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A Field Guide to Ecosystem Identification for the Boreal White and Black Spruce Zone of British Columbia e Best Place on Earth Ministry of Forests and Range Forest Science Program LAND MANAGEMENT HANDBOOK 65 2011

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  • A Field Guide to Ecosystem Identifi cation for the Boreal White and Black Spruce Zone of British Columbia

    The Best Place on Earth

    Ministry of Forests and Range

    Forest Science Program

    LAND MANAGEMENT HANDBOOK

    65

    2011

  • A Field Guide to Ecosystem Identification for the Boreal White and Black Spruce Zone of British Columbia

    S. Craig DeLong Allen Banner William H. MacKenzie Bruce J. RogersBenita Kaytor

    The Best Place on Earth

    Ministry of Forests and Range

    Forest Science Program

  • ii

    Library and Archives Canada Cataloguing in Publication

    A field guide to ecosystem identification for the boreal white and black spruce zone of British Columbia /C. DeLong ... [et al.].

    Includes bibliographical references. ISBN 978-0-7726-6268-2

    . Ecological zones--British Columbia, Northern. 2. Ecological regions-- British Columbia, Northern. 3. Biotic communities--British Columbia, North-ern. I. DeLong, S. C. (Craig) II. British Columbia. Ministry of Forests and Range

    SD397 S77 F55 20 577.097 C200-90742-0

    Citation

    DeLong, C., A. Banner, W. H. MacKenzie, B. J. Rogers, and B. Kaytor. 20. A field guide to ecosystem identification for the Boreal White and Black Spruce Zone of British Columbia. B.C. Min. For. Range, For. Sci. Prog., Victoria, B.C. Land Manag. Handb. No. 65. www.for.gov.bc.ca/hfd/pubs/Docs/Lmh/Lmh65.htm

    Prepared forB.C. Ministry of Forests and RangeResearch Branch, Victoria, B.C.

    Copies of this report can be obtained from:Crown Publications, Queen’s Printer PO Box 9452 Stn Prov Govt, 563 Superior Street, 2nd Floor, Victoria, BC V8W 9V7 800 663-605 www.crownpub.bc.ca

    For more information on Forest Science Program publications, visit: www.for.gov.bc.ca/scripts/hfd/pubs/hfdcatalog/index.asp

    © 20 Province of British ColumbiaWhen using information from this or any Forest Science Program report, please cite fully and correctly.

    The use of trade, firm, or corporation names in this publication is for the information and convenience of the reader. Such use does not constitute an official endorsement or approval by the Government of British Columbia of any product or service to the exclusion of any others that may also be suitable. Contents of this report are presented as information only. Funding assistance does not imply endorsement of any statements or information contained herein by the Government of British Columbia. Uniform Resource Locators (URLs), addresses, and contact information contained in this document are current at the time of printing unless otherwise noted.

  • ACKNOWLEDGEMENTS

    The classification and interpretations presented in this field guide have evolved over the past 30 years through the collective work of several individuals involved in the many phases of the provincial ecological classification project.

    Preliminary ecological classifications, description, and interpretation of units, and biogeoclimatic boundary refinement were developed in part by R. Annas, L. Jang, A. MacKinnon, D. Meidinger, J. Pojar, and E. Wright. In addition to these individuals, many others collected field data that form the basis of the classification presented in this guide. J. Pojar, G. Argus, F. Boas, T. Goward, and W. Noble identified difficult vascular plant, bryophyte, and lichen specimens.

    The format and content of this guide have benefited from guides from other forest regions in British Columbia and some of the aids to site and soil description have been adapted from these guides. V. Foord contrib-uted to the Climate Section of the guide.

    Review comments were provided by R. Kabzems, D. MacKillop, and D. Meidinger. Thanks to P. Nystedt and R. Scharf for co-ordinating layout, graphics, and publication.

    Plant illustrations are reprinted by permission of the University of Washington Press, Flora of the Pacific Northwest (Hitchcock and Cron-quist 973); of W. Schofield, for P. Drukker-Brammall’s moss illustra-tions from Some Common Mosses of British Columbia (Schofield 992); and of T. Goward for use of lichen illustrations from The Lichens of Brit-ish Columbia (Goward 999). The British Columbia Ministry of Forests and Range Northern Interior Region provided funds for preparation and publication of this guide.

    iii

  • CONTENTS

    Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . iii Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

    . Objectives/Scope . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .2 Regulatory and Standards Context . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3.3 Other Sources of Information . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4.4 Guide Content . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4.5 Training Courses . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5

    2 Biogeoclimatic Ecosystem Classification (BEC) System . . . . . . . . . . . 52. An Overview of the BEC Classification System . . . . . . . . . . . . . . . . . 52.2 Zonal (Climatic) Classification . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 62.3 Site Classification . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 82.4 Unit Numbering Convention . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9

    3 The BWBS Zone in British Columbia . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3. Vegetation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3.2 Disturbance . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 43.3 Physiography and Soils . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 53.4 Climate of the Boreal Forest . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 24

    4 Overview of Biogeoclimatic Units of the BWBS Zone . . . . . . . . . . . . . 324. BWBSdk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 324.2 BWBSmk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 364.3 BWBSmw . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 374.4 Murray BWBSwk (BWBSwk) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 384.5 Graham BWBSwk (BWBSwk2) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 384.6 Kledo BWBSwk (BWBSwk3) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 394.7 BWBSvk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 39

    5 How To Identify an Ecosystem . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 405. Field Procedures . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 405.2 Layout of Biogeoclimatic (BGC) Unit Sections . . . . . . . . . . . . . . . . . . 62

    6 Ecosystem Units of the BWBS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67 6. Dry Cool Boreal White and Black Spruce (BWBSdk) . . . . . . . . . . 67

    BWBSdk/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 75BWBSdk/02 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76BWBSdk/03 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77BWBSdk/04 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78BWBSdk/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79

    v

  • vi

    BWBSdk/ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80BWBSdk/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86BWBSdk/026B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 87BWBSdk/036B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88BWBSdk/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89BWBSdk/2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90

    6.2 Moist Cool Boreal White and Black Spruce (BWBSmk) . . . . . . . 9BWBSmk/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 99BWBSmk/02 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 00BWBSmk/03 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 0BWBSmk/04 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 02BWBSmk/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 03BWBSmk/ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 04BWBSmk/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 0BWBSmk/06B.2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . BWBSmk/026B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2BWBSmk/036B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3BWBSmk/046B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4BWBSmk/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5BWBSmk/2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6

    6.3 Moist Warm Boreal White and Black Spruce (BWBSmw) . . . . . . 7BWBSmw/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25BWBSmw/02 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26BWBSmw/03 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27BWBSmw/04 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28BWBSmw/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29BWBSmw/ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30BWBSmw/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36BWBSmw/026B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37BWBSmw/036B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38BWBSmw/046B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 39BWBSmw/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 40BWBSmw/6B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4BWBSmw/6B.2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42BWBSmw/2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 43

    6.4 Murray Wet Cool Boreal White and Black Spruce (BWBSwk) . 45BWBSwk/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 53BWBSwk/02 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 54BWBSwk/03 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55

  • BWBSwk/04 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56BWBSwk/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 57BWBSwk/ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 58BWBSwk/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 64BWBSwk/026B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 65BWBSwk/036B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 66BWBSwk/046B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67BWBSwk/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68BWBSwk/06B.2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69

    6.5 Graham Wet Cool Boreal White and Black Spruce (BWBSwk2) 7BWBSwk2/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79BWBSwk2/02 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80BWBSwk2/03 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8BWBSwk2/04 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 82BWBSwk2/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83BWBSwk2/ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84BWBSwk2/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90BWBSwk2/026B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9BWBSwk2/036B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92BWBSwk2/046B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93BWBSwk2/06B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94BWBSwk2/6B. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 95

    6.6 Kledo Wet Cool Boreal White and Black Spruce (BWBSwk3) . . 97BWBSwk3/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 204BWBSwk3/02 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 205BWBSwk3/03 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206BWBSwk3/0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 207BWBSwk3/ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 208

