heritability of education rises with intergenerational ... · our results indicate that social...

3
Heritability of education rises with intergenerational mobility Per Engzell a,b,c,1 and Felix C. Tropf a,d,e a Nuffield College, University of Oxford, Oxford OX1 1NF, United Kingdom; b Leverhulme Centre for Demographic Science, University of Oxford, Oxford OX1 1JD, United Kingdom; c Swedish Institute for Social Research, Stockholm University, 106 91 Stockholm, Sweden; d Laboratoire de Sociologie Quantitative, ´ Ecole Nationale de la Statistique et de l’Administration ´ Economique, 99120 Palaiseau, France; and e Department of Sociology, Center for Research in Economics and Statistics, 99120 Palaiseau, France Edited by Kenneth W. Wachter, University of California, Berkeley, CA, and approved November 13, 2019 (received for review August 9, 2019) As an indicator of educational opportunity, social scientists have studied intergenerational mobility—the degree to which chil- dren’s attainment depends on that of their parents—and how it varies across place or time. We combine this research with behav- ior genetics to show that societal variation in mobility is rooted in family advantages that siblings share over and above genetic transmission. In societies with high intergenerational mobility, less variance in educational attainment is attributable to the shared sibling environment. Variance due to genetic factors is largely constant, but its share as a part of total variance, heritabil- ity, rises with mobility. Our results suggest that environmental differences underlie variation in intergenerational mobility, and that there is no tension between egalitarian policies and the realization of individual genetic potential. educational attainment | intergenerational mobility | heritability I ntergenerational education mobility—how strongly educa- tional attainment persists from parent to child—is commonly used to indicate societies’ degree of openness or equality of oppor- tunity (1, 2). A limitation of this literature is that it often is silent on the channels of transmission. Yet, we may view genetic trans- mission differently from other advantages such as parents’ ability to pay for good neighborhoods, schools, or access to college (3, 4). Insofar as genetic factors capture relevant abilities, their influence is consistent with meritocratic norms (5–7). Such norms can be motivated on grounds of efficiency, as a society’s viability depends on its ability to attract competent leaders and innovators (8, 9). In other words, it matters not only whether education is inher- ited, but also how (3). One approach to the “how” question comes from behavior genetics (10). By comparing outcomes for family members with varying degree of genetic resemblance— typically, twins—we can partition variance in an outcome to that attributable to genetic factors (heritability, h 2 ), shared sib- ling environment (c 2 ), and idiosyncratic factors (e 2 ) (11). While such studies potentially tell us much about the distribution of opportunities, societal comparison has rarely been central to them. In this study, we ask: Where intergenerational mobil- ity is higher, does the balance of “nature” and “nurture” in educational attainment differ? Results Intergenerational Mobility. Fig. 1A plots the parent–offspring cor- relation in years of schooling for children born in the 1940s to 1980s in 10 countries for which genetic data are available. A lower correlation implies more intergenerational mobility. These data, from the World Bank, confirm previous research: Educa- tional mobility increased in the past and is higher in northern Europe and Australia—places with liberal welfare states—than in the United States or southern Europe (2, 12). Behavior Genetics Estimates. In Fig. 1 B and C, we link intergener- ational correlations with genetic data from twin studies (11). On the vertical axes, h 2 (Fig. 1B) tells us to what extent differences in educational attainment result from the genetic lottery, while c 2 (Fig. 1C) indicates the importance of other family background factors. In societies with high mobility—that is, a lower intergen- erational correlation—the relative explanatory power of genetic factors (h 2 ) is stronger, while that of shared sibling environment (c 2 ) is weaker. Table 1 shows that this result remains when con- trolling for gender and adjusting standard errors for correlation between male and female subsamples. The residual component e 2 is largely unrelated to mobility. Absolute and Relative Variance. That relative genetic influence h 2 is higher in more mobile societies could mean 2 things. Either 1) environmentally induced (and thereby total) variance is decreas- ing and/or 2) a lucky genetic draw is reaping higher rewards in absolute terms. If possibility 2, structural inequality may sim- ply be replaced by genetic inequality, which could be seen as no less troubling. To test this, we destandardize h 2 , c 2 , e 2 by total population variance (Table 1). Doing so supports possibil- ity 1 over possibility 2: the prevailing result is that high mobility goes together with a lower absolute influence of the shared envi- ronment, c 20 . In other words, h 2 and mobility correlate because mobility reduces total variance while genetic variance remains constant, not because genes matter more in an absolute sense. Discussion Our results indicate that social mobility is improved by reduc- ing social inheritance, a process that brings genetic influences to the fore. Besides their general interest, these findings speak to several academic debates. In social mobility research, there has been controversy about the extent to which policy can achieve lasting change. Critics have seen political attempts to increase mobility as either inadequate (13) or futile, adding the assump- tion that inheritance is largely genetic in origin (14). Our study challenges these views by showing that societal variation in mobility is not only substantial but firmly rooted in the removal of environmental barriers. Meanwhile, geneticists have surmised that more ample oppor- tunities may translate into a higher h 2 for developmental out- comes (15, 16). Based on this, it stands to reason that h 2 should be higher in societies where education policy promotes social mobility (17). Yet, comparative evidence has been lacking, and the canonical reference is to a study of Norwegian cohorts over time (18). A separate literature examines differences in h 2 between socioeconomic strata within a society, focusing mostly on IQ, but with mixed results (16, 17). Our study contributes to these debates using large-scale cross-national data on an outcome of high policy relevance. Author contributions: P.E. and F.C.T. designed research; P.E. performed research; P.E. analyzed data; and P.E. and F.C.T. wrote the paper.y The authors declare no competing interest.y This open access article is distributed under Creative Commons Attribution License 4.0 (CC BY).y 1 To whom correspondence may be addressed. Email: per.engzell@nuffield.ox.ac.uk.y First published December 2, 2019. 25386–25388 | PNAS | December 17, 2019 | vol. 116 | no. 51 www.pnas.org/cgi/doi/10.1073/pnas.1912998116 Downloaded by guest on March 24, 2020

