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    FAO SmartFish

    Publication

    PRELIMINARY RESULTS OF THE

    HYDROACOUSTIC SURVEY CONDUCTED

    ON LAKE KARIBASeptember 2014

    33

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    Cover photograph courtesy FAO SmartFish

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    REPORT/RAPPORT: SF-FAO/2014/33

    Preliminary Results of theHydroacoustic Survey conducted

    on Lake Kariba

    September 2014 

    GCP/RAF/466/EC SmartFish Project

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      3

    Executive Summary

    As a continuation of a hydroacoustic training in 2013 supported by SmartFish FAO, and in

    accordance with recommendations of the Fifth Technical Consultation on Development

    and Management of the Fisheries of Lake Kariba, SmartFish FAO supported a

    hydroacoustic survey in August 2014.

    The preliminary results of the survey that was undertaken to determine the standing

    biomass of Kapenta in Lake Kariba show that by adding the estimated biomasses of the

    basins and stratum, the estimated average standing biomass was 16,277 i.e. ± 9,730

    tonnes.

    In order to ascertain any changes over time and to draw substantial conclusions,

    comparisons across time is necessary. A regular hydroacoustic survey programme will

    have to be established and the relationship between densities and environmentalparameters should be assessed.

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     4

    Résumé exécutif

    Suite à la formation en hydroacoustique qui a été organisée en 2013 grâce au support de

    SmartFish FAO, et conformément aux recommandations de la Cinquième Consultation

    sur le Développement et la Gestion du Lac Kariba, SmartFish FAO a supporté laréalisation d’un relevé hydroacoustique en aout 2014.

    Ce relevé a été réalisé pour déterminer la biomasse du kapenta dans le Lac Kariba.

    Les résultats préliminaires montrent qu’en additionnant les relevés des biomasses des

    bassins et couches, la moyenne estimée de biomasse est de 16 277 c’est-à-dire +/- 9

    730 tonnes.

    Pour vérifier les changements dans le temps et tirer des conclusions fiables, il est

    nécessaire d’effectuer des comparaisons dans le temps. Un programme de relevés

    hydroacoustiques réguliers devra être établi et les relations entre les densités et les

    paramètres environnementaux devront être établies.

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      5

    Contents 

    Executive Summary ______________________________________________________________ 3

    Résumé exécutif _________________________________________________________________ 4

    Contents ________________________________________________________________________ 5

    List of Tables _____________________________________________________________________ 6

    List of Figures ____________________________________________________________________ 6

    List of Photos ____________________________________________________________________ 6

    1. Background___________________________________________________________________ 7 

    2. Material and methods _________________________________________________________ 7 

    2.1 Description of the study area _______________________________________________ 7

    2.2 Methods __________________________________________________________________ 82.3 Data analysis ____________________________________________________________ 11

    3. Results _______________________________________________________________________ 11 

    3.1 Average length and weight of Kapenta along the lake ____________________ 11

    3.2 Fish densities _____________________________________________________________ 11

    3.2.1 Zimbabwean side of Lake Kariba _______________________________________ 11

    3.2.2 Zambian side of Lake Kariba ____________________________________________ 13

    3.3 Fish biomass _____________________________________________________________ 13

    4. Conclusion and recommendations ____________________________________________ 15 

    Annex 1. List of participants and survey agenda __________________________________ 16 

    Annex 2. Pictures from the hydroacoustic survey __________________________________ 17 

    Annex 3. Data used to derive densities ___________________________________________ 24 

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     6

    List of Tables

    Table 1: Average length and weight of Kapenta along Lake Kariba ______________ 12

    List of Figures

    Figure 1: Location of Lake Kariba in Africa ________________________________________  8

    Figure 2: Research vessel, Namazambwe ________________________________________ 9

    Figure 3: The pre-defined sampling transects and basin boundaries _______________  10

    Figure 4: Trip track log __________________________________________________________  10

    Figure 5: Average Kapenta densities on the Zimbabwean side of Lake Kariba _____  12

    Figure 6: Correlation between Kapenta density and sampled depth in

    Leisure Bay (un-fished area) ____________________________________________________  12Figure 7: Average Kapenta densities on the Zambia side of Lake Kariba __________  13