    7 Wetlands and Non-forested Ecosystems of the BWBS Zone . . . . . . . . 2097. Site Classes in the BWBS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2097.2 Common Wetland and Non-forested Site Associations of

    the BWBS Zone . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 288 Literature Cited . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 239

    APPENDIX Table for conversion of old biogeoclimatic unit and site unit numbers

    and names to new BGC unit and site unit nubers and names . . . . . . . . . 244

    vii

  • viii

    tables2. System of naming and coding interior biogeoclimatic units . . . . . . . . . . 73. Environmental characteristics of the BWBS zone and adjacent

    zones in northern British Columbia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23.2 Comparison of climate variables for the BWBS zone and adjacent

    portions of zones in northern British Columbia . . . . . . . . . . . . . . . . . . . . 263.3 BWBS 97–2000 climate normals: Environment Canada weather

    stations with World Meteorological Organization Standards . . . . . . . . . 283.4 Summary of climate data for biogeoclimatic units within the

    BWBS zone of northern British Columbia . . . . . . . . . . . . . . . . . . . . . . . . . 294. Environmental characteristics of biogeoclimatic units within the

    BWBS zone of northern British Columbia . . . . . . . . . . . . . . . . . . . . . . . . . 335. General steps for site location and description . . . . . . . . . . . . . . . . . . . . . 435.2 Section and page numbers for biogeoclimatic unit sections . . . . . . . . . 485.3 Site and soil factors to be collected . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 495.4 Identification of upland humus forms . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55.5 Hand texturing guide . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 525.6 Properties of soil separates . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 525.7 Choice of procedures for mature and climax stands versus recently

    disturbed/early seral sites . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 565.8 Definitions of terms used in the keys to identification of relative

    soil moisture and nutrient regimes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 575.9 Table for estimation of soil nutrient regime . . . . . . . . . . . . . . . . . . . . . . . 65.0 Rules for identifying species values in the Characteristic Vegetation

    section of the site/seral unit summary page . . . . . . . . . . . . . . . . . . . . . . . . 667. Distribution and abundance of wetland, riparian, grassland, and

    other non-forested site classes in subzones of the BWBS . . . . . . . . . . . . 27.2 Wetland Site Associations occurrence by BGC unit . . . . . . . . . . . . . . . . . 297.3 Non-wetland Site Associations occurrence by biogeoclimatic unit . . . . 235

  • figures. Distribution of biogeoclimatic units of the Boreal White and

    Black Spruce zone in British Columbia . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22. Hierarchical relationship between climatic-level and

    site-level classifications. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83. Vegetation table comparing zonal sites for BWBS zone and

    adjacent zones. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73.2 Physiographic units and subunits of northern British Columbia . . . . . . 85. A flowchart outlining the procedure for identifying site/seral

    units as described in Sections 5..–5..3 . . . . . . . . . . . . . . . . . . . . . . . . . . . 45.2 Key to differentiation of biogeoclimatic units of the BWBS . . . . . . . . . . 455.3 Vegetation table for comparison of vegetation on zonal sites of

    the biogeoclimatic units of the BWBS . . . . . . . . . . . . . . . . . . . . . . . . . . . . 475.4 Slope position diagram . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 505.5 Soil texturing key . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 545.6 A key to the identification of relative soil moisture regimes . . . . . . . . . . 595.7 A key to the identification of relative soil nutrient regime . . . . . . . . . . . 606. Distribution of the BWBSdk within British Columbia . . . . . . . . . . . . . . 696.2 Distribution of the BWBSmk within British Columbia . . . . . . . . . . . . . . 936.3 Distribution of the BWBSmw within British Columbia . . . . . . . . . . . . . 96.4 Distribution of the BWBSwk within British Columbia . . . . . . . . . . . . . 476.5 Distribution of the BWBSwk2 within British Columbia . . . . . . . . . . . . . 736.6 Distribution of the BWBSwk3 within British Columbia . . . . . . . . . . . . . 987. Vegetation table of Bog and Coniferous Swamp Site Associations . . . . . 2237.2 Vegetation table of Fen, Marsh, and Low Shrub Bog Site Associations . 2277.3 Vegetation table of Flood and Tall Shrub Swamp Site Associations . . . . 2337.4 Vegetation table of Rock Outcrop, Grassland, and Brushland

    Site Associations . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 238

    ix

  • 1 INTRODUCTION

    1.1 Objectives/Scope

    This guide presents site identification information for ecosystems of the Boreal White and Black Spruce (BWBS) biogeoclimatic zone of British Columbia (Figure .).

    The guide has four main goals:• to present descriptions of British Columbia’s boreal ecosystems in a

    single guide;• to present a comprehensive revision to the classification of the BWBS;• to assist the user in classifying sites in the field; and• to provide insight into common site limitations and potential capa-

    bilities related to the ecological conditions of the classified units to assist in preparing management prescriptions.

    This guide represents a revision of existing classifications based on additions of new data and an inter-regional correlation of BWBS units. This guide replaces Land Management Handbook (LMH) 22 A Field Guide for Identification and Interpretation of Ecosystems of the North-east Portion of the Prince George Forest Region (DeLong et al. 990); the boreal sections of LMH 26, A Field Guide to Site Identification and Interpretation for the Prince Rupert Forest Region (Banner et al. 993); the BWBSdk section in LMH 54, A Field Guide to Site Identification and Interpretation for the North Central Portion of the Northern Interior Forest Region (DeLong 2004); and the Draft Field Guide Insert for Site Identification of Seral Populus Units of the Northeast Portion of the Prince George Forest Region.1

    The main changes arising from revisions are:• merged BWBSdk2 and BWBSmw2 into BWBSmk because vegetation

    similarities and climate data indicate that these regions are closely affiliated;

    • renamed BWBSdk as BWBSdk and BWBSmw as BWBSmw, as there is now only one variant of each subzone;

    • expanded classification of BWBSwk3 based on additional field sampling;

    British Columbia Ministry of Forests. 2002. Draft field guide insert of seral Populus units of the northeast portion of the Prince George Forest Region. Victoria, B.C. Field Guide Insert. Unpubl. draft.

  • BWBS Zone Field Guide2

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  • BWBS Zone Field Guide 3

    • major plot additions to BWBSdk and BWBSmw based on extensive data compilation, (historical and recent collections);

    • formal classification of mature broad-leaved ecosystems; and• revised system of numbering units based on province-wide correla-

    tion.

    See Appendix to determine equivalent previous biogeoclimatic and site units for new units described in this guide.

    1.2 Regulatory and Standards Context

    Section 6 of the Forest Planning and Practices Regulation (FPPR) (Province of British Columbia 2004a), requires forest stewardship plans (FSPs) to specify stocking standards for areas harvested under the plan. One of the factors that a person who prepares this plan should consider is that the stocking standards contain “the types of commercially valu-able and ecologically suitable species that should be established on the area” (Schedule (6) of the FPPR) (Province of British Columbia 2000, 2004b). To determine if a species is ecologically suitable for an area, the ecological characteristics of that area need to be determined. The gener-ally accepted method and current “best practice” of assessing these characteristics is biogeoclimatic site series mapping.

    The Forest and Range Practices Act (FRPA) requires that a site plan be prepared for areas that will be harvested (Province of British Columbia 2002). This site plan must identify the standards units for the area and the stocking standards that apply to those units (Section 34 of FPPR) (Province of British Columbia 2004c). The surveys guidebook states that ecosystem classification plays an important role in developing stan-dards units (Province of British Columbia 2000). In addition, stocking standards, which are a required component of an FSP, are commonly defined by the biogeoclimatic and site units over which they apply.

    Biogeoclimatic Ecosystem Classification (BEC) is also a standard that forms part of the many resource inventory procedures published by the provincial Resource Inventory Standards Committee (RISC), including standards for terrestrial ecosystem mapping and predictive ecosystem mapping and standards for mapping ecosystems at risk (www.ilmb.gov.bc.ca/risc/pubs/teecolo/index.htm). These procedures must be used for most inventory and environmental assessment projects in British Columbia.

    http://www.ilmb.gov.bc.ca/risc/pubs/teecolo/index.htmhttp://www.ilmb.gov.bc.ca/risc/pubs/teecolo/index.htm

  • BWBS Zone Field Guide4

    1.3 Other Sources of Information

    Numerous reports on vegetation, soils, wildlife, and ecosystem descrip-tion and classification exist for the BWBS of British Columbia and adjoining area. Some of these references can be found in Section 8 (Literature Cited).

