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Plant Organs: Leaves
Chapter 8
LEARNING OBJECTIVE 1
Describe the major tissues of the leaf (epidermis, mesophyll, xylem, and phloem)
Relate the structure of the leaf to its function of photosynthesis
“Typical” Leaf
Blade
Veins
Petiole
Stipules
Axillary bud
Stem
Fig. 8-1, p. 152
KEY TERMS
BLADE
Broad, flat part of a leaf
PETIOLE
Part of a leaf that attaches blade to stem
Leaf Morphology
Parallel
Pinnately
netted
Palmately netted
Bermuda grass
(Cynodon dactylon) Black willow
(Salix nigra)
Sweetgum
(Liquidambar
styraciflua)
(c) Venation patterns.
Stepped Art
Alternate Opposite
Whorled
American beech
(Fagus grandifolia) Sugar maple
(Acer saccharum) Southern catalpa
(Catalpa bignonioides) (b) Leaf arrangement on a stem.
Simple Pinnately compound
Palmately
compound
California white oak
(Quercus lobata) White ash
(Fraxinus americana)
Ohio buckeye
(Aesculus
glabra) (a) Leaf form: simple and compound.
Fig. 8-2, p. 154
KEY TERMS
PHOTOSYNTHESIS
The biological process that includes the capture of light energy and its transformation into chemical energy of organic molecules (such as glucose), which are manufactured from carbon dioxide and water
Tissues in a Leaf Blade
Animation: Leaf Organization
Epidermis
The transparent epidermis allows light to penetrate into the mesophyll, where photosynthesis occurs
KEY TERMS
CUTICLE
Waxy covering over epidermis of aerial parts (leaves and stems) of a plant
Enables the plant to survive in the dry conditions of a terrestrial environment
Trichomes
KEY TERMS
STOMA
Small pores in epidermis of stem or leaf
Permit gas exchange for photosynthesis and transpiration
Flanked by guard cells
GUARD CELL
Two guard cells form a pore (stoma)
Stomata
Stomata typically open during the day, when photosynthesis takes place, and close at night
KEY TERMS
MESOPHYLL
Photosynthetic ground tissue in the interior of a leaf
Contains air spaces for rapid diffusion of carbon dioxide and water into, and oxygen out of, mesophyll cells
Vascular Bundle
Leaf veins have
xylem to conduct water and essential minerals to the leaf
phloem to conduct sugar produced by photosynthesis to rest of plant
KEY TERMS
BUNDLE SHEATH
One or more layers of nonvascular cells (parenchyma or sclerenchyma) surrounding the vascular bundle in a leaf
LEARNING OBJECTIVE 2
Contrast leaf structure in eudicots and monocots
Bundle Sheath Extensions
Lower epidermis
Bundle sheath
extension
Midvein
Bundle sheath
Bundle sheath
extension
Upper epidermis
Fig. 8-5, p. 157
Leaf Cross Sections
Leaf Cross Sections
(a) Privet (Ligustrum vulgare), a eudicot, has a mesophyll with
distinct palisade and spongy sections.
Phloem Xylem Stoma
Lower epidermis
Air space
Spongy mesophyll
Lengthwise view of vein
Palisade mesophyll
Upper epidermis
Midvein
Privet
Fig. 8-6a, p. 158
Parallel vein
Midvein Bundle sheath cells
Mesophyll
Upper
epidermis
Lower
epidermis
Xylem Phloem
Fig. 8-6b, p. 158
Monocot and Eudicot Leaves
Monocot leaves
Usually narrow
Wrap around the stem in a sheath
Have parallel venation
Eudicot leaves
Usually have a broad, flattened blade
Have netted venation
Bulliform Cells
Large, thin-walled cells on upper epidermises of leaves of certain monocots (grasses)
Located on both sides of the midvein
May help leaf roll or fold inward during drought
Bulliform Cells
Midvein
Bulliform
cells
(a) A folded leaf blade.
The inconspicuous
bulliform cells occur in
the upper epidermis
on both sides of the
midvein.
Fig. 8-7a, p. 159
(b) An expanded leaf
blade. A higher
magnification of the
midvein region shows
the enlarged, turgid
bulliform cells.
Midvein
Mesophyll
cell
Bulliform
cells
Fig. 8-7b, p. 159
LEARNING OBJECTIVE 3
Outline the physiological changes that accompany stomatal opening and closing
Variation in Guard Cells
(a) Guard cells of eudicots and many monocots
are bean shaped.
Subsidiary
cells
Closed Open
Guard
cells
Fig. 8-8a, p. 160
Subsidiary
cells
Closed Open
Guard
cells
(b) Some monocot guard cells (those of grasses, reeds, and
sedges) are narrow in the center and thicker at each end.
