males during the mating season to prevent sexually selected infanticide

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D2.62 2012 Erwin Pier. Wageningen University, Wageningen, Netherlands, Master of Science thesis. Impact of climate change on bear-movement regarding berry foraging; Feedback on spatio-temporal data analyses.

D2.63 2012 Jürgen Joseph, Humboldt-Universität zur Berlin. Master of Science thesis: Zur Physiologie des Infantizides bei einem nicht sozial lebenden Carnivoren, dem Braunbären (Ursus arctos) in einer frei lebenden skandinavischen Population (On the physiology of infanticide in a non-social carnivore, the brown bear (Ursus arctos) in a Scandinavian free ranging population.

D2. 64 2012* Martin Müller. University of Natural Resources and Applied Life Sciences, Vienna. Master of Science thesis: Does food matter? Diet quality of Scandinavian brown bears in relation to their sex, age, reproductive status, and distance to settlements.

D2. 65 2012 Eva Filipczyková. Wageningen University, Wageningen, Netherlands, Master of Science thesis. To feed or to leave: The effects of bait sites on the behaviour of brown bears towards humans.

D2. 66 2012. Tommy Vestøl. Norwegian University of Life Sciences, Ås. Master thesis: Baiting for brown bears (Ursus arctos) in Sweden.

D2. 67 2013. Anna Maria Petré. Swedish University of Agricultural Sciences, Umeå. Master thesis in Biology (30 points): Bears (Ursus arctos) use of managed forests in autumn; a comparison between Sweden and Canada.

D2.68 2013* Anne Randi Græsli. University of Copenhagen. Master’s thesis in Veterinary Medicine: Haematological and biochemical reference values for medetomidine-tiletamine-zolazepam immobilized free-ranging brown bears (Ursus arctos) in Scandinavia and the influence of seasons and host characteristics.

D2.69 2013* Núria Fandos Esteruelas. Universidad de Pablo Olavide, Sevilla. Master’s thesis in Biodiversity and conservation biology: Leukocyte coping capacity as a tool for assessing capture and handling-induced stress in free-ranging Scandinavian brown bears (Ursus arctos).

D2.70 2013* Christine Reusch. University for Natural Resources and Applied Life Sciences, Vienna, Austria Master thesis: Habitat segregation in brown bears: testing the predation risk hypothesis with nutritional analyses.

D2.71 2014. Lisa A. Klestil. Universität Wien. Masterarbeit: Placental scar counts and the assessment of reproductive performance in female brown bears.

D2.72 2014* Jacinthe Gosselin. Université de Sherbrooke, Sherbrooke, Québec, Canada. Grade de maître ès sciences (M.Sc.). Influence de la chasse et de l’infanticide sur la dynamique de la population de l’ours brun scandinave (Ursus arctos).

D2.73 2014 Stine Emilie Nøding Hansen. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis. Behavior of Scandinavian brown bears when encountered by dogs and humans.

D2.74 2015. Nina Emilie Stenset. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis. Behaviour of brown bears (Ursus arctos) when repeatedly approached by humans on foot.

D2.75 2015 Juan Romairone. Pablo Olavide University, Seville, Spain. Master of Science thesis in Biodiversity and Biology of Conservation. Habitat characteristics between different clusters of wolf (Canis lupus) activity before and after brown bear (Ursus arctos) emergence in Central Sweden.

D2.76 2015 Linda Nowak. University for Natural Resources and Applied Life Sciences, Vienna, Austria Master thesis: Reproductive performance of Scandinavian female brown bears (Ursus arctos) in relation to the use of den-type.

D2.77 2015 Marion Lukkari Arnesen. Telemark University College, Bø. Master thesis: Use of experimental bait sites by wolverines.

D2.78 2015 Henriette Wathne Gelink. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource

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Management. Master Thesis: Risky berry business? Brown bear (Ursus arctos) foraging behaviour in a landscape of fear.

D2.79 2015 Hanna Kavli Lodberg-Holm. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis: When the hunter becomes the hunted: Impacts of hunting on the foraging behavior of the brown bear (Ursus arctos) in Sweden.

. Bachelor of Science level thesesD3.01 1988 Paul Klute & Tom Erik Ness, Statens skogskole, Evenstad. Undergraduate

report. Älvdalsbinna - arealbruk, forflyttninger og habitatbruk 1987. (Älvdalsbinna—use of area, movements and habitat use in 1987).

D3.02 1989 * Arne Söderberg, Öster-Malma jaktvårdsskola. Undergraduate final reseach report: Mortalitet hos älgkalvar under sommaren. (Moose calf mortality in summer; In Swedish)

D3.03 1992 Mats Ola Johnsson, Klarälvdalens Folkhögskola. Undergraduate two-year school final research report. Brunbjörn. En ankät undersökning över björnförekomst i Värmlands norra delar. Samt en sammanställning av det Svensk - Norska björnprojektet. (A questionnaire study of the occurrence of bears in northern Värmland and a description of the Swedish-Norwegian Brown Bear Project).

D3.04 1995 * Andreas Norin, Klarälvdalen Folkehögskola. Undergraduate two-year school final research report. Studier av näringsval och rörelsemönster hos radiomärkta brunbjörnar (Studies of food selection and movement patterns of radio-market brown bears; In Swedish)

D3.05 1997 Paul Anzjøn & Johan Henrik Castberg, Regional College in Nord-Trøndelag, Steinkjer, Norway. Undergraduate thesis: Omtalen av bjørn (Ursus arctos) i media i 1995. (Reporting about brown bears in the Norwegian media in 1995; In Norwegian).

D3.06 1998 Pilivi Vaajakari, Omnivet. Undergraduate final report: Reproduktionen hos brunbjörnar i Sverige. (Reproduction into Brown bear in Sweden. In Swedish)

D3.07 1999 Carina Sisell, Älvdalens naturbruksgymnasium. Undergraduate final report: Björnen, människan och turismen—går de att förena? (Bears, people and tourism—is it possible to combine them? In Swedish)

D3.08 2005 Kent Ove Moren, Høgskolen i Hedmark Bachelor thesis: Brun björnens (Ursus arctos) habitatvalg:kantsoner og habitatdiversitet. (Habitat selection in brown bears (Ursus arctos): edges and habitat diversity. In Norwegian)

D3.09 2009 Elinor Sahlén, Swedish University of Agricultural Sciences, Umeå (Examensarbete i biologi, 2009:15). Undergraduate thesis (15 points, C level): Brown bear (Ursus arctos) den site concealment in relation to human activity in Scandinavia

E) Rapports to management agencies, etc. E1 Bjärvall, A., F. Sandegren, & P. Wabakken. 1988. Rapport 1 från det svensk-norska

björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 1 1988 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E2 Bjärvall, A., F. Sandegren, & P. Wabakken. 1988. Rapport 2 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 2 1988 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E3 Bjärvall, A., F. Sandegren, & P. Wabakken. 1989. Rapport 1 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 1 1989 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E4 Bjärvall, A., F. Sandegren, & P. Wabakken. 1990. Rapport 1 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 1 from the Swedish-Norwegian Brown Bear Project. In Swedish)

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E5 Bjärvall, A., F. Sandegren, & P. Wabakken. 1990. Rapport 2 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 2 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E6 Bjärvall, A., F. Sandegren, & P. Wabakken. 1990. Rapport 3 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 3 1990 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E7 Franzén, R., F. Sandegren, & P. Wabakken. 1991. Årsrapport från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report of the year 1991 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E8 Söderberg, A., F. Sandegren. 1992. Rapport 1 från det svensk-norska björnprojektet. Våra Rovdjur Nr.2,Årg.9. (Report 1 1992 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E9 Franzén, R., F. Sandegren, & P. Wabakken. 1992. Rapport 2 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 2 1992 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E10 Wabakken, P. A. Bjärvall, R. Franzén, E. Maartmann, F. Sandegren, & A. Söderberg. 1992. Det svensk-norske bjørneprosjektet 1984-91. Norwegian Inst. Nature Res., Oppdragsmelding 146. (The Swedish-Norwegian Bear Research Project 1984-91. In Norwegian)

E11 Franzén, R., F. Sandegren, & P. Wabakken. 1993. Rapport 1 från det svensk-norska björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 1 1993 from the Swedish-Norwegian Brown Bear Project. In Swedish)

E12 Wabakken, P. & E. Maartmann. 1994. Slutrapport for bjørn-sauprosjektet i Hedmark 1990-93. Norwegian Inst. Nature Res., Forskningsrapport 58. (Final report for the bear-sheep project in Hedmark, 1990-93. In Norwegian)

E13 Swenson, J. E., F. Sandegren, P. Wabakken, A. Bjärvall, A. Söderberg and R. Franzén. 1994. Bjørnens historiske og nåværende status og forvaltning i Skandinavia. Norwegian Inst. Nature Res., Forskningsrapport 53. (The historic and present status and management of the brown bear (Ursus arctos L.) in Scandinavia; in Norwegian with English summary).