    Information at the biogeoclimatic zone level is available in Ecosystems of British Columbia (Meidinger and Pojar [editors] 99). The Wetlands of British Columbia: A Guide to Identification (MacKenzie and Moran 2004) provides more information and tools for identification of the wetlands described in this guide.

    The Field Manual for Describing Terrestrial Ecosystems (B.C. Ministry of Environment, Lands and Parks and B.C. Ministry of Forests 998) provides tools and useful information for assisting with collecting field data related to describing the ecosystems covered by this guide. Excel-lent references for plant identification, which complement this guide, are Plants of Northern British Columbia (MacKinnon et al. [editors] 9922) and Plants of the Western Boreal Forest and Aspen Parkland (Johnson et al. 995). When conducting a complete plant inventory Illustrated Flora of British Columbia should be used (Douglas et al. 998a, 998b, 999a, 999b, 2000, 200a, 200b, 2002). E-Flora BC (www.eflora.bc.ca/), which provides descriptions and colour images for most of British Columbia’s flora, is also a useful plant identification resource.

    Additional information relating to BEC, including the most recent biogeoclimatic mapping, links to BEC field guides, and background information on the BEC system, is on the provincial Ecology program website at: www.for.gov.bc.ca/hre/becweb/.

    1.4 Guide Content

    This guide consists of eight sections. Following the introduction is a brief discussion of the classification system (Section 2). Sections 3 and 4 provide an overview of the BWBS in British Columbia, including vegetation, physiography, climate, and wildlife, as well as descriptions of climatic units. Section 5 contains procedures for site description and identification. Section 6 contains detailed information to assist with ecosystem identification within each of the BWBS subzones/variants

    2 Revised second edition published in 999.

    http://www.eflora.bc.ca/http://www.eflora.bc.ca/http://www.for.gov.bc.ca/hre/becweb/

  • BWBS Zone Field Guide 5

    found within the guide area, tools for identification of climatic and ecosystem units, descriptions of coniferous and broad-leaved ecosys-tem units, and comments relating to identification and management. Section 7 describes common non-forested ecosystems of the BWBS. Applicable literature is cited in Section 8.

    The classification is based on 590 plots located in the BWBS within the Northern Interior Forest Region. These plots are generally well distrib-uted geographically (proportional to the size of the biogeoclimatic unit) except in areas with difficult access. Most ecosystem units are charac-terized by at least five plots, although certain less common ecosystems (e.g., on very dry or wet sites) may be characterized by fewer plots.

    1.5 Training Courses

    It is assumed that the user of this guide is familiar with the basic concepts and methods of site, soil, and vegetation evaluation and has completed the training programs offered by the Forest Service or educational institutions. These courses are offered annually in various locations within the Forest Region. For information about such training courses, please contact the Forest Research Section of the Northern Interior Forest Region, or the University of Northern British Columbia Continuing Studies Network, or check the following link www.for.gov.bc.ca/hre/becweb/resources/training/index.html.

    2 BIOGEOCLIMATIC ECOSYSTEM CLASSIFICATION (BEC) SYSTEM

    This section briefly describes the BEC system. For a more complete description refer to Ecosystems of British Columbia (Meidinger and Pojar [editors] 99) or Biogeoclimatic Ecosystem Classification in British Columbia (Pojar et al. 987).

    2.1 An Overview of the BEC Classification System

    The BEC system is a hierarchical classification scheme that combines three classifications: climatic (or zonal), vegetation, and site. For practi-cal purposes, users need only be concerned with the zonal and site clas-sifications (Figure 2.); the vegetation classification is used for ensuring consistency in naming conventions and for correlating ecosystems

    http://www.for.gov.bc.ca/hre/becweb/resources/training/index.htmlhttp://www.for.gov.bc.ca/hre/becweb/resources/training/index.html

  • BWBS Zone Field Guide6

    provincially, nationally, and internationally. The information presented in this guide will allow the user to apply BEC in the field. BEC principles evolved from the work of V.J. Krajina (965, 969) and are described in this section. The classification system was adapted for the province by the B.C. Ministry of Forests in the 970s (Pojar et al. 987). The objectives of the BEC system are: • to provide a framework for organizing ecological information and

    management experience about ecosystems; • to promote further understanding of identified ecosystems and the

    relationships among them; • to supply resource managers with a common language to describe

    forest sites; and • to improve the user’s ability to prescribe and monitor treatment

    regimes on a site-specific (ecosystem) basis.

    2.2 Zonal (Climatic) Classification

    Biogeoclimatic units are groups of ecosystems that have developed under the influence of the same regional climate. In the hierarchy of climatic units, the biogeoclimatic subzone is the basic unit. Subzones are grouped into zones, and divided into variants. Each biogeoclimatic subzone has a distinct climax (or near-climax) plant association on zonal sites. Zonal sites are defined as being those located in midslope positions, with deep, broadly loamy soils and mesic moisture regimes. The climax vegetation on zonal sites is considered to best reflect the regional climatic conditions of the subzone.

    Each subzone has a distinctive sequence of related ecosystems rang-ing from dry to wet sites. Sites that are wetter or drier than the zonal ecosystem are influenced not only by the regional climate, but also by soil and topographic features at local scales. For example, in the Moist Warm subzone of the Boreal White and Black Spruce zone (BWBSmw), zonal sites are dominated by a white spruce canopy with a diverse, moderately well-developed understorey of shrubs and herbs; dry sites are dominated by a lodgepole pine canopy with an understorey dominated by a limited number of shrub species, grasses, and a mix of lichens and mosses; and wet sites in the same subzone (climate) have a white spruce overstorey with an understorey dominated by a diverse, well-developed shrub layer and a wide variety of herbs.

    The biogeoclimatic variant was defined because subzones contain

  • BWBS Zone Field Guide 7

    considerable geographic variation. Variants reflect further differences in regional climate and are generally recognized for areas that are slightly drier, wetter, snowier, warmer, or colder than other areas in the subzone. For example, the Graham Wet Cool variant (BWBSwk2) of the BWBS is cooler than the Murray Wet Cool variant (BWBSwk) of the BWBS. These climatic differences result in corresponding differ-ences in vegetation, soil, and ecosystem productivity. The differences in vegetation are evident as distinct zonal climax or near-climax plant subassociations.

    Subzones with similar climatic characteristics and zonal ecosystems are grouped into biogeoclimatic zones. A zone is a large geographic area with a broadly similar type of climate. A zone has typical patterns of vegetation and associated similarities in nutrient cycling and soil climate. Zones also have one or more typical zonal climax species of tree, shrub, herb, or moss.

    Zones are usually named after one or more of the dominant climax spe-cies in zonal ecosystems and a geographic or climatic modifier (e.g., Bo-real White and Black Spruce zone). Zones are given a two- to four-letter code that corresponds to the name. For example, the Boreal White and Black Spruce zone is BWBS.

    Subzone names are derived from classes of relative precipitation and temperature. Subzone codes correspond to the climatic modifiers (Table 2.). For example, the BWBSmw refers to the Moist Warm (mw) subzone of the BWBS zone. Variants are named by geographic area and numbered from south to north or, if the most southerly extent is similar, from west to east. Hence, the BWBSwk variant extends farther south than the BWBSwk2 variant.

    Table 2. System of naming and coding interior biogeoclimatic units

    ZONEpt

    p = precipitation regime t = temperature regime x = very dry h = hot d = dry w = warm m = moist m = mild w = wet k = cool v = very wet c = cold v = very cold

  • BWBS Zone Field Guide8

    2.3 Site Classification

    Site series are the most commonly used units of site classification (Figure 2.) and are characterized within a biogeoclimatic unit. They are defined by using late seral or climax vegetation and represent site units having similar environmental properties and potential vegetation. The potential vegetation and selected environmental properties are used in this guide to characterize site series.

    FIGURE 2. Hierarchical relationship between climatic-level (zonal) and site-level classifications. Updated from Pojar et al. (1987).

