tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest...

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tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 30 25 20 15 10 5 0 -5 -10 -15 0 500 1000 1500 2000 2500 3000 3500 4000 4500 Annual Precipitation (mm) Annual Mean Temperature (C) temperate rain forest Figure 24.1 Ecological Climatology © 2008 G. Bon

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Page 1: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

tundradesert

woodlandshrubland

grassland

boreal forest

temperateforest

tropicalseasonal

forest

tropicalrain forest

thorn scrubsavanna

30 25 20 15 10 5 0 -5 -10 -150

500

1000

1500

2000

2500

3000

3500

4000

4500

An

nu

al P

reci

pita

tion

(m

m)

Annual Mean Temperature (C)

temperaterain forest

Figure 24.1Ecological Climatology © 2008 G. Bonan

Page 2: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Ann

ual N

et R

adia

tion,

Rn

(W m

-2)

Radiative Dryness Index (Rn/P)

0.5 1.0 1.5 2.0 2.5 3.00.00

20

40

60

80

100

120

140

Tropical

Wet savanna

Desert

Savanna

Subtropical

Forest

Mid-latitudedeciduous

Needleleaf

Tund

raSteppe

Steppe, prairie Semidesert

Figure 24.2Ecological Climatology © 2008 G. Bonan

Page 3: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

125

62.5

250

500

1000

2000

4000

8000

Annual Precipitation (mm

)

32.00 16.00 8.00 4.00 2.00 1.00 0.50 0.25 0.125superarid perarid arid semiarid subhumid humid perhumid superhumid

Pot

entia

l Eva

potra

nspi

ratio

n R

atio

0.25

0.50

1.00

2.00

4.00

8.00

16.00

32.00

0.125

Ann

ual M

ean

Bio

tem

pera

ture

(o C

)

1.5

3

6

12

24

polar

subpolar

boreal

cooltemperate

warm temperateor subtropical

tropical

LatitudinalRegions

1.5

3

6

12

24

nival

alpine

subalpine

montane

lowermontane

AltitudinalRegions

drytundra

moisttundra

wettundra

raintundra

desertdry

scrubmoistforest

wetforest

rainforest

desert desertscrub

steppe moistforest

wetforest

rainforest

desert desertscrub

thornwoodland

dryforest

moistforest

wetforest

rainforest

desert desertscrub

thornwoodland

very dryforest

dryforest

moistforest

wetforest

rainforest

Figure 24.3Ecological Climatology © 2008 G. Bonan

Page 4: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

0

500

1000

1500

2000

2500

3000

-15 -10 -5 0 5 10 15 20 25 30

Annual Mean Temperature (C)

NP

P (

g m

-2 y

r-1)

0

500

1000

1500

2000

2500

3000

0 500 1000 1500 2000 2500 3000 3500 4000

Annual Precipitation (mm)

NP

P (

g m

-2 y

r-1)

0

500

1000

1500

2000

2500

3000

0 200 400 600 800 1000 1200 1400

Annual Evapotranspiration (mm)

NP

P (

g m

-2 y

r-1)

(1.315 0.119 )

3000

1TNPP

e

0.0006643000 1 PNPP e

101.66 1.66log10

AETNPP

Figure 24.4Ecological Climatology © 2008 G. Bonan

Page 5: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.5Ecological Climatology © 2008 G. Bonan

Page 6: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.6Ecological Climatology © 2008 G. Bonan

Page 7: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

1 2

Desert Grass Shrub Savanna

3 5 9

Temperate Forest Tropical Forest

0

200

400

800

1600

Min

imu

m P

reci

pita

tion

(m

m y

r-1)

Ve

ge

tatio

n H

eig

ht

Leaf Area Index (m2 m-2)

Figure 24.7Ecological Climatology © 2008 G. Bonan

Page 8: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Tundra30 cm: 93%

Deepest: 0.5 mRoot-to-Shoot: 6.6

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Temperate Grassland30 cm: 83%