    Figure 8: Average standing biomass per basin on the Zimbabwean

    side of Lake Kariba. ____________________________________________________________  14

    Figure 9: Average standing biomass per basin on the Zambian side of Lake Kariba 14

    List of Photos

    Photo 1: Team photo taken during the official launch of the

    hydroacoustic survey held in the Sinasongwe District Council Hall ------------------------- 17

    Photo 2: The team members during preparations for the survey ---------------------------- 17

    Photo 3: Some members of the team monitoring the screen during calibration ------- 18

    Photo 4: Team members setting the sphere and the transducer during calibration -- 18

    Photo 5: View of the screen during calibration ---------------------------------------------------- 19

    Photo 6: Preparing the transducer --------------------------------------------------------------------- 19

    Photo 7: Team photo prior to departure from Sinazongwe for the survey ---------------- 20

    Photo 8: Launching CTD during the survey ---------------------------------------------------------- 20

    Photo 9: Kapenta fish sample taken from a Kapenta rig during the survey -------------- 21

    Photo 10: The biggest Kapenta collected during the survey ---------------------------------- 21

    Photo 11: Measuring and weighing Kapenta samples ------------------------------------------ 22

    Photo 12: The survey team just before disembarking “Namazambwe”

    at Siavonga, after the hydroacoustic survey ------------------------------------------------------- 22

    Photo 13: Data analysis being carried out by the survey team ------------------------------ 23

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 7

    1. Background

    Lake Kariba is jointly owned by the governments of Zambia in the north and Zimbabwe in

    the south. It lies between latitudes 16°28´ and 18°06´ south and longitude 26°40´ and

    29°03´ east. The full operation of the reservoir was reached in 1963. Although the lakewas created as a hydroelectric reservoir, there are accounts of a considerable number of

    activities taking place both in the lake and along its shores, including capture fisheries and

    aquaculture.

    In order to implement one of the recommendations of the Fifth Technical Consultation on

    Development and Management of the Fisheries of Lake Kariba, a hydroacoustic survey was

    conducted in Lake Kariba from 14 to 24 August 2014. The survey was conducted as a

    continuation of the training conducted in Songo in 2013 and, for this reason, involved all

    the personnel that attended this course, i.e: two officers from Zambia; two from

    Zimbabwe; two from Malawi and two from Mozambique. The survey was co-funded by FAOand SmartFish. The survey agenda and the list of those involved is given in Annex 1.

    The main objective of the hydroacoustic survey was to determine the standing biomass of

    Kapenta in Lake Kariba, given that the last hydroacoustic survey on Lake Kariba was

    carried out in 1997. Determination of the current biomass is thought to provide useful data

    for the management of the Kapenta fishery.

    In order to fulfill the aforementioned objective, the following activities were implemented:

    ♥ Calibration of the EK-60 split beam echo-sounder;

    ♥ Sampling with EK-60 along pre-established zigzag transects;

    ♥ Sampling of Kapenta from Kapenta fishing rigs;

    ♥ Sampling with CTD along the transects and;

    ♥ Data analysis.

    2. Material and methods

    2.1 Description of the study area

    Lake Kariba is a man-made lake which lies between latitudes 16°28´ and 18°06´ south

    and longitude 26°40´ and 29°03´ east. Figure 1 below, shows the geographic location of

    the lake.

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     Material and methods8

    Figure 1: Location of Lake Kariba in Africa

    Overlaid on NDVI by Henry Ndaimani

    The principal physical characteristic of the lake at the normal operating level of 485m

    above mean sea level can be summarized as follows:

    Catchment area 409,600 km²

    Surface area 5,364 km²

    Maximum length 320 km

    Maximum depth 120 m

    Maximum width 40 km

    For management purposes, the lake is divided into five basins (Mlibizi, Binga, Sengwa,

    Bumi/Chalala and Sanyati) on the Zimbabwean side and four strata numbered from 1 to 4,

    west-easterly, on the Zambian side (Figure 3).

    2.2  Methods

    The survey was conducted at night, on board the Department of Fisheries of Zambia ’s

    research vessel, “Namazambwe” (Figure 2). During sampling with EK-60, the transducer

    was towed alongside the vessel, which moved at an average speed of 7 knots.