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  • BWBS Zone Field Guide 9

    Each biogeoclimatic unit has a characteristic sequence of site series ac-cording to relative soil moisture regime (RSMR) and, to a lesser degree, relative soil nutrient regime (RSNR)3 along with other environmental drivers such as flooding regime and cold air ponding. RSMR is a relative scale of “available water” for plant growth within the climate of the biogeoclimatic unit. An eight-class scale is used ranging from 0 or very xeric (bare rock) to 7 or subhydric (water tables at or near the surface year-round). RSNR is a relative scale of “available nutrients” for plant growth. A five-class scale ranging from A (very poor) to E (very rich) is used.

    English names of one to four species are used to name site series, and tree species codes are usually substituted to shorten the name (e.g., SwPl – Soopolallie – Wildrye site series).

    Similar plant communities can occur in different biogeoclimatic units, but the RSMR and RSNR that they represent may differ between subzones. These communities belong to the same grouping of site series that is collectively called a site association,4 such as:

    BWBSwk1/Sb – Huckleberry – Lingonberry site series = BWBSwk1/104 BWBSwk3/Sb – Huckleberry – Lingonberry site series = BWBSwk3/103

    Both of these site series belong to the same site association, so their climax vegetation is similar, but their occurrence in the landscape, site conditions, and seral vegetation patterns differ between the two biogeo-climatic units.

    2.4 Unit Numbering Convention

    Historically, each forested site series was given a two-digit numeric code with the “0” reserved for the zonal ecosystem and then other units numbered sequentially from driest (02) to wettest. With new revisions ongoing in many parts of the province, a new three-digit system has been introduced to minimize confusion arising from coding reclassified units. In the three-digit system, the first number indicates

    3 The Site Identification section (Section 5.4) contains relative soil moisture and relative soil nutrient regime identification information.

    4 Site associations are not clearly recognized in the classification presented in this manual, except through the site series names. They are defined in Pojar et al. (987).

  • BWBS Zone Field Guide0

    the revision version (e.g., in this guide) and the second two numbers designate the site series with the “0” reserved for the zonal, 02–09 for units drier than zonal (ordered from driest to wettest, poorest to richest) and units 0 and greater applied to units wetter than zonal following the same numbering order. Units that are the same moisture regime as the zonal but poorer than zonal are coded as drier units. If the range in RSMR of a site series covers more than one class, the driest moisture regime is used for assignment. If future revisions are under-taken, all units will begin with a 2 (e.g., the zonal site will be coded as 20). Further label modifiers are: • BGC Phase: variability in mapped BEC variant (letter code);• Site Series Phase: same plant community, different site conditions; • Site Series Variation: different plant community, same site conditions

    (see LMH 25, Field Manual for Describing Terrestrial Ecosystems [B.C. Ministry of Environment, Lands and Parks, and B.C. Ministry of Forests 998]).

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    Seral ecosystems are numbered according to the site series with which the unit is affiliated, but are followed by a “” to designate that the unit is seral. The seral unit may then be further identified by the structural stage that the unit typically represents (see LMH 25, Field Manual for Describing Terrestrial Ecosystems [B.C. Ministry of Environment, Lands and Parks and B.C. Ministry of Forests 998] for structural stage descriptions); a stand composition modifier (B = broad-leaved, C = conifer, M = mixed); and a running number that is used to designate the seral vegetation community (seral plant association = seral varia-tion). Any one site series may have a number of seral variations within a structural stage/stand composition combination. The one that most commonly occurs will be seral variation , followed by the next most common, which would be seral variation 2, etc.

  • BWBS Zone Field Guide

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    Appendix contains a table to convert units from the old system to the new system.

    3 THE BWBS ZONE IN BRITISH COLUMBIA

    The BWBS zone in British Columbia covers about 6 million ha and extends from the lee slopes of the Coast Mountains in the west to the Alberta border in the east, and from just south of 54° N latitude in the south to the Northwest Territories border in the north (Figure .). The BWBS is part of the circumpolar boreal zone, one of the world’s major ecological zones, covering 29% of the North American Continent, north of Mexico (Brandt 2009). Table 3. compares environmental charac-teristics for selected subzones of the biogeoclimatic zones that share a common border with the BWBS.

    3.1 Vegetation

    The BWBS zone is a mixture of two main ecosystems: upland forests and muskeg. Upland forests are most common in better-drained por-tions of the Alberta Plateau in the east and in mountainous parts of the zone in the west. Muskeg is most extensive on the poorly drained northeastern lowlands.

    The BWBS contains several different upland forest types. The most common types are pure stands of trembling aspen, common in the southeast; mixed stands of trembling aspen, common in the northeast;

  • BWBS Zone Field Guide2

    Tabl

    e 3.

    En

    viro

    nmen

    tal c

    hara

    cter

    istic

    s of t

    he B

    WBS

    zon

    e an

    d ad

    jace

    nt z

    ones

    in n

    orth

    ern

    Briti

    sh C

    olum

    bia

    Zo

    neC

    hara

    cter

    istic

    BW

    BS

    SWB

    SBS

    ESSF

    IC

    H

    Biog

    eocl

    imat

    ic

    All

    All

    mc2

    , mk1

    , mk2

    , m

    c, m

    v1, m

    v2, m

    v3,

    mc2

    , wc,

    vc

    units

    con

    side

    red

    w

    k2, w

    k3

    mv4

    , wv

    for c

    ompa

    riso

    n

    Dis

    trib

    utio

    n

    rela

    tive

    to B

    WBS

    Occ

    urs a

    bove

    the

    Abu

    ts th

    e BW

    BS a

    t its

    O

    ccur

    s abo

    ve th

    e

    Abu

    ts th

    e BW

    BSdk

    BW

    BS o

    ver m

    ost

    sout

    hern

    ext

    ent i

    n BW

    BS in

    the

    at

    its s

    outh

    wes

    tern

    of it

    s ext

    ent n

    orth

    th

    e Ro

    cky

    Mou

    ntai

    ns

    sout

    hern

    par

    t of

    exte

    nt a

    long

    the

    of a

    ppro

    xim

    atel

    y

    and

    Foot

    hills

    , Roc

    ky

    its e

    xten

    t, so

    uth

    Isku

    t Riv

    er

    56

    ° 30'

    M

    ount

    ain

    Tren

    ch, a

    nd

    of a

    ppro

    xim

    atel

    y

    Om

    inec

    a M

    ount

    ains

    , 56

    ° 30'

    and

    at it

    s wes

    tern

    ex

    tent

    in th

    e St

    ikin

    e

    and

    Yuko

    n Pl

    atea

    usEl

    evat

    ion

    25

    0–12

    00

    900–

    1600

    10

    0–95

    0 80

    0–15

    00

    150–

    1000

    rang

    e (m

    )

    Terr

    ain

    Mou

    ntai

    n an

    d

    Gen

    tly ro

    lling

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    wer

    slop

    es a

    nd

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    tle to

    stee

    p,

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    er sl

    opes

    and

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    atea

    u va

    lleys

    pl

    atea

    us, w

    ide

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    s of m

    ajor

    ro

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    n flo

    ors o

    f maj

    or

    in th

    e w

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    ys to

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    avily

    gla

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    ains

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    st

  • BWBS Zone Field Guide 3

    Zo

    neC

    hara

    cter

    istic

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    BS

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    IC

    H

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    ate

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    ther

    n co

    nti-

    Con

    tinen

    tal

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    cont

    inen

    tal,

    Sub-

    cont

    inen

    tal

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    subc

    ontin

    en-

    ne

    ntal

    with

    su

    balp

    ine

    with

    w

    ith in

    crea

    sing

    suba

    lpin

    e; lo

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    tal,

    mod

    erat

    ed b

    y

    freq

    uent

    arc

    tic

    freq

    uent

    arc

    tic

    mar

    itim

    e in

    fluen

    ce

    cold

    , sno

    wy

    stro

    ng c

    oast

    al

    air m

    asse

    s; lo

    ng,

    air m

    asse

    s; lo

    ng

    tow

    ard

    the

    coas

    t;

    win

    ters

    and

    shor

    t, in

    fluen

    ces;