E14 Swenson, J. E., F. Sandegren, & P. Wabakken. 1994. Rapport 1 från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 1 1994 from the Scandinavian Brown Bear Project. In Swedish)

E15 Swenson, J. E., F. Sandegren, & P. Wabakken. 1994. Rapport 2 från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report 2 1994 from the Scandinavian Brown Bear Project. In Swedish)

E16 Swenson, J. E., F. Sandegren, & P. Wabakken. 1994. Årsrapport från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report of the year 1994 from the Scandinavian Brown Bear Project. In Swedish)

E17 Sagør, J. T., J. E. Swenson, and E. Røskaft. 1995. Bjørn og sauehold: Er det mulig å nå Rovviltmeldingenes målsettingene? (Bears and sheep husbandry: Is it possible to reach the goals in the Large Predator White Paper? In Norwegian). Norwegian Inst. Nature Res., Fagrapport 14.

E18 Swenson, J. E., F. Sandegren. 1995. Årsrapport från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report of the year 1995 from the Scandinavian Brown Bear Project. In Swedish)

E19 Swenson, J. E. 1995. How can research be an effective part of wildlife management?. Wirkungsvoller Naturschutz – Welche Rolle kann die SGW übernehmen? Jahrestagung der Schweizerischen Gesellschaft für Wildtierbiologie (SGW) 1995. S 4-9

E20 Sandegren, F., S. Brunberg, J. Swenson & A. Söderberg. 1996. Rapport angående arbetet med björnhonan Koska och hennes ungar. Rapport till länsstyrelsen i Dalarna. 3 pp. (Report about work with the bear female “Koska” and her young. In Swedish)

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E21 Swenson, J. E., F. Sandegren. 1996. Björnlakten i Sverige – Några rekomendationer. Rapport till Naturvårdsverket.

E22 Swenson, J. E., F. Sandegren & A. Söderberg. 1996. Utvärdering av björnobsen. Rapport till Svenska Jägareförbundet.

E23 Swenson, J. E., F. Sandegren, M. Heim, S. Brunberg, O. J. Sørensen, A. Söderberg, A. Bjärvall, R. Franzén, S. Wikan, P. Wabakken, and K. Overskaug. 1996. Er den skandinaviske bjørnen farlig? Norwegian Inst. Nature Res., Oppdragsmelding 404. (Is the Scandinavian brown bear dangerous? In Norwegian)

E24 Andersen, R., J. Linnell, and J. Swenson. 1996. Hovedrapport—Regionfelt Østlandet. Tema hjortevilt og rovvilt. Norwegian Inst. Nature Res., Oppdragsmelding 405. (Final report—Regionfelt Østlandet. Ungulates and carnivores. In Norwegian)

E25 Aanes, R., J. D. Linnell, J. E. Swenson, O. G. Støen, J. Odden, and R. Andersen. 1996. Menneskelig aktivitets innvirkning på klauvvilt og rovvilt. En utredning foretatt i forbindelse med Forsvarets planer for Regionfelt Østlandet, del 1. Norwegian Inst. Nature Res., Oppdragsmelding 412. (The effect of human disturbance on ungulates and carnivores: a study undertaken in connection with the proposal by the Department of Defense to establish a shooting range in southeastern Norway, part 1, in Norwegian)

E26 Linnell, J. D. C., B. Barnes, J. E. Swenson, and R. Andersen. 1996. Hvor sårbare er bjørner for forstyrrelser i hiperioden? En utredning foretatt i forbindelse med Forsvarets planer for Regionfelt Østlandet, del 2. Norwegian Inst. Nature Res., Oppdragsmelding 413. (How vulnerable are denning bears to disturbance? a study undertaken in connection with the proposal by the Department of Defense to establish a shooting range in southeastern Norway, part 2, in Norwegian)

E27 Swenson, J. E., T. M. Heggberget, P. Sandström, P. Wabakken, A. Bjärvall, A. Söderberg, R. Franzén, J. D. C. Linnell, and R. Andersen. 1996. Brunbjørnens arealbruk i forhold til menneskelig aktivitet. En utredning foretatt i forbindelse med Forsvarets planer for Regionfelt Østlandet, del 5. Norwegian Inst. Nature Res., Oppdragsmelding 416. (Use of area by brown bears in relation to human activity: a study undertaken in connection with the proposal by the Department of Defense to establish a shooting range in southeastern Norway, part 5, in Norwegian)

E28 Mysterud, I., J. E. Swenson, J. D. C. Linnell, A. O. Gautestad, I. Mysterud, J. Odden, M. E. Smith, R. Aanes, P. Kaczensky. 1996. Rovvilt og sauenæring i Norge: kunnskapsoversikt og evaluering av forebyggende tiltak. Report, Biological Institute, University of Oslo, Norway. (Carnivores and sheep farming in Norway: a survey of the information and evaluation of mitigating measures. In Norwegian)

E29 Aanes, R., J. E. Swenson, and J. D. C. Linnell. 1996. Rovvilt og sauenæring i Norge. 1. Tap av sau til rovvilt: en presentasjon av tapets omfang basert på brukeropplysninger. Norwegian Inst. Nature Res., Oppdragsmelding 434. (Carnivores and sheep farming in Norway. 1. Loss of sheep due to large predators: a presentation of the magnitude of loss based on information from sheep farmers. In Norwegian)

E30 Linnell, J. D. C., M. E. Smith, J. Odden, P. Kaczensky, and J. E. Swenson. 1996. Rovvilt og sauenæring i Norge (Carnivores and sheep farming in Norway). 4. Strategies for the reduction of carnivore—livestock conflicts: a review. Norwegian Inst. Nature Res., Oppdragsmelding 443. (In English)

E31 Linnell, J. D. C., M. E. Smith, J. Odden, P. Kaczensky, and J. E. Swenson. 1996. Rovvilt og sauenæring i Norge (Carnivores and sheep farming in Norway). 5. Strategier for å redusere rovdyr-husdyr konflikter: en litteraturoversikt. Norwegian Inst. Nature Res., Oppdragsmelding 444. (In Norwegian with English summary)

E32 Sæther, B.-E., S. Engen, J. E. Swenson, Ø. Bakke, and F. Sandegren. 1997. Levedyktighetsanslyser av skandinavisk brunbjørn. Norwegian Inst. Nature Res., Fagrapport 25. (Population viability analysis for the Scandinavian brown bear. In Norwegian).

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E33 Swenson, J. E., F. Sandegren. 1997. Årsrapport från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report of the year 1997 from the Scandinavian Brown Bear Project. In Swedish)

E34 Linnell, J. D. C., J. E. Swenson, A. Landa & T. Kvam. 1998. Methods for monitoring European large carnivores—a worldwide review of relevant experience Norwegian Inst. Nature Res., Oppdragsmelding 549.