Fig. 8-8b, p. 160
Fig. 8-8d, p. 160
Stomatal Opening 1
1. Blue light activates proton pumps
in guard-cell plasma membrane
2. Protons (H+) are pumped out of guard cells, forming a proton gradient
Charge and concentration difference on two sides of the guard-cell plasma membrane
KEY TERMS
PROTON GRADIENT
Difference in concentration of protons on the two sides of a cell membrane
Contains potential energy that can be used to form ATP or do work in the cell
Stomatal Opening 2
3. Gradient drives facilitated diffusion of potassium ions into guard cells
4. Chloride ions also enter guard cells through ion channels
Ions accumulate in vacuoles of guard cells
Solute concentration becomes greater than that of surrounding cells
KEY TERMS
FACILITATED DIFFUSION
Diffusion of materials from a region of higher concentration to a region of lower concentration through special passageways in the membrane
Stomatal Opening 3
5. Water enters guard cells from surrounding epidermal cells by osmosis
Increased turgidity changes the shape of guard cells, causing stoma to open
Stomatal Opening
Blue light
activates
proton
pumps.
Protons are
pumped
out of guard
cells,
forming proton
gradient.
Potassium
ions enter
guard cells
through
voltage-
activated ion
channels.
Chloride ions
also enter guard
cells through
ion channels.
Water enters
guard cells by
osmosis,and
stoma opens.
1 4 5 3 2
Fig. 8-9, p. 162
Stomatal Closing
As evening approaches, sucrose concentration in guard cells declines
Sucrose is converted to starch (osmotically inactive)
Water leaves by osmosis, guard cells lose their turgidity, pore closes
Adaptations to Environment
Blade Petiole
Fig. 8-10, p. 163
Mesophyll cell
(photosynthetic
parenchyma cell)
Vascular
bundle Phloem
Xylem
Endodermis
Resin duct
Epidermis and cuticle
Guard cells of
sunken stoma
Fig. 8-11, p. 164
LEARNING OBJECTIVE 4
Discuss transpiration and its effects on the plant
KEY TERMS
TRANSPIRATION
Loss of water vapor from a plant’s aerial parts
Transpiration
Occurs primarily through stomata
Rate of transpiration is affected by environmental factors
temperature, wind, relative humidity
Both beneficial and harmful to the plant
Transpiration
75% Water recycled by transpiration
and evaporation
25% Water seeps into ground or runs off
to rivers, streams, and lakes
p. 165
Wilting
Guttation
LEARNING OBJECTIVE 5
Define leaf abscission
Explain why it occurs and what physiological and anatomical changes precede it
KEY TERMS
ABSCISSION
Normal (usually seasonal) falling off of leaves or other plant parts, such as fruits or flowers
Leaf Abscission
In temperate climates, most woody plants with broad leaves shed leaves in fall
Helps them survive low temperatures of winter
Involves physiological and anatomical changes
Processes of Abscission 1
As autumn approaches, plant reabsorbs sugar
essential minerals are transported out of leaves
Chlorophyll is broken down
red water-soluble pigments are synthesized and stored in vacuoles of leaf cells (in some species)
Processes of Abscission 2
A protective layer of cork cells develops on the stem side of the abscission zone
Area where leaf petiole detaches from stem, composed primarily of thin-walled parenchyma cells
Processes of Abscission 3
Enzymes dissolve middle lamella in abscission zone
(“cement” that holds primary cell walls of
adjacent cells together)
After leaf detaches, protective layer of cork seals off the area, forming a leaf scar
Abscission Zone
Stem
Abscission
zone
Petiole
Bud scales
Axillary bud
Fig. 8-14, p. 167
LEARNING OBJECTIVE 6
List at least five modified leaves, and give the function of each
KEY TERMS
BUD SCALE
Modified leaf that covers and protects delicate meristematic tissue of winter buds
SPINE
Leaf modified for protection, such as a cactus spine
KEY TERMS
BRACT
Modified leaf associated with a flower or inflorescence but not part of the flower itself
TENDRIL
Leaf or stem that is modified for holding on or attaching to objects
Supports weak stems
KEY TERMS
BULB
A rounded, fleshy, underground bud that consists of a short stem with fleshy leaves
Specialized for storage
Leaf Modifications
Fig. 8-15a, p. 168
Fig. 8-15b, p. 168
Fig. 8-15c, p. 168
Fig. 8-15d, p. 168
Fig. 8-15e, p. 168
Fig. 8-15f, p. 168
Epiphytes
Flowerpot Plant
(a) The leaves of the flowerpot plant (Dischidia
rafflesiana) are modified to hold water and organic
material carried in by ants.
Pot (modified
leaf)
Stem
Fig. 8-16a, p. 169
(b) A cutaway view of a pot removed from
a plant reveals the special root that absorbs
water and dissolved minerals inside the pot.
Root
Fig. 8-16b, p. 169
Carnivorous Plants
Leaves modified to trap insects