plant histology 2

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    Plant Histology II

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    Primary Growth

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    MERISTEMATIC TISSUE

    Meristems: - Permanent regions of active cell division- (undifferentiated plant tissue)

    = plant stem cells

    3 types of meristematic tissuesi. Apical

    ii. Lateraliii. intercalary

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    Apical Meristems

    Found at the tips of roots and shoots. Increase in length as the apical meristems

    produce new cells (primary growth).

    Primary Meristems

    ProtodermGround MeristemProcambium

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    Lateral Meristems

    Produce tissues that increase the girth ofroots and stems .

    Involved in Secondary Growth

    Cork Cambium - Lies outside vascularcambium just inside the outer bark

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    Intercalary meristems

    Develop at intervals along stemswhere they add to stem length.

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    TISSUES PRODUCED BYMERISTEMS

    Parenchyma Collenchyma Sclerenchyma

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    Cross Section of a Stained Leaf

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    PARENCHYMA

    Main type of cell making up the cortex All plant tissues contain parenchyma. Found in roots, stems and leaves. Normally they are unsepcialized Fx = act as a packing tissue which

    = can later be modified.Loosely packed (Called aerenchyma)

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    Parenchyma

    Normally alive in maturity. In leaves: the chloroplasts are found in

    parenchyma tissue (called clorenchyma) When the cells are turgid, they provide

    important support and shape to the plant.

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    COLLENCHYMA

    Similar to parenchymain shape

    more elongated Can also be thickened in

    corners of cell wall(picture right)

    Important in youngplants, leaves, and olderstems of non-woodyplants whereverflexibility and support isimportant.

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    COLLENCHYMA

    Contains living cytoplasm Fx = Provides flexible support for organs. NB in young plants (< cellulose) found

    beneath epidermis

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    Stained & unstained chollenchymacells

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    SCLENENCHYMA

    Cells with thick, tough, secondary walls Normally impregnated with lignin.

    < Flexible & stronger than collenchyma Fx = mechanical (support ) & food storage.

    2 TYPES OF CELLS (based on cell-shape) Sclerids - Stone Cells Fibers - Contain Lumen

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    Scleroids

    Found in nut shells

    the hard part of seeds flexible floating leafblades of water plants.

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    Stone cells in pear fruit

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    Sclerenchyma fibres

    Elongated & thick-walled withflattened ends.

    E.g. Fibres from flaxand hemp are usedto make fabric andrope.

    The fibres also storefood like starch forthe plant

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    Sclerenchyma: Astroscleroidal & Fibre Cells

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    PLANT CONDUCTING TISSUE

    2 MAIN TYPES OF CELLS Xylem Phloem

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    Xylem & Phloem Cells

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    XYLEM

    = Chief conducting tissue for water and minerals absorbed by the roots.

    3 TYPES OF XYLEM TISSUE Vessels - Made of vessel elements.

    Long tubes open at each end. Tracheids - Tapered at the ends with pits Allows for water to pass between cells. Rays - Lateral conduction of water and minerals.

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    Xylem Vessels

    Xylem Tracheid Xylem Vessel Element

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    Xylem Microscopy

    Spiral thickenings in the secondary walls of vessels and tracheids givesthem the appearance of microscopic coils under high magnification with alight microscope.

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    Vascular Conducting Cells

    Conducting cells of the xylem; tracheids (left)are more primitive, while the various types ofvessels (the other three) are more advanced.

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    PHLOEM

    Fx = Conducts food materials produced byphotosynthesis throughout the plant.

    Phloem cells as seen inlongitudinal section.Note the longitudinalview of the sieve plateinside the large sievetube cell

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    4 TYPES OF PHLOEM CELLS

    1. Sieve Tube Members (makes up the sieve-tube functional unit)

    Large, cylindrical Sieve Plates - Porous region

    2. Companion Cells Provide the energy needed for the long distance

    transport of food.

    Also help determine the direction the foodshould be transported in. Narrow, tapered

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    Phloem Cells

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    3. Parenchyma Acts mainly as a storage place for food. They store many secondary metabolites

    such as tannins, starch, crystals, etc.

    4. Sclerenchyma

    Fibres are very common in phloem

    (scleroids less common)

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    Longitudinal View of Phloem Cells

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    Microscopy: Phloem Cells

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    OTHER PLANT TISSUES

    Neither vascular nor meristematic

    2 TYPES Epidermis Periderm

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    EPIDERMIS = Outermost layer of cells. (One cell-layer thick)

    * Most secrete fatty substance (cutin) on thesurface of the outer walls.Forms cuticle.

    Leave epidermal cells have stomata bordered bypairs of guard cells

    Root epidermal cells produce root hairs

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    EPIDERMIS

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    PERIDERM= Outer bark

    Primarily composed of cork cells.

    Lenticles: Living tissues under the cork layer(need oxygen) the periderm therefore possessesspecialized pores for gaseous exchange calledlenticles, made up of loosely packed parachymacells with large air spaces. They protrude throughthe epidermis, and allows for adequate gaseousexchange.

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    PERIDERM