E35 Linnell, J. D. C., J. E. Swenson, A. Landa & T. Kvam. 1998. Metodikk for bestandsovervåking av store rovdyr—en litteraturgjennomgang. Norwegian Inst. Nature Res., Oppdragsmelding 550. (Methods for monitoring European large carnivores—a worldwide review of relevant experience. In Norwegian)

E36 Brunberg, S., S. Nordlund & F. Sandegren. 1998. Rapport om björnen W9705 Björe. The Scandinavian Brown Bear Research Project, Unpubl. Rept. (Report about the bear W9705 Björe. In Swedish).

E37 Swenson, J. E., F. Sandegren. 1998. Årsrapport från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report of the year 1998 from the Scandinavian Brown Bear Project. In Swedish)

E38 Swenson, J. E., F. Sandegren, A. Bjärvall, P. Wabakken, R. Franzén & A. Söderberg. 1998. Bjørneungers oppvekstvilkår: hvilke faktorer er viktige for overlevelse av årsunger? Pages 180-184 in NINAs strategike instituttprogrammer 1991-95: Store rovdyrs økologi i Norge, Slutrapport. Norwegian Inst. Nature Res., Temahefte 8. (In Norwwegian)

E39 Swenson, J. E., F. Sandegren, P. Wabakken, A. Bjärvall, A. Söderberg R. Franzén & P. Segerström. 1998. Bjørneungers spredningsdynamikk. Pages 185-188 in NINAs strategike instituttprogrammer 1991-95: Store rovdyrs økologi i Norge, Slutrapport. Norwegian Inst. Nature Res., Temahefte 8. (In Norwwegian)

E40 Dahle, B., J. E. Swenson, O. J. Sørensen & E. H. Wedul.. 1998. Næringvalg hos brunbjørn i Sør-Skandinavia. Pages 189-195 in NINAs strategike instituttprogrammer 1991-95: Store rovdyrs økologi i Norge, Slutrapport. Norwegian Inst. Nature Res., Temahefte 8. (In Norwegian)

E41 Swenson, J. & F. Sandegren. 1999. Mistänkt illegal björnjakt i Sverige. Pages 201-206 in Bilagor till Sammanhållen rovdjurspolitik; Slutbetänkande av Rovdjursutredningen. Statens offentliga utredningar 1999:146. Stockholm, Sweden (Suspected illegal hunting of bears in Sweden. In Swedish)

E42 Swenson, J. E., F. Sandegren. 1999. Årsrapport från det Skandinaviska Björnprojektet. Rapport Svenska Jägareförbundet och Naturvårdsverket. (Report of the year 1999 from the Scandinavian Brown Bear Project. In Swedish)

E43 Swenson, J. & F. Sandegren. 2000. Ekologi och förvaltning av brunbjörnen i Skandinavien: slutrapport från Det skandinaviska björnprojektet. Rapport til Naturvårdsverket. 19 pp. (Ecology and management of the brown bear in Scandinavia: final report from the Scandinavian Brown Bear Research Project. In Swedish)

E44 Sandegren, F., J. Swenson. 2000. Comments to the swedish environmental protection agency from the scandinavian brown bear research project regarding the brown bear hunting season in 2000. Report from the Scandinavian Brown Bear Research Project to the Swedish Environmental Protection Agency. 2 pp.

E45 Swenson, J. E., B. Dahle and F. Sandegren. 2001. Bjørnens predasjon på elg. Norwegian Inst. Nature Res., Fagrapport 48. (In Norwegian: Brown bear predation on moose)

E46 Kristoffersson, M., S. Brunberg & J. Swenson. 2001. Försök att spåra björn med tillgängliga björneftersöksekipage under sommaren 2000. Rapport från Viltskadecenter. (Trials to track moose with available hunter and dog tracking teams during summer 2000. In Swedish).

E47 Linnell, J., Steuer, D., Odden, J., Kaczensky, P. & Swenson, J.E. 2002. European brown bear compendium. Safari Club International Foundation, Wildlife Conservation Issues Technical Series. 004A: 1-131.

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(http://www.lcie.org/Publicaciones/BB/European%20Brown%20Bear%20Compendium.pdf)

E48 Swenson, J. E., B. Dahle, J. M. Arnemo, S. Brunberg, H. Hustad, E. Nerheim, F. Sandegren, K. H. Solberg, and A. Söderberg. 2003. Utredninger i forbindelse med ny rovviltmelding: Status og forvaltning av brunbjørnen i Norge (In Norwegian with English summary: Reports for the large predator policy statement: Status and management of brown bears in Norway). Norwegian Institute for Nature Research, Fagrapport 60.

E49 Solberg, E. J., H. Sand, J. D. C. Linnell, S. M. Brainerd, R. Andersen, J. Odden, H. Brøseth, J.Swenson, Olav Strand, and P. Wabakken. 2003. Utredninger i forbindelse med ny rovviltmelding: Store rovdyrs innvirkning på hjortevilt i Norge: Økologiske prosesser og konsekvenser for jaktuttak og jaktutøvelse (In Norwegian with English summary: Reports for the large predator policy statement: The effects of large carnivores on wild ungulates in Norway: implications for ecological processes, harvest and hunting methods). Norwegian Institute for Nature Research, Fagrapport 63.

E50 Arnemo, J. M., S. Brunberg, and J. Swenson. 2003. Medetomidine-tiletamine-zolazepam in brown bears (Ursus arctos): test of high doses in captive animals. Report from the Scandinavian Brown Bear Research Project, Noppikoski, Sweden.

E51 Andersen, R., T. Bjerke, S. M. Brainerd, I. Bruteig, H. Brøseth, J. O. Gjershaug, H. Hustad, A. Landa, J. D. C. Linnell, T. Nygård, H. C. Pedersen, K. Skogen, E. J. Solberg, H. Svarstad, J. E. Swenson, B.-E. Sæther and T. Tveraa. 2003. Utredninger i forbindelse med ny rovviltmelding: Fremtidige FoU-oppgaver knyttet til store rovdyr i Norge. (In Norwegian with English summary: Reports for the large predator policy statement: Research and development requirements related to large carnivores in Norway). Norwegian Institute for Nature Research, Fagrapport 67.

E52 Bellemain, E. 2003. Results from the genetic analyses performed on faeces and hairs samples from Norwegian bears. Rapport to the Directorate for Nature Management. Laboratoire d’Ecologie Alpine, Grenoble.

E53 Lande, U. S., J. D. C. Linnell, I. Herfindal, V. Salvatori, H. Brøseth, R. Andersen, J. Karlsson, T. Willebrand, J. Persson, A. Landa, R. May, B. Dahle & J. Swenson. 2003. Utredninger i forbindelse med ny rovviltmelding. Potensielle leveområder for store rovdyr i Skandinavia: GIS-analyser på et økoregionalt nivå. Norwegian Institute for Nature Research, Fagrapport 64. (Report to the large predator policy statement. Potential habitat for large carnivores in Scandinvia: a GIS analysis at the ecoregion level.)

E54 Kindberg, J., J. Swenson, S. Brunberg, & G. Ericsson. 2004. Preliminär rapport om populationsutvickling och –storlek av brunbjörn i Sverige, 2004. Report from the Scandinavian Brown Bear Research Project to the Swedish Environmental Protection Agency. (Preliminary report on the trend and size of the brown bear poulation in Sweden, 2004, In Swedish).

E55. Swenson, J. E., J. M. Arnemo, E. Bellemain, S. Brunberg, B. Dahle, O.-M. Drageset, H. Hustad, J. Katajisto, J. Kindberg, E. Nerheim, K. H. Solberg, P. Segerström, O.-G. Støen, A. Söderberg and A. Zedrosser. 2005. Rovvilt og Samfunn (RoSa)—Det skandinaviske bjørneprosjektet. Oversikt over gjennomførte aktiviteter, Sluttrapport til Norges forskningsråd. (In Norwegian with English summary: Large Carnivores and Society (RoSa)—The Scandinavian Brown Bear Research Project. Overview of completed activites; Final report the the Research Council of Norway). Norwegian Institute for Nature Research, NINA Rapport 31.

