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Incomplete Gut- gut with only an opening at the blastopore (a.k.a blind gut)
Complete Gut- has two openings (mouth and anus)
Diploblastic- animals with two germ layers
Triploblastic- animals with three germ layersMesoderm- between the ectoderm and the endoderm (muscular system, reproductive system,
peritoneum, calcareous plates)
-initial cells of mesoderm come from endoderm
Coelom- body cavity completely surrounded by mesoderm
-when coelom formation is complete, the body has three germ layers and two cavities(gut and coelomic cavity)
D euterostome D evelopment:
Radial Cleavage- embryonic cells are arranged in radial symmetry around the animal vegetalaxis
Regulative Development- fate of cell is not fixed early in development; in early developmenteach cell is able to produce an entire embryo if separated from other cells
-blastopore becomes the anus
-coelom formation is enterocoely
Enterocoely- both mesoderm and coelom are mad at the same time
-there are variations in deuterostome cleavage
*Bilateral Cleavage- (ex. Ascidian chordates) anteroposterior axis is established prior tofertilization; first cleavage furrow passes through the animal-vegetal axis
Rotation Cleavage- (isolecithcal) cells at the second cell division appeqar to have turned relativeto each other (90 degrees)
Trophoblast- form embryonic portion of the placenta when the embryo implants in the uterinewall
Inner cell mass- cells that give rise to the embryo proper form here (in mice)
Discoidal Cleavage- (telolecithal) due to great mass of yolk, cleavage is confined to a small discof cytoplasm lying atop a mound of yolk (in chickens)
-coelomic vesicles: (deuterostome) pinch off from archenteron to form left and rightcoelomic compartments
*-primitive streak: the anteroposterior axis of the embryo and the center of early growth
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-Blastoderm consists of two layers: epiblast and hypoblast with blastocoels betweenthem
-Epiblast and hypoblast end up forming endoderm and mesoderm
Protostome D evelopment:
Spiral Cleavage- blastomeres cleave at 45-degree angles; cells lie between cells of the underlyinglayer
Mosaic Development- embryonic development is characterized by each part of the embryo
Morphogenetic Determinants- determined by distribution of certain proteins and mRNAs (this iswhy Protostome is mosaic)
-blastopore becomes the mouth
-coelom formation is schizocoely
Schizocoey- coelom formed by the splitting of embryonic mesoderm
-some protostomes do not develop a coelom
Acoelomate- animals without a coelom
Pseudocoelom- a body cavity formed from a persistent blastocoels and linked with mesoderm ononly one side
Lophotrochozoan Protostomes- (segmented worms & mollusks) have lophophore or trochophorelarva
Ecdysozoan Protostomes- (arthropods) members shed their cuticle as they grow
*Superficial Cleavage- centrally located mass of yolk restricts cleavage to the cytoplasmic rim of the egg
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Chapter 9
Grades of Organization
1. Protoplasmic Grade (unicellular)
2. Cellular Grade3. Cell-Tissue Grade
4. Tissue-Organ Grade
5. Organ System Grade
Metazoa- multicellular animals
Eumetazoan- multicellular animals with distinct germ layers that form true tissues
Parenchyma- chief functional cells of an organ
Stroma- supportive tissue
Organ Systems: skeletal, muscular, urinary, reproductive, nervous, digestive, integumentary,respiratory, circulatory, excretory, endocrine and immune
-Animal body plans differ in the grade of organization, in body symmetry, in the number of embryonic germ layers and in the number of body cavities
*Spherical Symmetry- any plane passing through the center divides a body intoequivalent/mirrored image; chiefly among unicellular organisms
*Radial Symmetry- can be cut in more than one way to be symmetrical*Biradial Symmetry-
Bilateral Symmetry- animals that can be divided along a sagittal plane into two mirrored portions
Cepalization- sensory organs and specialized appendages become localized in the head end of animals
Anterior- head area Posterior- tail end
Dorsal- back side Ventral- front/belly side
Lateral- the sides Distal- parts farther from the middle of the body
Proximal- parts closer to the body
Frontal Plane- divides body into dorsal and ventral
Sagittal Plane- divide animal into right and left halves
Transverse Plane- separates anterior from posterior at middle of body
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Pectoral- chest region Pelvia- hip region
Acoelomate- mesodermal cells completely fill the blastocoels, leaving the gut (archenteron) asthe only body cavity
*parenchyma- a spongy mass of space-filling cells between the ectodermal epidermis andthe endodermal digestive tract
Pseudocoelomate- mesodermal cells line the outer edge of the blastocoels, leaving two bodycavities
Pseudocoelom- a false coelm, where the blastocoels was
Peritoneum- thin cellular membrane derived from medoderm lining it
Mesenteries- suspended organs in the coelom
-Pseudocoelom lacks a peritoneum
Segmentation (metamerism)- serial repetition of similar body segments along longitudinal axisof the body
Metamere/Somite- the segments that make up the segmentation
-segmentation permits greater body mobility and complexity of structure and function
-Metazoan animals contain two noncellular components
1. Body Fluids (separated into two fluid components)
-Intracellular Space- within the bodys cells
-Extracellular Space- outside the cells
-blood plasma- fluid portion of body
-interstitial fluid-(tissue fluid) occupies the space surrounding cells
2. Extracellular Structural Elements (supportive material of the organism) ex. Looseconnective tissue, cartilage, bone and cuticle
Tissue- group of similar cells specialized for performance of common function
Histology- study of tissues
Epithelial Tissue
-sheet of cells that covers an external or internal surface
-a large variety of transport molecules are located on epithelial cell membranes
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Connective Tissue
-diverse group of tissues that serve carious binding and supportive functions
-specialized connective tissue: blood, lymph, adipose, cartilage and bone
Ground Substance- fibers are suspendedLoose Connective Tissue- composed of fibers and both fixed and wandering cellssuspended in a viscous fluid ground substance
Dense Connective Tissue- composed largely of densely packed fibers and little groundsubstance (ex. Tendons and ligaments)
Collagen- protein of great tensile strength
Muscular Tissue
-the most abundant tissue in the body o most animals
-originates from mesoderm
-its unit is the muscle fiber
-Sarcoplasm- unspecialized cytoplasm of muscles
-Myofibrils- contractile elements within the fiber
Nervous Tissue
-specialized for reception of stimuli and conduction of impulses from one region to
another -Neurons- basic functional unit of nervous system
-Neuroglia- variety of nonnervous cells that insulate neuron membranes and servevarious supportive functions