Upload: others

Post on 18-Mar-2020

8 views

Category:

Documents


0 download

TRANSCRIPT

Heritability of education rises withintergenerational mobilityPer Engzella,b,c,1 and Felix C. Tropfa,d,e

aNuffield College, University of Oxford, Oxford OX1 1NF, United Kingdom; bLeverhulme Centre for Demographic Science, University of Oxford, Oxford OX11JD, United Kingdom; cSwedish Institute for Social Research, Stockholm University, 106 91 Stockholm, Sweden; dLaboratoire de Sociologie Quantitative,Ecole Nationale de la Statistique et de l’Administration Economique, 99120 Palaiseau, France; and eDepartment of Sociology, Center for Research inEconomics and Statistics, 99120 Palaiseau, France

Edited by Kenneth W. Wachter, University of California, Berkeley, CA, and approved November 13, 2019 (received for review August 9, 2019)

As an indicator of educational opportunity, social scientists havestudied intergenerational mobility—the degree to which chil-dren’s attainment depends on that of their parents—and how itvaries across place or time. We combine this research with behav-ior genetics to show that societal variation in mobility is rootedin family advantages that siblings share over and above genetictransmission. In societies with high intergenerational mobility,less variance in educational attainment is attributable to theshared sibling environment. Variance due to genetic factors islargely constant, but its share as a part of total variance, heritabil-ity, rises with mobility. Our results suggest that environmentaldifferences underlie variation in intergenerational mobility, andthat there is no tension between egalitarian policies and therealization of individual genetic potential.

educational attainment | intergenerational mobility | heritability

Intergenerational education mobility—how strongly educa-tional attainment persists from parent to child—is commonly

used to indicate societies’ degree of openness or equality of oppor-tunity (1, 2). A limitation of this literature is that it often is silenton the channels of transmission. Yet, we may view genetic trans-mission differently from other advantages such as parents’ abilityto pay for good neighborhoods, schools, or access to college (3, 4).Insofar as genetic factors capture relevant abilities, their influenceis consistent with meritocratic norms (5–7). Such norms can bemotivated on grounds of efficiency, as a society’s viability dependson its ability to attract competent leaders and innovators (8, 9).

In other words, it matters not only whether education is inher-ited, but also how (3). One approach to the “how” questioncomes from behavior genetics (10). By comparing outcomes forfamily members with varying degree of genetic resemblance—typically, twins—we can partition variance in an outcome tothat attributable to genetic factors (heritability, h2), shared sib-ling environment (c2), and idiosyncratic factors (e2) (11). Whilesuch studies potentially tell us much about the distribution ofopportunities, societal comparison has rarely been central tothem. In this study, we ask: Where intergenerational mobil-ity is higher, does the balance of “nature” and “nurture” ineducational attainment differ?