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 9

    Figure 2: Research vessel, Namazambwe

    Calibration was done at the bay, close to the harbour. The actual sampling began where

    the first transect starts, close to the town of Binga. Figures 3 and 4 show the pre-definedtransects and the trip log.

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     Material and methods10

    Figure 3: The pre-defined sampling transects and basin boundaries

    Figure 4: Trip track log

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 11

    CTD casts and fish samples were taken along the transects. CTD provides profile readings

    of environmental parameters such as temperature, conductivity and depth, as well as

    dissolved oxygen and chlorophyll. Fish samples were taken to derive the average weight of

    fish in order to determine the biomass.

    2.3  Data analysis

    Sampled data from EK-60 were converted and analyzed using data analysis software

     “SONAR 4”. Converted samples were analyzed for fish densities per transect and late

    average densities per basin were calculated using Excel. The results of data sampled in the

    different transects are presented in Annex 3.

    In order to derive the average length and weight, weighted length and weight were

    calculated per basin.

    3. Results

    3.1  Average length and weight of Kapenta along the lake

    Both the average length and weight of Kapenta differ across basins and across stratum

    (Table 1). Bigger individuals are found at the Sanyati basin, probably (possibly) as a result

    of a spillover from the Leisure Bay, which is an un-fished area.

    Table 1: Average length and weight of Kapenta along Lake Kariba

    Basin/Stratum Average fishweight (g) Average fishlength (mm)

    Mlibizi

    Binga 1.06 52.23

    Sengwa 1.01 50.76

    Bumi-Chalala 0.85 49.79

    Sanyati 1.14 50.72

    Stratum 1 1.06 52.23

    Stratum 2 1 49.77

    Stratum 3 0.97 51.44

    Stratum 4 1.14 50.72

    3.2  Fish densities

    3.2.1 Zimbabwean side of Lake Kariba 

    Fish densities for the Zimbabwean side of Lake Kariba are presented in Figure 5. The

    highest densities were observed in Leisure Bay at an average of 50,000 fish/ha. However,

    it should be noted that there was a significant variation of Kapenta fish densities at

    different sampled points along the transects, which is consistent with an uneven

    distribution of Kapenta along the lake.

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     Material and methods12

    In fact, it was observed that, for the un-fished area, there was a strong correlation

    between fish density and sampled depth (Figure 6). This scenario was not observed in

    fished areas, which would suggest that other factors such as fishing are affecting fish

    densities.

    Figure 5: Average Kapenta densities on the Zimbabwean side of Lake Kariba

    Figure 6: Correlation between Kapenta density and sampled depth in Leisure Bay

    (un-fished area)

    0

    10,000

    20,000

    30,000

    40,000

    50,000

    60,000

    70,000

    80,000

    90,000

        M    l   i    b   i   z   i

        B   i   n   g   a

        S   e   n   g   w   a

        B   u   m

       i  -    C    h   a    l   a    l   a

        S   a   n   y   a   t   i

        L   a   i   s   u   r   e    b   a   y

        K   a   p   e   n   t   a    d   e   n   s    i

       t   y    (    f    /    h   a    )

    Basin

    Kapenta fish density on the Zimbabwe side of Lake

    Kariba

    y = -1109.8x + 111644R² = 0.9252

    0

    20,000

    40,000

    60,000

    80,000

    100,000

    120,000

    0 20 40 60 80 100 120

        K   a   p   e   n   t   a    d   e   n   s    i   t   y    (    f    /    h   a    )

    Sampled depth (m)

    Correlation between depth and fish density at Leisure

    bay

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 13

    3.2.2 Zambian side of Lake Kariba 

    On the Zambian side of Lake Kariba, the Kapenta densities seemed highest on the first

    stratum and lowest on the third. These results are somehow comparable with those of the

    Zimbabwean side of the lake, with the exception of the 4 th  stratum, which presents

    relatively lower densities than its closest counterpart, the Sanyati basin.

    Figure 7: Average Kapenta densities on the Zambia side of Lake Kariba

    3.3  Fish biomass

    The average Kapenta biomasses for the different basins on both sides of Lake Kariba are

    presented in Figure 8 and Figure 9. Just as for the fish densities, the standing biomasses

    vary across the basins and are highest in the Sanyati basin on the Zimbabwean side of the

    lake.