E56. Andrén, H., H. Lindén, A. Lundvall, B.-E. Sæther, and J. Swenson. 2005. Utvärdering av Norges nationella övervakningsprogram för stora rovdjur. Direktoratet for naturforvaltning, Utredning 2005-7. (In Swedish; Evaluation of Norway’s national monitoring program for large carnivores)

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E57. Bellemain, E. and P. Taberlet. 2005. Results from the genetic analyses performed on feces samples from bears in Västernorrland, January-May 2005. Report from Laboratiore d’Ecologie Alpine, Université Joseph Fourier, Grenoble.

E58. Durand, A. and P. Taberlet. 2005. Results from the genetic analyses performed on feces samples from bears in Västerbotten county, Sweden. Report from Laboratiore d’Ecologie Alpine, Université Joseph Fourier, Grenoble.

E59. Swenson, J. E., R. Fujita, A. Zedrosser and A. Söderberg. 2005. The use of hunting methods for brown bear hunting in Sweden and the effect of the ban on baiting. Rapport från Skandinaviska Björnprojektet till Naturvårdsverket.

E60. Kindberg, J., J.E.Swenson, G. Ericsson. 2005 Populationsutveckling för björnstammen i Sverige 2005. Rapport till Naturvårdsverket från Skandinaviska björnprojektet (in swedish)

E61. Swenson, J. E. and J. Katajisto. 2005. Estimating the total population size of brown bears in an area based on the number of annual reproductions. Report No. 2005-1 from the Scandinavian Brown Bear Research Project.

E62. Fahlman, Å. 2005. Pilotstudie:Fysiologisker effekter av sövning av vilda djur. Slutrapport till Naturvårdsverket. (In Swedish)

E63. Taberlet, P. 2005. Results from the genetic analyses performed on tissue samples from bears shot during 2005 in Västernorrland County, Sweden Persons in charge of genetics analyses. Report from Laboratiore d’Ecologie Alpine, Université Joseph Fourier, Grenoble.

E64. Arnemo, J. M. (editor), Ahlqvist, P., Andersen, R., Andrén, H., Brunberg, S., Dypsund, P., Fahlman, Å., Gangås, L., Landa, A., Liberg, O., Odden, J., Persson, J., Ranheim, B., Segerström, P., and. Swenson, J. E. 2006. Biomedical protocols for free-ranging brown bears, gray wolves, wolverines and lynx. Norwegian School of Veterinary Science, Tromsø, Norway.

E65. Brainerd, S., D. Bakka, J. Swenson and S. Brunberg. 2006. Jakt på bjørn i Norge. Norges Jeger- og Fiskerforbund.

E66. Brunberg, S. and J. Swenson. 2006. Inventering av björn med hjälp av spillningsmetoden. Report No. 2006-1 from the Scandinavian Brown Bear Research Project. (In Swedish; Estimate bear population size from collection of feces).

E67. Kindberg, J. and J. Swenson. 2006. Populationsberäkning av björnstammen i Sverige 2005. Report No. 2006-2 from the Scandinavian Brown Bear Research Project. (In Swedish; Estimate of the bear population in Sweden in 2005).

E68. Kindberg, J. and J. E. Swenson. 2006. Results from the genetic analyses performed on feces samples from bears in Västerbotten County, Sweden. Part II Population estimate. Report No. 2006-3 from the Scandinavian Brown Bear Research Project.

E69. Sahlén, V., J. Swenson, S. Brunberg and J. Kindberg. 2006. Björnen i Sverige. En rapport från Skandinaviska Björnprojektet till den svenska Rovdjursutredningen. Report No. 2006-4 from the Scandinavian Brown Bear Research Project.

E70. Swenson, J. E. 2007. The Scandinavian brown bear, summary of knowledge and research needs. Report to the Wildlife Research Committee, Swedish Environmental Protection Agency. Report No. 2007-1 from the Scandinavian Brown Bear Research Project.

E71. Arnemo, J. M.and Fahlman, Å (editors), Ahlqvist, P., Andersen, R., Andrén, H., Brunberg, S., Landa, A., Liberg, O., Odden, J., Persson, J., Segerström, P., and. Swenson, J. E. 2007. Biomedical protocols for free-ranging brown bears, gray wolves, wolverines and lynx. Norwegian School of Veterinary Science, Tromsø, Norway.

E72. Taberlet, P. 2007. Results from the genetic analyses performed on 3000 faeces samples from bear in Jämtland country, Sweden January - October 2007. Report from Laboratiore d’Ecologie Alpine, Université Joseph Fourier, Grenoble.

E73. Sahlén, V. 2007. Åteljakt på björn—status och erfarenheter från Nordamerika och Europa (inklusive Ryssland). (In Swedish with English summary: Bait hunting of

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bears; status and experiences from North America and Europe (including Russia). Report No. 2007-2 from the Scandinavian Brown Bear Research Project.

E74. Swenson, J. E. and S. Brunberg. 2008. Årsrapport fra Det skandinaviske bjørneprosjektet 2007. (In Norwegian: Annual report of the Scandinavian Brown Bear Research Project 2007). Report No. 2008-1 from the Scandinavian Brown Bear Research Project.

E75. Støen, O.-G. 2008. Årsrapport fra prosjektet: The effects of human disturbance on brown bears; an experimental approach 2007. (In Norwegian: Annual report of the project: The effects of human disturbance on brown bears; an experimental approach 2007). Report No. 2008-2 from the Scandinavian Brown Bear Research Project.

E76. Swenson, J. E., J. Kindberg and S. Brunberg. 2008. Jaktens effekter på bjørnebestanden i Dalarna. (In Norwegian: The effects of hunting on the brown bear population in Dalarna, Sweden). Report No. 2008-3 from the Scandinavian Brown Bear Research Project.

E 77. Kindberg, J. and Swenson, J.E. 2008. Resultat från Spillningsinventering av björn i Jämtland 2006 . Populationsberäkning. Report No. 2008-4 from the Scandinavian Brown Bear Research Project.

E 78. Swenson, J.E. and Støen, O.-G. 2008. Possibilities to estimate the population size of brown bears in Norway area based on the number of annualreproductions. Report No. 2008-5 from the Scandinavian Brown Bear Research Project

E 79.  Swenson, J. E., V. Sahlén, S. Brunberg and J. Kindberg. 2008. Björnen i Sverige – kunskapsläget idag. Slutrapportering från Skandinaviska Björnprojektet till Naturvårdsverket (Viltforskningskommitten). (In Swedish: The Bear in Sweden – our current knowledge. Final report to the Wildlife Research Committee, Swedish Environmental Protection Agency).  Report No. 2008-6 from the Scandinavian Brown Bear Research Project.

E 80. Vang, S., A. Zedrosser, J. Swenson and S. Brunberg. 2009. Sporing av bjørn—en empirisk studie av ettersøksekvipasjer på bjørn sommeren 2007 og 2008. (In Norwegian: Tracking bears; an empirical study of tracking bears by dog-man teams in summer 2007 and 2008. Report No. 2009-1 from the Scandinavian Brown Bear Research Project.

E81. Kindberg, J., J. E. Swenson and G. Ericsson. 2009. Björnstammens storlek i Sverige 2008 — länsvisa uppskattningar och trender. (In Swedish: The size of the brown bear population in Sweden in 2008; estimates of size and trend for each county.) Report No. 2009-2 from the Scandinavian Brown Bear Research Project.

E82. Bischof, R. and J. E. Swenson. 2009. Preliminary predictions of the effect of increasing hunting quotas on brown bear population growth in Sweden. Report No. 2009-3 from the Scandinavian Brown Bear Research Project.