ResultsIntergenerational Mobility. Fig. 1A plots the parent–offspring cor-relation in years of schooling for children born in the 1940s to1980s in 10 countries for which genetic data are available. Alower correlation implies more intergenerational mobility. Thesedata, from the World Bank, confirm previous research: Educa-tional mobility increased in the past and is higher in northernEurope and Australia—places with liberal welfare states—thanin the United States or southern Europe (2, 12).

Behavior Genetics Estimates. In Fig. 1 B and C, we link intergener-ational correlations with genetic data from twin studies (11). Onthe vertical axes, h2 (Fig. 1B) tells us to what extent differences ineducational attainment result from the genetic lottery, while c2

(Fig. 1C) indicates the importance of other family backgroundfactors. In societies with high mobility—that is, a lower intergen-erational correlation—the relative explanatory power of geneticfactors (h2) is stronger, while that of shared sibling environment(c2) is weaker. Table 1 shows that this result remains when con-trolling for gender and adjusting standard errors for correlationbetween male and female subsamples. The residual componente2 is largely unrelated to mobility.

Absolute and Relative Variance. That relative genetic influence h2

is higher in more mobile societies could mean 2 things. Either 1)environmentally induced (and thereby total) variance is decreas-ing and/or 2) a lucky genetic draw is reaping higher rewards inabsolute terms. If possibility 2, structural inequality may sim-ply be replaced by genetic inequality, which could be seen asno less troubling. To test this, we destandardize h2, c2, e2 bytotal population variance (Table 1). Doing so supports possibil-ity 1 over possibility 2: the prevailing result is that high mobilitygoes together with a lower absolute influence of the shared envi-ronment, c2′. In other words, h2 and mobility correlate becausemobility reduces total variance while genetic variance remainsconstant, not because genes matter more in an absolute sense.

DiscussionOur results indicate that social mobility is improved by reduc-ing social inheritance, a process that brings genetic influences tothe fore. Besides their general interest, these findings speak toseveral academic debates. In social mobility research, there hasbeen controversy about the extent to which policy can achievelasting change. Critics have seen political attempts to increasemobility as either inadequate (13) or futile, adding the assump-tion that inheritance is largely genetic in origin (14). Our studychallenges these views by showing that societal variation inmobility is not only substantial but firmly rooted in the removalof environmental barriers.

Meanwhile, geneticists have surmised that more ample oppor-tunities may translate into a higher h2 for developmental out-comes (15, 16). Based on this, it stands to reason that h2 shouldbe higher in societies where education policy promotes socialmobility (17). Yet, comparative evidence has been lacking, andthe canonical reference is to a study of Norwegian cohortsover time (18). A separate literature examines differences in h2

between socioeconomic strata within a society, focusing mostlyon IQ, but with mixed results (16, 17).

Our study contributes to these debates using large-scalecross-national data on an outcome of high policy relevance.

Author contributions: P.E. and F.C.T. designed research; P.E. performed research; P.E.analyzed data; and P.E. and F.C.T. wrote the paper.y

The authors declare no competing interest.y

This open access article is distributed under Creative Commons Attribution License 4.0(CC BY).y1 To whom correspondence may be addressed. Email: [email protected]

First published December 2, 2019.

25386–25388 | PNAS | December 17, 2019 | vol. 116 | no. 51 www.pnas.org/cgi/doi/10.1073/pnas.1912998116

Dow

nloa

ded

by g

uest

on

Mar

ch 2

4, 2

020

BRIE

FRE

PORT

SOCI

AL

SCIE

NCE

S

AUS

DEU

DNK

ESPFINGBR

ITA

NOR

SWEUSA

AUS

DEUDNK

ESP

FINGBR

ITA

NOR

SWE

USA

.2

.3

.4

.5

.6

.2

.3

.4

.5

.6

1940 1960 1980

Daughters

Sons

Inte

rgen

erat

iona

l cor

rela

tion

(less

mob

ility

)