    0

    10,000

    20,000

    30,000

    40,000

    50,000

    60,000

    70,000

    Stratum 1 Stratum 2 Stratum 3 Stratum 4

        K   a   p   e   n   t   a    d   e   n   s    i   t   y    (    f    /

        h   a    )

    Stratum

    Kapenta densities on the Zambia side of lake Kariba

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     Results14

    Figure 8: Average standing biomass per basin on the Zimbabwean side of Lake

    Kariba.

    Figure 9: Average standing biomass per basin on the Zambian side of Lake Kariba

    0

    1000

    2000

    3000

    4000

    5000

    6000

    7000

        K   a   p   e   n   t   a   s   t   a

       n    d    i   n   g    b    i   o   m   a   s   s    (   t   o   n   n   e   s    )

    Basin

    Kapenta standing biomass on the Zimbabwe side oflake kariba

    0

    500

    1000

    1500

    2000

    2500

    3000

    3500

    4000

    4500

    Stratum 1 Stratum 2 Stratum 3 Stratum 4

        K   a   p   e   n   t   a   s   t   a   n    d    i   n   g    B    i   o   m   a   s   s    (   t   o   n   n   e   s    )

    Stratum

    Kapenta standing biomass on the Zambia side of Lake

    Kariba

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 15

    Total standing biomass

    By adding the estimated biomasses of the basins and stratum, the estimated average

    standing biomass was 16,277 i.e. ± 9,730 tonnes.

    4. Conclusion and recommendations

    Comparisons between the results reported in this study and those from previous surveys

    are necessary in order to ascertain any changes over time.

    Further analysis on the samples collated is required in order to assess densities/depth

    variation along the transects.

    The relationship between densities and environmental parameters should be assessed.

    Fish densities versus distribution of Kapenta fishing efforts should be determined andmapped.

    A regular hydroacoustic survey programme will have to be established.

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     Annexes16

    Annex 1. List of participants and survey agenda

    List of participants

    Name Country

    Jorge Mafuca  Mozambique 

    Antonia Manica da Costa  Mozambique 

    Daniel Mueleque Mozambique 

    Emmanuel Silwimba  Zambia 

    Taylor Banda Zambia 

    Mebelo Wamulume  Zambia 

    Itai Tendaupenyu Zimbabwe 

    Michael Tiki Zimbabwe 

    Stanley Mvula Malawi Innocent Gumurila Malawi 

    Nobuhle Ndhlovu FAO 

    Drivers (2) FAO 

    Boat crew (3) Zambia 

    Survey agenda

    Day Date Activity

    1 10/08/2014 Arrival of participants in Harare

    2 14/08/2014 Departure to Sinazongwe, Zambia via Kariba (to pick

    up Kariba and Siavonga participants)

    3 15/08/2014 Official opening

    Recap. for participants

    4 16/08/2014 Calibration of equipment

    5 17/08/2014 Calibration of equipment

    6 18/08/2014 Night survey

    7 19/08/2014 Night survey

    8 20/08/2014 Night survey

    21/08/2014 Night survey

    22/08/2014 Night survey

    23/08/2014 Arrival in Siavonga, disembarcation from boat and

    departure for Kariba, Zimbabwe.