    mild

    ,

    very

    col

    d w

    inte

    rs

    very

    col

    d w

    inte

    rs

    moi

    ster

    sum

    mer

    s co

    ol, r

    athe

    r wet

    m

    oist

    sum

    mer

    s

    and

    shor

    t, dr

    y,

    and

    very

    shor

    t, an

    d m

    ilder

    win

    ters

    su

    mm

    ers

    and

    cold

    , sno

    wy

    qu

    ite w

    arm

    co

    ol su

    mm

    ers

    than

    BW

    BS

    w

    inte

    rs

    sum

    mer

    s

    Tree

    spec

    ies

    Maj

    or c

    onife

    rous

    Sw

    , Pl,

    Bl, S

    ba

    Sw, S

    b, P

    l, Bl

    Sx

    w, P

    l, Sb

    , Bl

    Se, B

    l, Pl

    , Sb,

    H

    w, B

    l, Sx

    w, P

    l t

    rees

    (Hm

    , Hw

    ) M

    ajor

    bro

    ad-

    At,

    Acb

    or A

    ct,

    Will

    ows,

    A

    t, Ep

    , Act

    At,

    Act

    , Ep

    lea

    ved

    tree

    s Ep

    A

    t, A

    cb

    a Tr

    ee sp

    ecie

    s cod

    es: A

    cb =

    bal

    sam

    pop

    lar,

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    t = tr

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    mar

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  • BWBS Zone Field Guide4

    and mixed stands of lodgepole pine and white spruce, common in the western and montane areas of the zone. Open lodgepole pine and lichen forests occupy drier sites, whereas poorly drained wet sites give rise to communities of black spruce and moss. White spruce and balsam pop-lar flourish on rich, well-drained river floodplains, which contain the most productive forests of the zone. Other tree species that have more limited ranges within the guide area are discussed in the introductory comments for each biogeoclimatic unit.

    The term “muskeg” describes the peatland combination of bogs and nutrient-poor fens that cover extensive parts of northeast British Columbia. The most common trees in this ecosystem are stunted black spruce and tamarack. Muskeg occurs over deep layers of peat where permafrost is often found.

    The BWBS zone contains several other minor but important ecosys-tems: in the northeast, nutrient-rich swamps and fens are dominated by tamarack, sedges, and brown mosses; boreal grassland and scrub communities occupy steep, south-facing slopes; and small but produc-tive marsh and shallow lake ecosystems are found throughout the zone. Figure 3. compares characteristic vegetation of the BWBS with that of selected areas of adjacent zones (listed in environmental summary – Table 3.).

    3.2 Disturbance

    Fire is the key stand-replacing disturbance agent operating in the BWBS zone, with the exception of flooding along broad fluvial terraces adjacent to larger rivers. The disturbance rate from fire is generally esti-mated to be about % of the total forested area per year (i.e., fire cycle of 00 years), but will vary in particular locations depending on climatic and topographic factors. Historically, large wildfires (> 000 ha) domi-nated the landscape, and upland sites were regenerated quickly by dense trembling aspen, trembling aspen and spruce, or lodgepole pine, result-ing in large patches of relatively even-aged forests. On wetlands, black spruce, tamarack, Alaska paper birch, and occasionally white spruce regenerate after fire. Initially, wetland stands tend to be very open, filling in over time, except in upland transition areas where stands are often denser. In small areas where past fires were intense, stands may regenerate to willow or alder.

  • BWBS Zone Field Guide 5

    During stand development, increasing amounts of white spruce and black spruce will occur in many stands originally dominated by trem-bling aspen or lodgepole pine. This increase occurs more rapidly and these species become a more dominant portion of the canopy on wetter sites. Post-fire stands are very dense except on the wettest sites, and self thin over time.

    Tomentosus root disease is thought to be a key disturbance agent af-fecting white spruce and in some localized areas may cause conversion from spruce-dominated stands to aspen-dominated stands over the course of 20–40 years. Eastern spruce budworm may also cause signifi-cant mortality of mature or immature spruce and lead to conversion of mixed to almost pure aspen stands.

    Floodplains are often extensive along major rivers in the BWBS. Seasonal inundation is common in these areas, but stand replacement flood cycles are unknown. Where stand replacement occurs, the typical floodplain succession pattern shifts from willow to cottonwood on the lower benches and from cottonwood to spruce on the higher benches.

    Stand ages rarely exceed 200 years. Relatively large patches (> 00 ha) of older forest (40–80 years) that were found scattered across the landscape have been reduced by harvesting over the past 40–50 years. Although patches of old forest (> 40 years) likely always occurred in the landscape, their position would have moved around over time due to regular disturbances.

    3.3 Physiography and Soils

    The distribution and extent of boreal forests in northern British Co-lumbia is to a large degree controlled by physiography. The BWBS zone is a lowland to montane zone found at elevations below 200 m; it is extensive on the lowlands and subdued terrain of the Alberta Plateau in the northeast corner of the province but becomes a valley-bottom phenomenon in the more mountainous western extent.

    Holland (976) divides northern British Columbia into 3 physiographic units; most of these are further divided into subunits (Figure 3.2). This framework will be used to describe the diverse physiography of north-ern British Columbia from east to west. Further information relevant to the physiography and soils of British Columbia’s boreal forest can be

  • BWBS Zone Field Guide6

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  • BWBS Zone Field Guide 7

    FIGU

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  • BWBS Zone Field Guide8

    found in Annas (997), Valentine et al. (editors, 978), Pojar et al. (983), Meidinger and Lewis (983), and Meidinger et al. (986).

    3.3. Alberta Plateau

    The Alberta Plateau is an extensive area of low relief resulting from the erosion of sedimentary sandstones and shales. This large plateau lies mostly between 900 and 200 m in elevation but is incised by the Liard and Peace Rivers and their tributaries to elevations as low as 300 m.

    The Alberta Plateau represents the most extensive area of the BWBS zone in British Columbia. The BWBSmk and BWBSmw subzones domi-nate the plateau area with the BWBSwk occurring at elevations above 950–050 m in the west near the Rocky Mountain Foothills. Upland trembling aspen or mixed trembling aspen–white spruce forests domi-nate the southern Alberta Plateau, while the Fort Nelson Lowlands por-tion of the plateau that lies mostly below 600 m in the northeast corner of the province is dominated by poorly drained black spruce muskeg.

    FIGURE 3.2 Physiographic units and subunits of northern British Columbia (after Holland 1976).

  • BWBS Zone Field Guide 9

    Fine-textured Luvisols dominate in the southern half of the plateau where morainal and (glacio)lacustrine parent materials occur, grad-ing to Gleysols where drainage is impeded. Organic soils (Mesisols and Fibrisols) dominate the Fort Nelson Lowlands where discontinu-ous permafrost occurs north of 58º N latitude. Brunisols occur on coarser-textured glaciofluvial deposits, and Regosols dominate the river floodplains as well as the (often unstable) valley sides above floodplains. Limited areas of grassland Chernozems and saline soils (Solonetzic Order) occur along the Peace River near the Alberta border. Significant portions of the Peace River lowlands near larger centres such as Fort St. John and Dawson Creek have been converted to agriculture.

    As is typical throughout the boreal forest, there is a tendency for Hemi-mor humus forms, derived mainly from feathermosses, to build up to considerable thicknesses as stands age and the broad-leaved tree com-ponent drops out. Repeated fire disturbance plays an important role in controlling this “smother moss” build-up that insulates the soil surface, triggering a gradual decline in soil temperature, drainage, nutrient turnover, and tree productivity, and eventually leading to bog develop-ment. This phenomenon, called paludification, is especially evident on cooler aspects across the north.

    3.3.2 Rocky Mountain Foothills and Rocky Mountains

    The transition from the flat lying/gently dipping sedimentary rocks of the Alberta Plateau to folded and faulted sedimentary rocks (lime-stones, siltstones, sandstones, and shales) underlying the slopes to the west delimits the boundary of the Rocky Mountain Foothills. The foothills stretch from the Liard River in the north, southeast to the Alberta border and beyond. The foothills give way to the more rugged Rocky Mountains to the west that are dominated by older sedimentary and metamorphic rocks, mainly limestones, quartzites, schists, and slates. The Rocky Mountain Foothills rise in elevation from about 000 m in the east to summits and ridges in the west with elevations of 800 to over 200 m. The Rocky Mountains to the west consist of the rugged Muskwa Ranges north of the Peace River, where peaks above 2400 m are typical, and the more subdued Hart Ranges in the south.