Birth cohort

A

AUSAAAAAAUUUU 004000

S50US5AA S5

AUS60AU 0

FIN40FS4EUDEE FINFINFF

IIIITTTTTTA40A40AAAATTTT

NORR40NORRNOR 0000

OR50RNOO4040000US40US400004040005050N4N4S4S4OU40U4044EUEUUUFINFINFINFINFFFF 000

ESP500

N

US 0SA400S 0

USA80U

AUS40U NNN

US50US5AUUS5S5S5USUSAAAA SS

AUS60AUDNK70DNK70KAUS4AUS4UU

450N44FIN40FFIN40FIN40FFFINFINFINFINFFFF 00000040ESPESP

DEU40U40D

ITA40044440000TT

NOR400DDDNOR50R50RNNUS60US60US60US6000DEUDEUDEUDEUDD00000000000

ESP500E40E40WE40WE40SWSWWE40E40SSSSSSSSSSSS 0000S40S40S40S400000EEEES4S4S4S4SSWWWWWW0000404040400000WW4W0WW

NOONO00GBR4GBR400RRRRR4R4444444040000000RRBRR 00000000000000000WWWWWWWSWSWSWSWSWSWSWWSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSS

GBR60GBR6066USA40UUUUUUUUSSS

SUSSA80SUSUSUSA4USA4USS0000UU

0

.2

.4

.6

.8

h2 (g

enet

ic e

ndow

men

t mat

ters

mor

e )

.2 .3 .4 .5 .6 .7

B

A

AUS50AA SSSSS5

AUS600

FIN4NF N40N40N4N

U404DEU404U404DEDDEE505055555500

ITA40AATT

NOR40R

FFNOR50R

P500ESP500P500EEEEE

USA4A4000A405050

USA80U

AUS40AUS4UAA

0US50U 0AUSUAUUUUUSSSSS5S5AUS5AUS5AUUUSUS

AUS60AUAU

ADNK70KNK70KNK70K

ESIN40NNNNESPESPEEEEEII

DEU400000

IIITTTA400TT

NOR40OR400OR40OORRNOR50RNONONN

ESP500EEIIIEE40E40E40E404444GBGBBR40BR400BB 404440AUSAUSAUSAUSSSSSSSSS 0000RRR 0000000000000000000000S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S40S4S4S4S4S4S4S4S4S4S4S4S4S4S40S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4SSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSWWWW00000000000000000000000000000000000RR4S4S4S4SSSSRRSSSSRSSSSSSSSSSSSRSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSSBR60BR60BR60BR606600000000AUSAUSAUSAUS

FINFINFINFINFINFINFINFINFFFF NNNNNNNNGGGGGGGGAUAUAUAUAUAUAU SWSWSWSWSWSWSWWWWWWWWWWWWWWWWWWWWWWWWWWS40S40S4S4S40S4S4S4S4S4S4S4S4S4S400004040404040404040404040404040000000404044404444444444404044404444444444000000000000000000000000000000000000000000000000000000000AUSAUSAUSAUSAUSAUSAUSAUSAUSAUSAUSAUSAUSAUSAUSS4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S4S440444404444444444444444444444440444404444444444444444444444444440000444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444444000000000000000000000000000000000000000000000000000000000000000000000000000000000000000000000UUUSA40USA40UUUUUU

SP4SPSSPSSUSA80AA88800E40E40E40E400000000000EEEEE4EE4EE40E40E40E40E40E40E4044444444444444444440000400000000000000EEEEEEEE4EEEE4EEWWWWWWW000000000000000000WEWEWEWEWEWEWEWWWWWWWWWWW0000000000000000000000000000WEWEWEWEEEEWEWEWEWEEEEEWWWWWWW000000000000000000000000000000000000000000000000000000000000WWWWWWWWWWWWWWW000000000000000000000000000000000000000000000000000000000000000

U

0

.2

.4

.6

.8

c2 (s

iblin

g en

viro

nmen

t mat

ters

mor

e )

.2 .3 .4 .5 .6 .7

DaughtersSons

C

Intergenerational correlation (less mobility )

Fig. 1. Genetic and environmental influences on educational attainment. (A) Trends in intergenerational mobility across 10 countries: Australia (AUS),Denmark (DNK), Finland (FIN), Germany (DEU), Italy (ITA), Norway (NOR), Spain (ESP), Sweden (SWE), United Kingdom (GBR), and United States (USA).(B and C) Association of intergenerational mobility with heritability (h2) and shared environmental influences (c2). Superimposed lines show the least-squaresline of best fit, with 95% confidence intervals indicated by shaded areas; marker labels encode the country and decade of birth for each cohort.

Nevertheless, educational attainment is an incomplete proxy forsocial standing, and our results might have looked different withaccess to comparable data on income, wealth, or occupationalprestige. Notably, research typically finds larger environmen-tal components for economic outcomes (19). The intergenera-tional correlation is also distinct from causal effects of parentaleducation, or the sibling correlation (20).