E83. Aarnes, S. G., E. Bellemain, H. G. Eiken, and I. Wartiainen. 2009. Interlaboratory comparison of genetic profiles of brown bears from Sweden (Laboratoire d’Ecologie Alpine) and Norway (Bioforsk Svanhovd). Bioforsk Report Vol. 4, No. 133.

E84. Nygård, T., Brainerd, S. M., Brøseth, H., Bertsen, F., Bjørnes, E., Kvam, T., Nilsen, E. B., Pedersen, P. H., Stokke, S., Sørensen, O. J. & Swenson, J. E. 2009. Bjørn i Nord-Trøndelag. (In Norwegian: The brown bear in Nord-Trøndelag, central Norway). Norwegian Institute for Nature Research, NINA Rapport 472.

E85. Karamanlidis, A., and Zedrosser, A. 2009. Status of brown bears (Ursus arctos) in Albania and FYROM. Final report of a project supported by a research and conservation grant from the International Bear Association (IBA) on the status of brown bears (Ursus arctos) in Albania and the Former Yugoslav Republic of Macedonia (FYROM), 53pp

E86. Bischof, R. and J. E. Swenson. 2010. Estimating the number of annual reproductions based on the number of female brown bears documented in Norway in 2008 and 2009. Report No. 2010-1 from the Scandinavian Brown Bear Research Project.

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E.87. Moen, G. K. og O.-G. Støen. 2010. Statusrapport for prosjektet: Metodeutvikling for studier av bjørners predasjon på klauvdyr. Rapport 2010 – 2 fra Det skandinaviske bjørneprosjektet.

E88. Swenson, J. E., O.-G. Støen, A. Zedrosser, J. Kindberg, S. Brunberg, J. M. Arnemo & V. Sahlén. 2010. Bjørnens status and økologi i Skandinavia. Report No. 2010-3 from the Scandinavian Brown Bear Research Project.

E89. Kindberg, J. & J. E. Swenson. 2010. Beräkning av björnstammens storlek i Västerbotten 2009. Report No. 2010-4 from the Scandinavian Brown Bear Research Project.

E90. Kindberg, J. & J. E. Swenson. 2010. Skattning av björnstammens storlek i Sverige åren 1994 och 2000. Report No. 2010-5 from the Scandinavian Brown Bear Research Project.

E91. Steyaert, S. M. J. G. & S. C. Frank. 2010. Wolf-brown bear interactions in Central Sweden; pilot study 05-06 2010. Report No. 2010-6 from the Scandinavian Brown Bear Research Project.

E92. Støen, O.-G. 2010. Årsrapport—Brown bear behaviour and human perceptions of human bear encounters – implications for management (2009-2012). Kontraktnr: 06040039, 2009. Report No. 2010-7 from the Scandinavian Brown Bear Research Project.

E93. Ericsson, G., Sandström, C., Kindberg, J. & Støen, O-G. 2010. Om svenskars rädsla för stora rovdjur, älg och vildsvin. Rapport 2010:1. Institutionen för vilt, fisk och miljö, SLU, Umeå. (In Swedish: The Swedes’ fear of large carnivores, moose, and wild boar).

E94. Arnemo, J. M., A. Evans & Å. Fahlman, (editors), Ahlqvist, P., Andrén, H., Brunberg, S., Liberg, O., Linnell, J. D. C., Odden, J., Persson, J., Sand, H., Segerström, P., Sköld, K., Strømseth, T. H., Støen, O.-G., Swenson, J. E & Wabakken, P. 2011. Biomedical protocols for free-ranging brown bears, wolves, wolverines and lynx. Report.

E95. Sand, H., N. Jonzén, H. Andrén, J. Månsson, J. E. Swenson, & Jonas Kindberg. 2011. Beskattningsstrategier på älg med och utan rovdjur. Swedish University of Agricultural Sciences, Dnr. SLU ua.FE. 2011.1.5-711.

E96. Støen, O.-G. 2011. Årsrapport fra prosjektet: Brown bear behaviour and human perceptions of human bear encounters – implications for management (2009-2012). Kontraktnr: 06040039, 2010. Report No. 2011-1 from the Scandinavian Brown Bear Research Project.

E97. Swenson, J. E. & J. Kindberg. 2011. Arealkrav til en hunnbjørnbestand med 20 ynglinger årlig i Norge. Report No. 2011-2 from the Scandinavian Brown Bear Research Project.

E98. Swenson, J. E., A. Zedrosser, S. Brunberg & P. Segerström. 2011. Causes of mortality, especially illegal killing, among Swedish brown bears, 1984-2010. Report No. 2011-3 from the Scandinavian Brown Bear Research Project.

E99. Støen, O.-G. 2011. Statusrapport for prosjektet: Metodeutvikling for studier av bjørners predasjon på klauvdyr. Report No. 2011-4 from the Scandinavian Brown Bear Research Project.

E100. Swenson, J. E. & J. Kindberg. 2011. The management and conservation of large carnivores in Sweden and Norway. Report No. 2011-5 from the Scandinavian Brown Bear Research Project.

E101. Kindberg, J. & J. E. Swenson. 2011. Beräkning av björnstammens storlek i Norrbotten 2010. Report No. 2011:6 from the Scandinavian Brown Bear Research Project.

E102. Swenson, J. E. & J. Kindberg. 2011. Final Report: The Scandinavian Brown Bear Research Project, 2009-2011. Report No. 2011:7 from the Scandinavian Brown Bear Research Project.

E103. Swenson, J. E. & J. Kindberg. 2011. Slutrapport: Skandinaviska Björnprojektet 2009-2011. Report No. 2011:8 from the Scandinavian Brown Bear Research Project.

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E104. Sand, H., N. Jonzén, H. Andrén, J. Månsson, J. E. Swenson, & J. Kindberg. 2011. Strategier för beskattning av älg, med och utan rovdjur. SLU-rapport. 47 s. http://www.slu.se/algforvaltning , Dnr. SLU ua.FE. 2011.1.5-711.

E105. Sand, H., H. Andrén, J. E. Swenson, & J. Kindberg. 2011. Adaptiv älgförvaltning nr 16: Flera jägare på älgpopulationen – predationsmönster hos varg och björn. Fakta Skog, Rön från Sveriges Lantbruksuniversitet Nr 25, 2011.

E106. Arnemo, J. M., A. Evans & Å. Fahlman, (editors), Ahlqvist, P., Andrén, H., Brunberg, S., Liberg, O., Linnell, J. D. C., Odden, J., Persson, J., Sand, H., Segerström, P., Sköld, K., Strømseth, T. H., Støen, O.-G., Swenson, J. E & Wabakken, P. 2012. Biomedical protocols for free-ranging brown bears, wolves, wolverines and lynx. Report.

E107. Støen, O.-G. 2012. Årsrapport fra prosjektet: Brown bear behaviour and human perceptions of human-brown bear encounters – implications for management (2009-2012). Kontraktnr: 06040039-3. 2011. Report No. 2012:1 from the Scandinavian Brown Bear Research Project.

E108. Zedrosser, A & J. Kindberg. 2012. Åtling Report No. 2012:2 from the Scandinavian Brown Bear Research Project.

E109. Zedrosser, A., R. Bischof & J. E. Swenson. 2012. The influence of the regulation protecting families of brown bears from hunting for the bear population in Sweden. Report No. 2012:3 from the Scandinavian Brown Bear Research Project.

E110. Swenson, J. E. 2012. The importance of genetic analyses of hunter-killed brown bears for management and research in Scandinavia. Report No. 2012:4 from the Scandinavian Brown Bear Research Project.

E111. Støen, O.-G. 2012. Årsrapport fra prosjektet: Brown bear behaviour and human perceptions. Kontraktnr: 12040017 2012. Report No. 2012:5 from the Scandinavian Brown Bear Research Project.