    The BWBS zone (mainly the BWBSwk) becomes more restricted in the foothills. It extends onto slopes of the eastern flanks to about 200 m

  • BWBS Zone Field Guide20

    and penetrates limited distances up some of the major river valleys that originate in the Rocky Mountains and flow east onto the Alberta plateau (e.g., Murray, Pine, Sukunka, Graham, Halfway, Sikanni Chief, Prophet, and Muskwa Rivers and the Peace Reach of Williston Lake). The BWBSdk occurs in the river valleys (e.g., the Akie, Kwadacha, Warneford, and Gataga Rivers) that drain the Rocky Mountains to the west into the Rocky Mountain Trench. The BWBSmk occurs in the valley bottoms of the northern Rockies where they drain into the Liard River. Near the divide of the Rocky Mountains, the BWBS gives way to the Sub-Boreal Spruce (SBS) zone. At elevations above 200 m in the foothills and the Rockies, the BWBS gives way to the Engelmann Spruce – Subalpine Fir (ESSF) zones in the south, and the Spruce – Willow – Birch (SWB) zone in the north.

    Medium- to fine-textured glacial till dominates valley bottoms where the BWBS occurs. Glacial meltwaters flowing east out of the Rockies deposited coarser-textured glaciofluvial deposits in some lower slope and valley-bottom locations. Localized, medium- to coarse-textured colluvial and fluvial deposits also occur throughout.

    3.3.3 Liard Plateau

    The Liard Plateau lies north of the Rocky Mountains with the south-erly extent of the Liard River forming the boundary. This plateau is underlain by folded sedimentary rocks, but, in contrast to the Rocky Mountains, consists of rounded and flat-topped summits lying mostly below 500 m in elevation. The Liard River has carved through the sedimentary bedrock to the south of the Plateau to elevations of 450 m, creating the Grand Canyon of the Liard. The BWBS extends through the Liard River Corridor in the southern portion of the plateau and penetrates up the Grayling, Scatter, and Crow Rivers that incise the plateau farther north. The BWBSmk occurs below 900–000 m and the BWBSwk occurs above 900 m in the eastern portions. The SWB zone is extensive on the Liard Plateau at elevations above 000–00 m. Medium- to fine-textured Luvisols derived from glacial till dominate the BWBS portion of this landscape.

    3.3.4 Liard Plain

    The Liard Plain is an extensive area of low relief west of the Liard

  • BWBS Zone Field Guide 2

    Plateau, within the drainage of the Liard River. With elevations largely between 750 and 050 m, this area is almost exclusively comprised of forests of the BWBS zone (BWBSmk subzone). This plain is surrounded on all sides by higher mountain and plateau areas. It is underlain by sedimentary bedrock, but the surface is shaped largely by glacial fea-tures and deposits such as eskers, drumlinized till, and shallow lakes. Coarser-textured Brunisols are associated with the eskers and other glaciofluvial deposits, while Luvisols are typical of the fine- to medium-textured till and glaciolacustrine deposits.

    3.3.5 Northern Rocky Mountain Trench

    The Northern Rocky Mountain Trench is a straight, narrow valley 3–4 km wide that extends from the headwaters of the Parsnip River in the south to the Liard Plain in the north along the western edge of the Rocky Mountains. Valley-bottom elevations range from 700 m in the north and south to a little more than 000 m at the divide (Sifton Pass) between the northward-flowing Kechika River and the southward-flow-ing Fox and Finlay Rivers.

    The BWBS (mk in the north, dk in the south) extends from the Liard Plain southward in the trench to the north end of Williston Lake, where it abuts the SBS zone in the valley bottom. The BWBS also extends up several river valleys draining the Rocky Mountains to the east and the Cassiar/Omineca Mountains to the west, such as the Ingenika, Finlay, Frog, Turnagain, Gataga, and Akie Rivers. The floor of the trench con-tains deep morainal as well as fluvial and glaciofluvial deposits.

    3.3.6 Cassiar and Omineca Mountains

    The mountains that rise up east of the Northern Rocky Mountain Trench and the Liard Plain are divided into two ranges: the Cassiar Ranges in the north and the Omineca Ranges south of the Finlay River. The Cassiar Mountains consist of folded sedimentary (includ-ing limestone and slate) and volcanic rocks as well as a core of granitic rock called the Cassiar Batholith where the highest peaks occur (e.g., Sharktooth Mountain west of Dall Lake at 2670 m). The BWBS zone occupies several large glacial valley systems within the Cassiar Ranges below 00–200 m elevation, including the Turnagain and Frog Rivers and their tributaries. The SWB and Boreal Altai Fescue Alpine (BAFA) zones dominate both the Cassiar and Omineca Ranges.

  • BWBS Zone Field Guide22

    The Omineca Mountains extend south from the Cassiar Mountains on the west flank of the trench to the southern end of Williston Lake, where they grade into the Nechako Plateau. These mountains are simi-lar in lithology to the Cassiar Mountains, with the largest peaks in the north being of granitic origin (e.g., Mount Cushing at 2650 m). Large river valleys such as the Finlay, Ingenika, Mesilinka, Osilinka, and Omineca contain significant areas of BWBSdk. In the south the BWBS gives way to the SBS zone and at higher elevations the ESSF replaces the SWB. The most southerly extent of the BWBS zone in the Omineca Mountains is at Germansen Lake.

    Soils in the valleys of both the Cassiar and Omineca Mountains are derived mostly from glacial till as well as colluvium on the valley sides and glaciofluvial and fluvial materials along the rivers.

    3.3.7 Stikine Plateau

    To the west of the Cassiar and Omineca Mountains lies the Stikine Pla-teau, a large area of dissected plateau country in the rainshadow of the Coast Mountains. This area is drained mostly by the Stikine and Taku Rivers and their tributaries. For the most part this area lies at or above 500 m in elevation but generally below the level of the surrounding mountains. The Stikine Plateau is subdivided into seven geographically and geologically distinct subunits: the Tahltan Highlands in the west, which is transitional to the Coast Mountains; the central Taku, Nahlin, and Klastline Plateaus; and the eastern Kawdy, Tanzilla, and Spatzizi Plateaus. Some units, such as the Taku Plateau and Tahltan Highlands, are more incised and dissected compared with the gentler relief of the Nahlin, Kawdy, and Spatzizi Plateaus. The highest peaks occur in the Tahltan Highlands, the most impressive of which is the shield volcano of Mount Edziza rising to 2790 m, with the brilliantly coloured Spec-trum Range extending to the south. The Grand Canyon of the Stikine River slices 300 m through the Stikine Plateau for 75 km upstream of Telegraph Creek and separates the Klastline and Tanzilla Plateaus.

    Sedimentary and volcanic rocks dominate throughout the Stikine Plateau. Major centres of past volcanic activity include Mount Edziza, Level Mountain, and the tuyas of the Kawdy Plateau. With much of the Stikine Plateau surface occurring above 500 m in elevation, it is domi-nated by the SWB and BAFA zones. The BWBSdk occurs below 200 m

  • BWBS Zone Field Guide 23

    in the major river valleys that dissect the plateau, including the Stikine, Spatzizi, Iskut, and Tanzilla in the south and the Nakina, Inklin, Sheslay, Nahlin, and Dease in the north. These valleys are dominated by glacial till and fluvial and glaciofluvial parent materials with some colluvial materials associated with the steeper terrain. Fine-textured Luvisols dominate on the tills, and coarser-textured Brunisols (and Regosols) on the fluvial and colluvial parent materials.

    Where major river valleys (e.g., the Stikine, Iskut, and Inklin/Taku Rivers) reach the western edge of the Plateau and enter the Coast Mountains, the BWBS zone gives way to the SBS and Interior Cedar – Hemlock (ICH) zones and eventually the CWH zone over a fairly rapid continental/maritime climate transition.