    9 24/08/2014 Data analysis at Lake Kariba Fisheries Research

    Institute

    10 25/08/2014 Departure for Harare and participants from Zambia

    travel back to Zambia

    11 26/08/2014 Participants travel back to respective countries

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 17

    Annex 2. Pictures from the hydroacoustic survey

    Photo 1: Team photo taken during the official launch of the hydroacoustic survey

    held in the Sinasongwe District Council Hall

    Photo 2: The team members during preparations for the survey

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     Annexes18

    Photo 3: Some members of the team monitoring the screen during calibration

    Photo 4: Team members setting the sphere and the transducer during calibration

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 19

    Photo 5: View of the screen during calibration

    Photo 6: Preparing the transducer

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     Annexes20

    Photo 7: Team photo prior to departure from Sinazongwe for the survey

    Photo 8: Launching CTD during the survey

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 21

    Photo 9: Kapenta fish sample taken from a Kapenta rig during the survey

    Photo 10: The biggest Kapenta collected during the survey

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     Annexes22

    Photo 11: Measuring and weighing Kapenta samples

    Photo 12: The survey team just before disembarking “Namazambwe” at

    Siavonga, after the hydroacoustic survey

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 23

    Photo 13: Data analysis being carried out by the survey team

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     Annexes24

    Annex 3. Data used to derive densities

    Transect Basin StratumSample

    NumberDepth Total f/ha

    1 Binga 1 1 37.95 62,911

    1 Binga 1 2 27.7 63,165

    1 Binga 1 3 17.17 2,589

    1 Binga 1 4 12.74 8,523

    2 Binga 1 1 31.86 6,410

    2 Binga 1 2 44.6 70,144

    2 Binga 1 3 36.84 15,534

    2 Binga 1 4 40.17 58,346

    2 Binga 1 5 55.68 80,577

    2 Binga 1 6 42.94 49,058

    2 Binga 1 7 38.78 26,685

    2 Binga 1 8 32.41 45,652

    3 Binga 1 1 32.69 32,396

    3 Binga 1 2 45.43 29,552

    3 Binga 1 3 42.94 39,131

    3 Binga 1 4 40.17 48,515

    3 Binga 1 5 31.30 11,560

    3 Binga 1 6 28.53 32,602

    3 Binga 1 7 26.59 19,7073 Binga 1 8 18.00 22,351

    3 Binga 1 9 24.65 31,071

    3 Binga 1 10 21.61 48,450

    4 Binga 2 1 22.44 35,981

    4 Binga 2 2 19.94 28,229

    4 Binga 2 3 19.11 11,079

    4 Binga 2 5 23.55 11,232

    4 Binga 2 6 18.56 2,662

    4 Binga 2 7 27.15 19,327

    4 Binga 2 8 23.55 10,406

    5 Sengwa 2 1 26.04 253

    5 Sengwa 2 2 24.1 7,408

    5 Sengwa 2 3 32.69 31,087

    5 Sengwa 2 4 40.72 12,889

    5 Sengwa 2 5 37.95 8,003

    5 Sengwa 2 6 31.58 43,511

    5 Sengwa 2 7 23.27 26,204

    6 Sengwa 2 1 23.55 14,794

    6 Sengwa 2 2 40.04 79,8776 Sengwa 2 3 51.25 42,046

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 25

    Transect Basin StratumSample

    NumberDepth Total f/ha

    6 Sengwa 2 4 65.93 2,691

    6 Sengwa 2 5 36.46 21,652

    7 Sengwa 2 1 56.01 7,177

    7 Sengwa 2 2 43.69 9,092

    7 Sengwa 2 3 49.31 22,747

    7 Sengwa 2 4 50.69 34,911

    7 Sengwa 2 5 67.31 17,411

    7 Sengwa 2 6 27.98 1,274

    8 Sengwa 2 1 31.03 49,374

    8 Sengwa 2 2 60.39 46,636

    8 Sengwa 2 3 53.74 35,055

    8 Sengwa 2 4 31.3 27,250

    8 Sengwa 2 1 19.94 27,9449 Sengwa 2 2 31.86 40,508

    9 Sengwa 2 3 42.11 10,522

    9 Sengwa 2 4 39.61 83,331

    9 Sengwa 2 5 50.97 92,396

    9 Sengwa 2 6 57.34 10,320

    9 Sengwa 2 7 53.14 24,202