E112. Kindberg, O.-G. Støen, G. R. Rauset & J. Karlsson. 2012. Brunbjörnars predation på älgkalvar i Norrbottens län—rapport utarbetad på uppdrag av regeringen (L2011/1478). Rapport, Sveriges lantbruksuniversitet, Institutionen för vilt, fisk och miljö, Umeå.

E113. Støen, O.-G. 2012. Metodeutvikling for studier av bjørners predasjon på klauvdyr. INA fagrapport 21. Institutt for naturforvaltning, Universitetet for miljø- og biovitenskap, Ås.

E114. Karlsson, J., O.-G. Støen, P. Segerström, R. Stokke, L.-T. Persson, L.-H. Stokke, S. Persson, N. A. Stokke, A. Persson, E. Segerström, G.-R. Rauset, J. Kindberg, R. Bischof, T. R. Sivertsen, A. Skarin, B. Åhman, I. Ängsteg, & J. Swenson. 2012. Björnpredation på ren och potentiella effekter av tre förebyggande åtgärder. Rapport från Viltskadecenter 2012:6.

E115. Zedrosser, A., J. Kindberg, and J. E. Swenson. 2013. Final report of bait hunting trials in Västerbotten, Norrbotten, and Dalarna counties, Sweden, 2010-2012. Report No. 2013:1 from the Scandinavian Brown Bear Research Project.

E116. Kindberg, J. & J. E. Swenson. 2013. Jaktens inverkan på björnstammen i Norrbottens län. Report No. 2013:2 from the Scandinavian Brown Bear Research Project.

E117. Zedrosser, A., S. M. J. G. Steyaert, S. Brunberg, J. E. Swenson & J. Kindberg. 2013. The effects of baiting for hunting purposes on brown bears and their behavior. Report No. 2013:3 from the Scandinavian Brown Bear Research Project.

E118. Kindberg, J. & J. E. Swenson. 2013. Beräkning av björnstammens storlek i Värmland, Dalarnas och Gävleborgs län. Report No. 2013:4 from the Scandinavian Brown Bear Research Project.

E119. Støen, O.-G. 2013. Årsrapport fra prosjektet: Brown bear behaviour and human perceptions. Kontraktnr: 12040017 2013. Report No. 2013:5 from the Scandinavian Brown Bear Research Project.

E120. Steyaert, S. 2014. Effects of GPS relocation intervals on behavior metrics in brown bears. Report No. 2014:1 from the Scandinavian Brown Bear Research Project.

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E121. Kindberg, J. & J. E. Swenson. 2014. Björnstammens storlek i Sverige 2013 – länsvisa skattningar och trender. Rapport 2014:2 from the Scandinavian Brown Bear Research Project.

E122. Zedrosser, A., M. Lukkari-Arnesen, S. M. J. G. Steyaert, J. Kindberg, J. Persson, M. Aronsson, and J. E. Swenson. 2014. Use of experimental bait sites by wolverines in Dalarna, Gävleborg and Västerbotten counties, 2010-2012, final report. Rapport 2014:3 from the Scandinavian Brown Bear Research Project and the Swedish Wolverine Project.

E123. Støen, O.-G & M. Johansson. 2014. Årsrapport fra prosjektet: Brown bear behaviour and human perceptions. Kontraktnr: 12040017 2014. Report No. 2014:4 from the Scandinavian Brown Bear Research Project.

E124. Linnell, J. D. C., O.-G. Støen, I. Hansen, S. Eilertsen, Ø. Flagstad, V. Gervasi, H. Brøseth, J. Swenson, J. & J. Odden. 2015. Utredning om endring i yngleområdene for jerv og bjørn i rovvilt region 6. NINA Rapport 1123. 36 s.

E125. Boitani, L. et al. 2015. Key actions for large carnivore populations in Europe. Report to DG Environment, European Commission. Contract 07.0307/2013/654446/SER7B3.

E126. Brunberg, S. 2015. Studien om överlevnad av ensamma årsungar 2014. Rapport 2015:1 from the Scandinavian Brown Bear Research Project.

E127. Swenson, J. E. and J. Kindberg. 2015. Final Report: The Scandinavian Brown Bear Research Project 2012-2014. Rapport 2015:2 from the Scandinavian Brown Bear Research Project.

E128. Johansson; M., I. Ferreira, O.-G. Støen, J. Frank, and A. Flykt. 2015. Interventions targeting human fear of large carnivores – many ideas but scarce data. Rapport 2015:3 from the Scandinavian Brown Bear Research Project.

E129, Elfström, M. and O.-G. Støen. 2015. Occurrence of large carnivores near settlements: a review of mechanisms and preventive measures. Rapport 2015:4 from the Scandinavian Brown Bear Research Project.

E130. Bischof, R. 2015. Approaches for assessing illegal hunting of brown bears and other large carnivores in Sweden. Rapport 2015:5 from the Scandinavian Brown Bear Research Project.

E131. Kindberg, J. & J. E. Swenson. 2015. Björnstammens storlek i Västerbotten 2014. Rapport 2015:6 from the Scandinavian Brown Bear Research Project.

E132. Linnell, J. D. C., T. Tveraa (redaktører), J. Swenson, O. G. Støen, I. Hansen, H. Andrén, J. Persson, H. Sand, C. Wikenros, B. Zimmermann, J. Odden, A. Stien, B. Ytrehus, O. Kleven, H. Brøseth, J. Kindberg, J. Mattisson, E. J. Solberg, E. B. Nilsen, T. Nygård, K. O. Jacobsen. 2015. Kunnskapsstatus og kunnskapsbehov for forvaltning av rovvilt i Norge. 2015. – NINA Rapport 1195. 80 s.

E133. Anon. 2015. Bidrag till Norsk institutt for naturforskning (NINA) för arbete inom Skandinaviska björnprojektet säsong 2014 (NV-04578-14). Report from the Scandinavian Brown Bear Project 2015-02-31.

E134. Swenson, J. E. and J. Kindberg. (2015). Effects of combined predation by brown bears and gray wolves on moose population dynamics; what more do we need to know? Bidrag till Norsk institutt for naturforskning (NINA) för arbete inom Skandinaviska björnprojektet säsong 2014 (NV-04578-14). Report from the Scandinavian Brown Bear Project.

F) Popular publicationsF1 Bengtsson, G. & F. Sandegren. 1985. Björnen anfaller. Svensk Jakt. 123:726-729.

(The bear attacks. In Swedish)F2 Bjärvall, A. & P. Ahlqvist. 1985. Första björnen med radiosändare. Svensk Jakt.

123:278-281. (The first bear with a radio transmitter. In Swedish)F3 Sandegren, F. & A. Bjärvall. 1985. Här radiomärks en bjässe. Svensk Jakt. 123:622-

623. (A hefty chap is radio-marked. In Swedish)

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F4 Ahlqvist, P. & G. Markgren. 1987. Ett björnmärkningsprojekt. Viltnytt 24:6-12. (A bear-marking project. In Swedish)

F5 Bjärvall, A. & P. Westman. 1987. Licensjakten på björn. Svensk Jakt. 125:986-989. (The license system for bear hunting. In Swedish)

F6 Bjärvall, A., F. Sandegren, P. Ahlqvist, R. Franzén, O. Persson, & L. Pettersson. 1987. Björnmärkningen. Svensk Jakt. 125:208-213. (Bear marking. In Swedish)

F7 Pettersson, L. & F. Sandegren. 1987. Hur gammal blir björnen? Från Hav till Fjäll 11(3):20-21. (How old do bears become? In Swedish)

F8 Bjärvall, A. & F. Sandegren. 1988. Vi behöver veta mer om björnen. Dalajägaren 40 år. Pp. 82-92. (We must know more about bears. In Swedish)

F9 Bjärvall, A. 1989. Alltjämt ovisst för de fyra. Sveriges Natur 80(3):3. (Still uncertain for the four large predators. In Swedish)