    3.3.8 Yukon Plateau

    The Yukon Plateau occurs to the northwest of the Stikine Plateau. In British Columbia, it is dominated by the central Teslin Plateau, which is flanked to the west by the Tagish Highland and to the east by the Nisutlin Plateau. The Yukon Plateau lies in the rainshadow of the St. Elias Mountains and Boundary Ranges to the west. Bedrock geology is dominantly sedimentary and volcanic with a large area of igneous rock occurring east of Atlin Lake.

    The Teslin Plateau is a gently sloping and dissected upland area that stretches from Atlin Lake in the west to Teslin Lake in the east. Mountains rise to 200 m and though much of the terrain is above 00 m, where the SWB and BAFA zones dominate, significant areas of BWBSdk occupy the broad valley bottoms of Atlin Lake and Teslin Lake and the narrower valleys in between. Glacial till, drumlin features, and lakes formed in glacial depressions characterize the Teslin Plateau where Luvisols (and Brunisols) are the typical soils.

    On the Nisultan Plateau to the east of Teslin Lake and in the Swift River and Jennings River drainages, the relatively low-lying terrain gives rise to extensive areas of BWBSdk. The Tagish Highland occurs north of the Taku River in the western Yukon Plateau and represents a narrow transition between the Teslin Plateau and the Coast Mountains, where dissection increases and mountain peaks are somewhat higher. Here the BWBSdk occurs extensively in the glacial till–dominated valley bot-toms of Tagish and Atlin Lakes.

  • BWBS Zone Field Guide24

    3.3.9 Skeena Mountains

    The Skeena Mountains occur south of the Stikine Plateau and constitute the major mountainous area in the northern Interior of British Colum-bia, extending approximately 400 km south to the Bulkley Valley at the northern edge of the Interior Plateau. The Skeena Mountains form the headwaters of three major northern river systems: the Skeena, Stikine, and Nass drainages. The BWBS zone extends only into the extreme northern portion of the Skeena Mountains as narrow fingers in the upper Klappan and Spatzizi Rivers. South of these areas, the ESSF, ICH, and SBS zones dominate below the extensive alpine zone in the Skeena Mountains.

    3.3.0 Coast Mountains and St. Elias Mountains

    North of the Nass River, the Coast Mountains are represented by the Boundary Ranges: rugged, granitic, ice-capped mountains that extend the length of the British Columbia–Alaska border. Maritime influences extend up the major river valleys (Iskut, Stikine, and Taku Rivers) that dissect the Boundary Ranges, and thus the BWBS zone does not pen-etrate west into the Coast Mountains. In the extreme northwest corner of the province (the Haines Triangle), however, where the Alsek and Tatshenshini Rivers flow through the ice-draped St. Elias Mountains, a unique subzone of the BWBS occurs along the valley bottoms. This very wet, cool (BWBSvk) subzone experiences strong maritime/conti-nental gradients with heavy snowpacks and strong winds. The Bound-ary Ranges and St. Elias Mountains, both heavily influenced by moist Pacific air masses, create a rainshadow effect that yields the relatively dry boreal climate typical of the Yukon and Stikine Plateaus to the east.

    3.4 Climate of the Boreal Forest

    Most of the BWBS zone experiences a continental climate, dominated by both continental arctic and continental polar air masses (Ahrens 994). This zone is characterized by long, cold winters and short grow-ing seasons (Meidinger and Pojar [editors] 99). The mean annual temperature ranges from –2.4 to 3.6°C and annual precipitation ranges between 34 and 897 mm (Table 3.2). The BWBS is generally drier, especially in the winter, and warmer in the summer than the adjacent

  • BWBS Zone Field Guide 25

    portions of the zones (Table 3.2). The BWBS is also colder in the winter than the adjacent portions of lowland zones (Table 3.2).

    Approximately 32% of precipitation falls as snow, which may seem low considering that any day of the year can see temperatures below freezing. In the portion of the BWBS east of the Rockies, winters generally receive the least amount of precipitation due to cold, dry, arctic high-pressure systems. The inter-annual variability of the Arctic Oscillation controls the amount and the extent of cold arctic air in the winter. Summer rain makes up most of the precipitation and is typically double any other season. This is due mainly to moist weather systems from the Pacific Ocean flowing across the Rocky Mountains, but also from temperatures warm enough to generate convective showers and thundershowers. When inverted troughs of low pressure from Alberta force air to flow west and condense against the mountains, they bring high precipitation amounts in short time periods. Areas of the BWBS in and west of the Rocky Mountains typically experience milder condi-tions in the winter but are cooler in the summer due to the influence of the mountainous topography, which blocks the influence of the arctic fronts in winter but allows cold air to seep down from higher elevations in the summer. In this area, annual precipitation is comprised more evenly of rain and snow; however, rain still dominates as most precipi-tation falls in the summer.

    Temperatures in the BWBS can be extreme. Fort Nelson, for example, experiences one of the largest spreads in temperature of the province, with summer highs reaching above 30°C and winter lows commonly below –40°C (Heidorn 2004). The climate station at the Fort Nelson air-port recorded both the highest extreme maximum temperature (36.7°C, 942) and the lowest extreme minimum temperature (–5.7°C, 947) for the BWBS (Table 3.3). The warmest month is July, in which mean temperatures range from .7 to 6.7°C. The coldest month is January; mean temperatures range from –23.8 to –7.4°C. The temperature spread throughout the year is less extreme in western portions of the BWBS.

    Climate conditions of the BWBS subzones from 97 to 2000 are described in Table 3.4. The warmest and the wettest subzone is the BWBSwk, with a mean annual temperature of 2.9°C and mean annual precipitation of 743 mm. The coldest subzone is the BWBSmk, with a

  • BWBS Zone Field Guide26

    TABL

    E 3.

    2 Co

    mpa

    rison

    of c

    limat

    e va

    riabl

    es fo

    r the

    BW

    BS z

    one

    and

    adja

    cent

    por

    tions

    of z

    ones

    in n

    orth

    ern

    Briti

    sh C

    olum

    biaa

    Clim

    ate

    char

    acte

    rist

    ics

    BW

    BSb

    SBSb

    SW

    Bc

    ESSF

    c IC

    Hc

    Ann

    ual p

    reci

    pita

    tion

    (mm

    ) M

    ean

    535

    708

    778

    1125

    10

    14

    SDd

    96

    257

    292

    788

    375

    R

    ange

    34

    1–89

    7 46

    3–18

    93

    496–

    1691

    45

    5–32

    26

    580–

    2445

    Mea

    n gr

    owin

    g se

    ason

    M

    ean

    317

    296

    408

    424

    362

    prec

    ipita

    tion

    (mm

    ) SD

    48

    67

    11

    8 19

    6 94

    (May

    –Sep

    tem

    ber)

    R

    ange

    14

    5–44

    3 19

    2–54

    0 23

    0–59

    0 19

    3–94

    2 25

    3–74

    2A

    nnua

    l sno

    wfa

    ll (c

    m)

    Mea

    n 16

    9 31

    6 36

    4 56

    8 43

    3

    SD

    41

    166

    223

    479

    184

    R

    ange

    11

    0–33

    2 14

    8–10

    51

    161–

    1019

    16

    3–24

    24

    183–

    938

    Ann

    ual t

    empe

    ratu

    re (°

    C)

    Mea

    n 1.7

    2.

    2 -1

    .0

    1.1

    2.6

    SD

    1.0

    0.

    6 1.1

    1.6

    1.7

    R

    ange

    -2

    .4–3

    .6

    0.7–

    4.2

    -2.5

    –1.1

    -3.5

    –6.6

    -0

    .7–6

    .3M

    ean

    war

    mes

    t mon

    th

    Mea

    n 14

    .7

    13.8

    11

    .6

    12.3

    13

    .5te

    mpe

    ratu

    re (°

    C)

    SD

    0.8

    0.8

    1.4

    1.6

    1.4

    Ran

    ge

    11.7

    –16.

    7 11

    .4–1

    5.9

    9.4–

    13.5

    7.

    5–16

    .6

    10.3

    –16.

    2M

    ean

    cold

    est m

    onth

    M

    ean

    -12.