    9 Sengwa 2 8 22.99 10,626

    9 Sengwa 2 9 24.93 6,225

    10 Sengwa 3 1 38.5 52,541

    10 Sengwa 3 2 48.75 58,574

    10 Sengwa 3 3 46.81 88,998

    10 Sengwa 3 4 59.83 33,237

    11 Sengwa 3 1 39.61 6,967

    11 Sengwa 3 2 46.26 10,623

    11 Sengwa 3 3 52.35 25,863

    11 Sengwa 3 4 31.58 7,863

    11 Sengwa 3 5 32.69 6,284

    11 Sengwa 3 6 56.23 31,636

    11 Sengwa 3 7 20.78 9,54211 Sengwa 3 8 25.76 37,385

    11 Sengwa 3 9 21.88 14,166

    12 Bumi-Chalala 3 1 24.38 12,831

    12 Bumi-Chalala 3 2 42.66 10,284

    12 Bumi-Chalala 3 4 60.94 13,836

    12 Bumi-Chalala 3 5 62.88 19,907

    12 Bumi-Chalala 3 6 53.35 22,767

    12 Bumi-Chalala 3 7 40.17 13,713

    12 Bumi-Chalala 3 8 32.96 8,475

    12 Bumi-Chalala 3 9 39.9 6,486

    12 Bumi-Chalala 3 10 21.88 12,840

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     Annexes26

    Transect Basin StratumSample

    NumberDepth Total f/ha

    12 Bumi-Chalala 3 11 17.17 12,540

    12 Bumi-Chalala 3 12 15.79 9,000

    12 Bumi-Chalala 3 13 13.85 41,560

    12 Bumi-Chalala 3 14 30.47 18,000

    12 Bumi-Chalala 3 15 41.27 8,013

    12 Bumi-Chalala 3 16 26.87 8,146

    12 Bumi-Chalala 3 17 65.37 21,562

    12 Bumi-Chalala 3 18 39.34 26,136

    13Tashinga/SambaKaluma 4 1 41.27 21,146

    13Tashinga/SambaKaluma 4 2 64.82 38,896

    13

    Tashinga/Samba

    Kaluma 4 3 85.04 27,428

    13Tashinga/SambaKaluma 4 4 91.69 45,232

    13Tashinga/SambaKaluma 4 5 80.61 7,539

    13Tashinga/SambaKaluma 4 6 81.72 10,622

    14 Sanyati 1 65.10 38,900

    14 Sanyati 2 55.12 45,048

    14 Sanyati 3 27.42 43,301

    14 Sanyati 4 30.75 48,811

    14 Sanyati 5 51.25 64,80314 Sanyati 6 46.54 60,275

    14 Sanyati 7 52.63 38,168

    14 Sanyati 8 23.55 68,756

    14 Sanyati 9 23.82 56,231

    15 Sanyati 1 26.04 60,050

    15 Sanyati 2 33.52 34,891

    15 Sanyati 3 34.09 58,704

    15 Sanyati 4 37.95 32,025

    15 Sanyati 5 61.50 31,964

    15 Sanyati 6 66.48 35,280

    15 Sanyati 7 66.48 36,247

    15 Sanyati 8 32.13 69,106

    15 Sanyati 9 71.47 60,538

    16 Sanyati 1 60.66 41,794

    16 Sanyati 2 54.45 42,988

    16 Sanyati 3 50.14 40,477

    16 Sanyati 4 52.08 53,436

    16 Sanyati 5 25.21 64,130

    16 Sanyati 6 36.01 51,29317 Leisure Bay 1 15.51 87,234

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     Preliminary results of the hydroacoustic survey conducted on Lake Kariba 27

    Transect Basin StratumSample

    NumberDepth Total f/ha

    17 Leisure Bay 2 19.39 98,747

    17 Leisure Bay 3 23.27 98,468

    17 Leisure Bay 4 35.73 68,499

    17 Leisure Bay 5 57.06 39,552

    17 Leisure Bay 6 63.16 34,746

    17 Leisure Bay 7 71.47 27,211

    17 Leisure Bay 8 81.44 19,940

    17 Leisure Bay 9 77.84 20,413

    17 Leisure Bay 10 95.57 21,859

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    SmartFish is a regional fisheries project managed by the Indian Ocean

    Commission, funded by the European Union and co-implemented by theFood and Agriculture Organization of the United Nations. SmartFish, which

    operates in 20 countries throughout the East and Southern Africa - Indian

    Ocean region, focuses on fisheries governance, management, monitoring,

    control and surveillance, trade, and food security.

    As a continuation of a hydroacoustic training in 2013 and in accordance

    with recommendations of the Fifth Technical Consultation on Development

    and Management of the Fisheries of Lake Kariba, SmartFish supported a

    hydroacoustic survey in August 2014. This report documents the

    preliminary results of the survey that was undertaken to determine the

    standing biomass of Kapenta in Lake Kariba.