F10 Bjärvall, A. 1989. Björnens vandringar klarnar. Svensk Jakt 127:902-904. (The wanderings of bears become more clear. In Swedish)

F11 Sandegren, F. & A. Söderberg. 1989. Provtagning på björn. Svensk Jakt 127:878. (Taking samples from bears. In Swedish)

F12 Wabakken, P., A. Bjärvall, & F. Sandegren. 1989. Bjørner med radio tvers over riksgrensa—et felles svensk-norsk forskningsprosjekt. Jakt og Fiske 118(4):38-39. (Bears with radio transmitters over the national border—a common Swedish-Norwegian research project. In Norwegian)

F13 Bjärvall, A. 1990. Projekt björn. Camera Natura 1(3):2-9. (Project Bear. In Swedish)

F14 Wabakken, P. & A. B. Vaag. 1990. Bjørneprosjektet. Naturnytt 2:40-41. (The bear project. In Norwegian)

F15 Sandegren, F. & A. Söderberg. 1991. Björnens idegång i Norrbotten. Jakt i norr 1:5-7. (Denning by bears in Norrbotten. In Swedish)

F16 Sandegren, F. & A. Söderberg. 1991. Norrbottenbjörnars hemområde i september. Jakt i norr 1:7-8. (The September home ranges of bears in Norrbotten. In Swedish)

F17 Wabakken, P. 1991. Hedmark Sau- og geitalslag aktivt med i bjørneforskningen. Sau og Geit 44:108-111. (The Hedmark Sheep and Goat Raisers Organization participate actively in bear research. In Norwegian)

F18 Söderberg, A. & F. Sandegren. 1992. Avskjutningsstatestik 1991 för björn i Sverige. Våra Rovdjur 9(2):24. (Bear harvest statistics for Sweden in 1991. In Swedish)

F19 Sandegren, F. & A. Söderberg. 1992. Brown bear. Swedish game; biology and management. The Swedish Association for Hunting and Wildlife Management, Spånga. Pp. 115-116.

F20 Sandegren, F. & J. Swenson. 1992. Björnjakten 1992--var den så dålig? Svensk Jakt 1992(12): 88-89. (The 1992 bear hunting season--was it so bad? In Swedish)

F21 Swenson, J., F. Sandegren, A. Söderberg & P. Wabbakken. 1992. Hur många björnar? Svensk Jakt 1992(10):37-42. (How many bears? In Swedish)

F22 Swenson, J., F. Sandegren, A. Söderberg & P. Wabbakken. 1992. Hur många björnar har vi i Sverige? Våra Rovdjur 9(3):6-12. (How many bears do we have in Sweden? In Swedish)

F23 Sandegren, F. & A. Söderberg.1993. Brunbjörn. Sverige, jakten och EG. Svenska jägareförbundet, Spånga. Pp. 148-149.

F24 Sandegren, F. & J. Swenson. 1993. Jo, björnräkningen stämmer visst! Svensk Jakt 1993(1):89-90. (Yes, the bear population estimate is correct! In Swedish)

F25 Swenson, J. & F. Sandegren. 1993. Björnen i Sverige. Svensk Jakt 1993(9):978-979. (The brown bear in Sweden. In Swedish)

F26 Söderberg, A., F. Sandegren, and J. Swenson. 1994. Hur stor var björnen?. Svensk Jakt 1994(8):832-834. (How large was the bear? In Swedish)

F27 Swenson, J. E., F. Sandegren, R. Franzén, S. Brunberg, K. A. Olander, P. Wabakken, A. Söderberg, and A. Bjärvall. 1994. Bjørnen, elgen og elgjakta. Elgen 1994:28-29. (Bears, moose, and moose hunting. In Norwegian)

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F28 Swenson, J., F. Sandegren, A. Söderberg, and P. Wabakken. 1994. Björnen i Skandinavien. Pp. 143-148 in G. Markgren, H. Andrén and H. Sand, eds. Skogsvilt II. Grimsö Wildlife Research Station, Swedish University of Agricultural Sciences, Riddarhyttan. (The brown bear in Scandinavia. In Swedish)

F29 Swenson, J. E. and F. Sandegren. 1995. Brunbjörnens genetik i Skandinavien. Svensk Jakt 1995(5):493-497. (The genetics of brown bears in Scandinavia. In Swedish)

F30 Söderberg, A. and J. E. Swenson. 1996. Björn på villovägar? ABC Jakt 10:6-7. (Lost bears? In Swedish)

F31 Sagør, J. T. and J. Swenson. 1996. Sau i bjørneland—kilde til konflikt! Pp. 8-23 in K. H. Brox (ed.). Natur 96/97. Tapir forlag, Trondheim, Norway. (Sheep in bear country—the source of conflict! In Norwegian, Invited chapter)

F32 Swenson, J., Sandegren, F., Wallin, K. & Cederlund, G. 1998. Karhun ja hirven yhteiselo Skandinaviassa. Riistantutkimuksen tiedote 149:3-4. (The coexistence of brown bears and moose in Scandinavia. In Finnish)

F33 Swenson, J. 1998. Forebyggende tiltak mot rovdyrskader. Pp. 374-379 in Husdyrforsøksmøtet 1998, Norwegian Agricultural University, Ås. 709 pp. (Mitigating measures against predator depredation. in Norwegian)

F34 Sandegren, F. and J.Swenson. 1998. Nya rön från det Skandinaviska Björnprojektet—projektet om den svenska björnen. Allt om djur och natur 1998(3):20-24. (New results from the Scandinavian Brown Bear Research Project—the project about the Swedish brown bear. in Swedish)

F35 Busk, H., B. Dahle, A. Jansson, T. Johansen, A. Norin, O. Opseth, I.-L. Persson, R. Riig, L. Stabell, A. Söderberg, J. Swenson, F. Sandegren, K. Wallin, and G. Cederlund. 1999. Bjørnen som predator på elg. Elgen 1999:12-15. (The brown bear as a predator on moose. in Norwegian).

F36 Söderberg, A., F. Sandegren & J. Swenson. 1999. En ovanlig vandringsväg. Våra Rovdjur 2: 8-9. (An unusual dispersal route. in Swedish)

F37 Sandegren, F. & J. Swenson. 2000. Björnen som älgjägare. Svensk Jakt 2000(2/3):62-65. (The brown bear as a moose hunter. in Swedish)

F38 Olson, J. E. 2000. Är björnen farlig? Björnprojektet i Orsa, Orsa. 24 pp. (Is the brown bear dangerous? In Swedish)

F39 Swenson, J. E. 2000. Rovdyr og rovdyrforvaltning. Pp. 325-329 in I. S. Kristiansen, chief editor. Store Norske Årbok 2000, Kunnskapsforlaget, Oslo, Norway. 543 pp. (Large predators and their management; in Norwegian)

F40 Swenson, Jon E. 2000. Rovdyr med konsekvenser. Verdens natur 15(2):8-11. (Large predators with consequences; in Norwegian)

F41 Brunberg, S. 2000. Om du möter en björn. Broschyr från Viltskadecenter. 8 sidorF42 Brunberg, S. 2000. If you encounter a bear . Pamphlet from Viltskadecenter. 8 ppF43 Brunberg, S. 2000.Wenn Sie einen Bären begegnen. from Viltskadecenter. 8 ppF44 Olson, J. E. 2001. Is the brown bear dangerous? Björnexperten i Orsa, Orsa,

Sweden. 24 pp. F45 Olson, J. E. 2001. Ist der Braunbär gefährlich? Björnexperten i Orsa, Orsa, Sweden.