    4 -1

    0.5

    -12.

    9 -1

    0.0

    -8.9

    tem

    pera

    ture

    (°C

    ) SD

    2.

    5 1.0

    1.3

    2.

    0 2.

    5

    Ran

    ge

    -23.

    8–(-

    7.4)

    -1

    4.4–

    (-7.

    7)

    -16.

    8–(-

    11.0

    ) -1

    4.2–

    (-4.

    4)

    -13.

    1–(-

    3.8)

    Extr

    eme

    min

    imum

    M

    ean

    -44.

    2 -4

    1.9

    -44.

    3 -4

    0.9

    -38.

    4te

    mpe

    ratu

    re (°

    C)

    SD

    2.3

    1.3

    1.2

    2.6

    4.0

    R

    ange

    -4

    9.7–

    (-37

    .3)

    -46.

    4–(-

    38.0

    ) -4

    7.3–

    (-42

    .5)

    -45.

    3–(-

    31.5

    ) -4

    3.9–

    (-29

    .2)

  • BWBS Zone Field Guide 27

    Clim

    ate

    char

    acte

    rist

    ics

    BW

    BSb

    SBSb

    SW

    Bc

    ESSF

    c IC

    Hc

    Gro

    win

    g de

    gree

    day

    s M

    ean

    1451

    11

    47

    1737

    12

    50

    979

    (< 0

    °C)

    SD

    288

    130

    191

    286

    308

    R

    ange

    93

    5–26

    82

    870–

    1730

    14

    17–2

    161

    422–

    2031

    41

    1–15

    85G

    row

    ing

    degr

    ee d

    ays

    Mea

    n 11

    49

    1016

    63

    0 78

    5 99

    7(>

    5°C

    ) SD

    14

    0 12

    9 20

    4 25

    6 24

    9

    Ran

    ge

    702–

    1421

    66

    9–13

    65

    382–

    946

    181–

    1711

    46

    8–15

    83D

    ate

    of g

    row

    ing

    degr

    ee

    Mea

    n M

    ay 16

    M

    ay 2

    6 Ju

    n 12

    Ju

    n 06

    M

    ay 2

    5da

    y >

    5°C

    reac

    hes 1

    00

    SD

    7 8

    10

    15

    13(b

    udbu

    rst)

    R

    ange

    M

    ay 7

    –Jun

    8

    May

    10–J

    un 19

    M

    ay 2

    7–Ju

    n 26

    A

    pr 2

    5–Ju

    l 25

    Apr

    28–

    Jun

    26Fr

    ost-

    free

    per

    iod

    M

    ean

    92

    83

    56

    73

    95(d

    ays)

    SD

    9

    12

    19

    27

    22

    Ran

    ge

    54–1

    11

    50–1

    08

    13–8

    8 2–

    161

    48–1

    43N

    umbe

    r of f

    rost

    -fre

    e

    Mea

    n 15

    5 15

    4 12

    4 14

    3 16

    4da

    ys

    SD

    8 9

    15

    23

    26

    Ran

    ge

    126–

    175

    128–

    177

    102–

    153

    78–2

    27

    94–2

    19

    a Bi

    ogeo

    clim

    atic

    uni

    ts u

    sed

    with

    in th

    e ad

    jace

    nt z

    ones

    as f

    ollo

    ws:

    SBS

    (Sub

    -Bor

    eal S

    pruc

    e) =

    mc2

    , mk

    , mk2

    , wk2

    , wk3

    ; SW

    B (S

    pruc

    e –

    Will

    ow

    – Bi

    rch)

    = d

    k, m

    k, v

    k; E

    SSF

    (Eng

    elm

    ann

    Spru

    ce –

    Sub

    alpi

    ne F

    ir =

    mc,

    mv

    , mv2

    , mv3

    , wv;

    ICH

    (Int

    erio

    r Ced

    ar –

    Hem

    lock

    ) = m

    c2, w

    c, v

    c.

    b U

    pdat

    ed fr

    om R

    eyno

    lds (

    989

    ) for

    97

    –200

    0 cl

    imat

    e no

    rmal

    s usi

    ng C

    limat

    eBC

    (Wan

    g et

    al.

    2006

    ).c

    Supp

    lem

    ente

    d Re

    ynol

    ds (

    989)

    wea

    ther

    stat

    ion

    loca

    tions

    with

    add

    ition

    al d

    ata.

    d

    Stan

    dand

    dev

    iatio

    n of

    the

    mea

    n.

  • BWBS Zone Field Guide28

    TABL

    E 3.

    3 BW

    BS 19

    71–2

    000

    clim

    ate

    norm

    als:

    Env

    ironm

    ent C

    anad

    a w

    eath

    er st

    atio

    ns w

    ith W

    orld

    Met

    eoro

    logi

    cal O

    rgan

    izat

    ion

    Stan

    dard

    sa

    El

    evat

    ion

    M

    ATb

    Extr

    eme

    Extr

    eme

    MA

    Pb

    Ann

    ual

    Ann

    ual

    DD

    Stat

    ion

    (m)

    Subz

    one

    (°C

    ) m

    ax (°

    C)

    min

    (°C

    ) (m

    m)

    rain

    (mm

    ) sn

    ow (c

    m)

    > 5

    °C

    Atli

    n 64

    7 dk

    0.

    5 30

    .6

    -50.

    0 34

    7 19

    2 15

    5 83

    0D

    awso

    n C

    reek

    A

    655

    mw

    1.6

    34

    .5

    -49.

    2 48

    2 32

    6 17

    4 12

    36D

    ease

    Lak

    e 80

    7 dk

    -0

    .8

    35.3

    -5

    1.2

    426

    265

    218

    774

    Fort

    Nel

    son

    A

    382

    mk

    -0.7

    36

    .7

    -51.7

    45

    2 32

    0 17

    8 13

    20Fo

    rt S

    t. Jo

    hn A

    69

    5 m

    w

    2.0

    33.6

    -4

    7.2

    466

    313

    186

    1312

    Ger

    man

    sen

    Land

    ing

    766

    dk

    1.0

    34.5

    -4

    8.8

    538

    321

    243

    968

    a En

    viro

    nmen

    t Can

    ada

    (200

    8).

    b M

    AT

    = m

    ean

    annu

    al te

    mpe

    ratu

    re, M

    AP

    = m

    ean

    annu

    al p

    reci

    pita

    tion.

  • BWBS Zone Field Guide 29

    TABL

    E 3.

    4 Su

    mm

    ary

    of c

    limat

    e da

    ta fo

    r bio

    geoc

    limat

    ic u

    nits

    with

    in th

    e BW

    BS z

    one

    of n

    orth

    ern

    Briti

    sh C

    olum

    biaa

    Clim

    ate

    char

    acte

    rist

    ics

    BWBS

    dk

    BWBS

    mk

    BWBS

    mw

    BW

    BSw

    k1

    BWBS

    wk2

    BW

    BSw

    k3

    Ann

    ual p

    reci

    pita

    tion

    M

    ean

    498

    499

    515

    743

    571

    570

    (mm

    ) SD

    b 85

    46

    64

    62

    43

    10

    R

    ange

    34

    1–64

    4 43

    1–58

    4 42

    4–74

    9 64

    4–89

    7 48

    5–62

    5 56

    0–57

    9M

    ean

    grow

    ing

    seas

    on

    Mea

    n 26

    6 32

    2 31

    5 38

    9 39

    5 39

    8pr

    ecip

    itatio

    n (m

    m)

    SD

    55

    51

    27

    21

    37

    10(M

    ay–S

    epte

    mbe

    r)

    Ran

    ge

    145–

    443

    229–

    416

    266–

    404

    356–

    421

    317–

    428

    388–

    407

    Ann

    ual s

    now

    fall

    (cm

    ) M

    ean

    189

    155

    156

    241

    152

    156

    SD

    47

    40

    26

    35

    24

    4

    R

    ange

    11

    0–25

    8 12

    1–24

    8 11

    0–24

    2 19

    7–33

    2 13

    4–21

    0 15

    2–15

    9

    Ann

    ual t

    empe

    ratu

    re

    Mea

    n 1.2

    -0

    .6

    1