Lastly, it remains debatable whether genetic advantagesshould be seen as less troubling than socially inherited ones.While social inheritance may be easier to subvert, we should wantto compensate for genetic bad luck when we can (21). Researchusing molecular genetic data can shed further light on the mech-anisms underlying genetic influence (10, 22). Such research tellsus that individual genetic variation contributes to social mobil-ity and not just inheritance (23). More work in this vein willhelp grow the evidence base for policies aimed at amelioratingdisadvantage.

Materials and MethodsIntergenerational Mobility. Estimates are from the Global Database on Inter-generational Mobility (GDIM) (24) compiled by the World Bank (2) fromvarious sample surveys. The main respondent is the daughter or son whoalso reports about the parents. We use the correlation between child (yc)and parent (yp) years of schooling,

rIG = Corr(yc, yp) =Cov(yc, yp)√

Var(yc)√

Var(yp), [1]

Table 1. Correlation of standardized and destandardizedvariance components with the intergenerational correlation

Standardized Destandardized

h2 c2 e2 h2′ c2′ e2′

Coefficient −0.512 0.574 −0.011 −0.056 0.726 0.342Robust SE 0.142 0.121 0.179 0.284 0.147 0.207t statistic −3.61 4.74 −0.06 −0.20 4.94 1.65P> |t| 0.003 0.000 0.951 0.848 0.000 0.121

The table shows correlations (standardized β) between rIG and standard-ized and destandardized versions of h2, c2, and e2, including a fixed effectfor gender. Standard errors are clustered for each birth cohort within acountry (15 clusters).

for the parent with highest education. We smooth country trends byaveraging the value for each cohort with the adjacent ones.

Behavior Genetics Estimates. Genetic variance components are from a meta-analysis by Branigan et al. (11). The h2 (heritability) and c2 (shared envi-ronmental variance) are estimated from the correlation in educationalattainment among identical (rMZ ) and fraternal (rDZ ) twin pairs,

h2= 2× (rMZ − rDZ ), [2]

c2= rMZ − h2, [3]

e2= 1− h2− c2, [4]

where e2 is a residual component reflecting nonshared environmentand measurement error. To compare absolute and relative variances, wedestandardize the components,

h2′= h2 ·Var(yc), [5]

and do likewise for c2 and e2. Total population variance Var(yc) is estimatedfrom the GDIM.

Twin studies make some strong model assumptions such as no geneticparental assortative mating, no gene–environment interaction, and that dif-ferent types of twins share environmental influences effectively to the sameextent. In our comparative study design, we mainly need to assume thatpotential model violations do not correlate with mobility patterns.

Quality Control and Matching. The GDIM spans cohorts born from the1940s through the 1980s; genetic variance components span cohorts bornthroughout the century. We match each twin estimate to the GDIM basedon country, gender, and the closest year of birth. Of the 34 estimates avail-able, we exclude 2 cohorts that lack overlap in the GDIM data, and a further3 that are from local or nonrepresentative samples in the United States. Thisleaves a total of 26 data points. The data contain a few negative varianceestimates, which we set to zero.

Data Availability. Data and code to replicate all findings can be found athttps://osf.io/c549j/. There we also show that our results are robust to ana-lytical decisions taken, including the choice of parent, whether to smoothover cohorts, whether to exclude nonrepresentative US samples, whetherto recode negative variance estimates, and how to account for clustering.Moreover, we confirm that our results are not driven by any one country, byrepeating analyses excluding each.

ACKNOWLEDGMENTS. P.E. was supported by Nuffield College, theLeverhulme Centre for Demographic Science, The Leverhulme Trust, andFORTE Swedish Research Council for Health, Working Life and Welfare

Engzell and Tropf PNAS | December 17, 2019 | vol. 116 | no. 51 | 25387

Dow

nloa

ded

by g

uest

on

Mar

ch 2

4, 2

020

(Grant #2016-07099). F.C.T. was supported by the Laboratory of Excel-lence in Economics and Decision Sciences (LabEx Ecodec) funded under theFrench National Research Agency (ANR) Investissements d’Avenir (Grant

#ANR-11-LABX-0047). We thank the World Bank and authors of the heri-tability metaanalysis (Amelia R. Branigan, Kenneth J. McCallum, and JeremyFreese) for making their estimates available.