24 pp. F46 Sandegren, F. & J. Swenson. 2001. Åteljakt på björn—bra eller dåligt? Svensk Jakt

2001 (2/3):64-65. (Bait hunting for brown bears—good or bad? in Swedish)F47 Hustad, H. & J. E. Swenson. 2001. Tildeling av skadefellingstillatelser på bjørn som

tar sau. Sau og Geit 2001(5):60-63. (Issuance of kill permits for bears that kill sheep; in Norwegian)

F48 Swenson, J. E. 2002. Nå våkner bjørnen! Villmarksliv 29(4):56-59. (Bears are waking now!; in Norwegian).

F49 Zedrosser, A. & B. Dahle. 2002. Brown bear attack in central Sweden. International Bear News 11/2:9.

F50 Olson, J. E. 2003. Er bjørnen farlig? Björnprojektet i Orsa, Orsa. 24 pp. (Is the brown bear dangerous? In Norwegian)

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F51 Olson, J. E. 2004. Lórso bruno é pericoloso? Björnprojektet i Orsa, Orsa. 24 pp. (Is the brown bear dangerous? In Italian)

F52 Katajisto, J.K. 2004. Näin pärjäät karhun naapurina, Tiede 3/2004 (Having bears as neighbour, Finnish science magazine, in Finnish).

F53 Swenson, J., J. Kindberg, E. Bellemain, S. Brunberg & G. Ericsson. 2004. Björnstammen är större än man trott. Våra Rovdjur 2004(3):14-15. (The population of brown bears is larger than previously thought; in Swedish).

F54 Swenson, J., J. Kindberg, E. Bellemain, S. Brunberg & G. Ericsson. 2005. Brunbjørn i Sverige 2004. Våre Rovdyr 19(1):14-15. (The brown bears in Sweden 2004; in Norwegian).

F55 Zedrosser, A. 2005. Mord, Intrigen und Sex. - Über die Partnerwahl bei Braunbären. Österreichs Weidwerk 6: 60-61. (Murder, intrigue, and sex—On mate choice in brown bears; in German)

F56 Zedrosser, A. 2005. Bäriges. Österreichs Weidwerk 8: 46-47. (About bears; in German)

F57 Zedrosser, A. 2007. Kindesmord zu Fortpflanzungszwecken. Wildbiologie 1/2007: 1-12. (Infanticide for the purpose of reproduction; in German)

F58 Sahlén, V., Swenson, J., Brunberg, S. og Kindberg, J. 2007. Bjørnen i Sverige. Våre Rovdyr 21(3): 80-94. (The brown bear in Sweden: in Norwegian)

F59 Arnemo, J. & S. Stokke. 2008. Myter og sannheter: hardskutte dyr. Jakt 2008 (8): 28-30.

F60 Stokke, S., J. Arnemo, M. Kristoffersson & A. Söderberg. 2008. Märkbart många björnar skadeskjuts. Våre Rovdjur 2008 (2): 18-20.

F61 Ericsson, G., Kindberg, J. & Swenson, J. 2008. Jägarna vet hur många björnar det finns! Svensk Jakt 2008: 920-921. (The hunters know how many bears there are!: in Swedish)

F62 Moen, G. K., O.-G. Støen, S. Brunberg & J. Swenson. 2009. Björnen går undan för skogsvandrare. Våra Rovdjur 2009 (3): 3-7. (Bears avoid people walking in the forest: in Swedish)

F63 Zedrosser, A. 2009. L’infanticide à des fin de reproduction. Chasse et nature 2: 32-34.F64 Moen, G. K., Støen, O.-G., Brunberg, S. og Swenson, J. 2010. Når mennesker møter

brunbjørn i skogen. Våre Rovdyr 24 (1): 10-11. (When people meet a brown bear in the forest: in Norwegian)

F65 Moen, G. K., Støen, O.-G., Brunberg, S. og Swenson, J. 2010. Når mennesker møter brunbjørn i skogen. Hjorteviltet 20: 84-85. (When people meet a brown bear in the forest: in Norwegian)

F66 Moen, G. K., Støen, O.-G., Brunberg, S. og Swenson, J. 2010. Når mennesker møter brunbjørn i skogen. Norsk Skogbruk 2010 (7/8): 32-33. (When people meet a brown bear in the forest: in Norwegian)

F67 Brunberg, S., O.-G. Støen & V. Sahlén. 2010. Capturing denning brown bears in Sweden. International Bear News 19(4):13.

F68 Swenson, J. E. 2010. Faktaark brunbjørn Ursus arctos truet art. Artsdatabankens faktaark ISSN1504-9140 nr. 101 (Fact sheet, brown bear, Ursus arctos, endangered species: in Norwegian).bankens faktaark

F69 Vang, S., A. Zedrosser, S. Brunberg & J. Swenson. 2011. Björnhundar behöver bättre träning. Svensk Jakt 2011: 185-188. (Bear dogs need better training: in Swedish).

F70 Vang, S., A. Zedrosser, S. Brunberg & J. Swenson. 2011. Ettersøksekvipasjene ikke gode nok. Jakt og Fiske 2011 (1-2): 64-68. (Dog-hunter tracking teams are not good enough; in Norwegian).

F71 Nilsen, E. B., V. Gervasi, J. D. C. Linnell, J. Odden, H. C. Pedersen, P. Wabakken, B. Zimmermann, & J. Swenson. 2011. Effekten av predasjon på hjortebestander. Hjorteviltet 21:91-93.

F72 Swenson, J. E. 2012. A new management policy for brown bears in Norway. International Bear News 21(2):21-22.

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F73 Støen, O.-G., P. Segerström, and J. Karlsson. 2012. Bjørnepredasjon på tamrein. Hjorteviltet 22:82-83.

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F76 Zedrosser, A., and Kindberg, J. 2013. Åtejakt på bjørn. Våre Rovdyr 27: 96-97.F77 Rønning, A. 2014. Bamsefar som er så snill og rar? Våre Rovdyr 28(1):22-25.F78 Elfstöm, M. & V. Sahlén. 2014. Björn och människa—inkräktare på varandras

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carnívoros? Quercus 341(Julio):14-21.F80 Gosselin, J. 2014. Hunting’s hidden impacts. Biosphere December 2014: 42-49.F81 Johansson, M., A. Flykt, J. Frank & Ole-Gunnar Støen. 2015. Redsel for bjørn og ulv

blant mennesker som bor i rovdyrområder. Rovdyrviten 3:12-13.F82 Sahlén, V. 2015. Når bjørn og menneske møtes. Rovdyrviten 3:14-15.F83 Støen, O.-G., P. Segerström, B. Åhman, L. T. Persson, & J. Frank. 2015. Bjørnens

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http://atlasofscience.org/brown-bear-habitat/F86 Steyaert, S. 2015. Infanticide bij beren uitgeplozen. Natuur.focus 2015 (maart): 34-35

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Year Sci. Books Other papers

PhD MS Rept Popular

1985 0 0 0 0 0 0 31986 0 0 0 0 0 0 01987 1 0 0 0 0 0 41988 0 0 0 0 1 2 11989 0 0 0 0 1 1 41990 2 1 0 0 1 3 21991 0 0 0 0 0 1 31992 0 0 0 0 1 3 51993 0 0 0 0 1 1 31994 1 0 1 0 0 5 31995 3 0 0 0 2 3 11996 2 0 1 0 2 12 21997 4 1 5 0 3 2 01998 6 0 2 0 4 7 31999 7 2 2 0 2 2 22000 5 2 3 0 6 2 72001 8 0 3 0 7 2 42002 1 1 1 0 4 1 22003 6 1 2 1 2 6 12004 6 0 0 3 4 1 32005 5 2 0 0 5 9 32006 12 0 0 3 3 6 02007 11 0 0 0 4 4 22008 11 0 0 3 4 6 32009 13 1 1 2 4 6 22010 8 0 0 2 3 8 52011 11 0 0 1 5 12 32012 20 0 0 1 6 9 22013 14 0 1 2 4 5 32014 22 0 0 0 3 4 42015 17 0 0 3 6 11 62016 5 0 0 0 0 0 0total 201 11 22 21 88 134 86in press 5

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