1. R. Breen, J. O. Jonsson, Inequality of opportunity in comparative perspective: Recentresearch on educational attainment and social mobility. Annu. Rev. Sociol. 31, 223–243 (2005).

2. A. Narayan et al., Fair Progress?: Economic Mobility Across Generations Around theWorld (The World Bank, 2018).

3. C. Jencks, L. Tach, “Would equal opportunity mean more mobility?” in Mobilityand Inequality: Frontiers of Research from Sociology and Economics, S. L. Morgan,D. B. Grusky, G. S. Fields, Eds. (Stanford University Press, Stanford, CA, 2006), pp.23–58.

4. D. Conley et al., Is the effect of parental education on offspring biased or moderatedby genotype?Soc. Sci. 2, 82–105 (2015).

5. G. Guo, E. Stearns, The social influences on the realization of genetic potential forintellectual development. Soc. Forces 80, 881–910 (2002).

6. F. Nielsen, Achievement and ascription in educational attainment: Genetic andenvironmental influences on adolescent schooling. Soc. Forces 85, 193–216(2006).

7. R. Plomin, Blueprint: How DNA Makes Us Who We Are (MIT Press, 2018).8. E. Dal Bo, F. Finan, O. Folke, T. Persson, J. Rickne, Who becomes a politician?Q. J. Econ.

132, 1877–1914 (2017).9. A. Bell, R. Chetty, X. Jaravel, N. Petkova, J. Van Reenen, Who becomes an inventor in

America? The importance of exposure to innovation. Q. J. Econ. 134, 647–713 (2018).10. D. Cesarini, P. M. Visscher, Genetics and educational attainment. npj Sci. Learning 2,

4 (2017).11. A. R. Branigan, K. J. McCallum, J. Freese, Variation in the heritability of educational

attainment: An international meta-analysis. Soc. Forces 92, 109–140 (2013).12. R. Breen, R. Luijkx, W. Muller, R. Pollak, Nonpersistent inequality in educational

attainment: Evidence from eight European countries. Am. J. Sociol. 114, 1475–1521(2009).

13. R. Erikson, J. Goldthorpe, The Constant Flux: A Study of Class Mobility in IndustrialSocieties (Clarendon Press, Oxford, United Kingdom, 1992).

14. G. Clark, The Son Also Rises: Surnames and the History of Social Mobility (PrincetonUniversity Press, 2014).

15. S. Scarr-Salapatek, Race, social class, and IQ. Science 174, 1285–1295 (1971).16. D. C. Rowe, K. C. Jacobson, E. J. G. Van den Oord, Genetic and environmental influ-

ences on vocabulary IQ: Parental education level as moderator. Child Dev. 70,1151–1162 (1999).

17. E. M. Tucker-Drob, T. C. Bates, Large cross-national differences in gene ×socioeconomic status interaction on intelligence. Psychol. Sci. 27, 138–149(2016).

18. A. C. Heath et al., Education policy and the heritability of educational attainment.Nature 314, 734–736 (1985).

19. K. Majlesi, P. Lundborg, S. Black, P. Devereux, Poor little rich kids? The role of natureversus nurture in wealth and other economic outcomes and behaviors. Rev. Econ.Stud., 10.1093/restud/rdz038 (2019).

20. A. Bjorklund, K. G. Salvanes, “Education and family background: Mechanisms andpolicies” in Handbook of the Economics of Education, E. A. Hanushek, S. Machin,L. Woessmann, Eds. (Elsevier, 2011), vol. 3, pp. 201–247.

21. A. S. Goldberger, Heritability. Economica 46, 327–347 (1979).22. J. J. Lee et al., Gene discovery and polygenic prediction from a genome-wide asso-

ciation study of educational attainment in 1.1 million individuals. Nat. Genet. 50,1112–1121 (2018).

23. D. W. Belsky et al., Genetic analysis of social-class mobility in five longitudinal studies.Proc. Natl. Acad. Sci. U.S.A. 115, E7275–E7284 (2018).

24. Development Research Group, GDIM. Global Database on Intergenerational Mobility(World Bank, 2018). https://www.worldbank.org/en/topic/poverty/brief/what-is-the-global-database-on-intergenerational-mobility-gdim. Accessed 25 July 2018.

25388 | www.pnas.org/cgi/doi/10.1073/pnas.1912998116 Engzell and Tropf

Dow

nloa

ded

by g

uest

on

Mar

ch 2

4, 2

020