Deepest: 2.6 mRoot-to-Shoot: 3.7

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Boreal Forest30 cm: 83%

Deepest: 2.0 mRoot-to-Shoot: 0.32

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Tropical Decid. Forest30 cm: 70%

Deepest: 3.7 mRoot-to-Shoot: 0.34

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Tropical Ever. Forest30 cm: 69%

Deepest: 7.3 mRoot-to-Shoot: 0.19

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Temp. Decid. Forest30 cm: 65%

Deepest: 2.9 mRoot-to-Shoot: 0.23

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Tropical Savanna30 cm: 57%

Deepest: 15.0 mRoot-to-Shoot: 0.7

Cumulative Root Fraction

So

il D

ep

th (

cm)

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Temp. Conif. Forest30 cm: 52%

Deepest: 3.9 mRoot-to-Shoot: 0.18

0

50

100

150

200

0.00 0.25 0.50 0.75 1.00

Desert30 cm: 53%

Deepest: 9.5 mRoot-to-Shoot: 0.7, 4.5

Figure 24.8Ecological Climatology © 2008 G. Bonan

Page 9: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

0

100

200

300

400

500

600

700

800

900

1000

0 5 10 15 20 25 30

Annual Mean Temperature (C)

Litt

erf

all

(g m

-2 y

r-1)

Broadleaf deciduous

Broadleaf evergreen

Needleleaf deciduous

Needleleaf evergreen

0

100

200

300

400

500

600

700

800

900

1000

0 500 1000 1500 2000 2500 3000

Annual Precipitation (mm)

Litt

erf

all

(g m

-2 y

r-1)

Broadleaf deciduous

Broadleaf evergreen

Needleleaf deciduous

Needleleaf evergreen

Figure 24.9Ecological Climatology © 2008 G. Bonan

Page 10: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

50

60

70

80

90

100

-10 -5 0 5 10 15 20 25 30

Annual Mean Temperature (C)

Mas

s R

emai

ning

(%

)

Desert

Grassland

Agriculture

Marsh

Tundra

Broadleaf forest

Conifer forest

Tropical forest50

60

70

80

90

100

0 1000 2000 3000 4000

Annual Precipitation (mm)

Mas

s R

emai

ning

(%

)

50

60

70

80

90

100

0 250 500 750 1000 1250 1500

Annual Evapotranspiration (mm)

Mas

s R

emai

ning

(%

)

Figure 24.10Ecological Climatology © 2008 G. Bonan

Page 11: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Fir

Sp

ruce

Pin

e

Oa

k

Ash

Hic

kory

Ho

rnb

ea

m

Boreal Forest Deciduous Forest

0 25 50 75 100

Pollen (%)

0

2

4

6

8

10

12

14

16

Th

ou

san

ds

Of Y

ea

rs B

efo

re P

rese

nt

Figure 24.11Ecological Climatology © 2008 G. Bonan

Page 12: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.12Ecological Climatology © 2008 G. Bonan

Page 13: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.13Ecological Climatology © 2008 G. Bonan

Page 14: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Ice

Forest Tundra

Tundra

Boreal Forest

Mixed Forest

Deciduous Forest

Aspen Parkland

Prairie

Southeast Forest

No Analog

No Data

18000 12000

9000 6000

3000 Modern

Figure 24.14Ecological Climatology © 2008 G. Bonan

Page 15: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Mean January Temperature (C)

Mean July Temperature (C)

Annual Precipitation (mm)

0-6

-18

-12

16

16

-18-12 -6

0

-12 -18-12

-60

66

24

20

-12

-60

-12

-12-18

-12

-6

06

-24-30 -18

-18-12

-60 6

-24-30

-18-12

-606

-24

1212

812

16

24

20

812

16

24

20

28

1281216

24

20

28

1216

24

20

20

16

1620

24

28

16

24 20

28

600

1000

400

800

2424

-12 -6-12

600

12001000800

600

1200

1400

1000

800

800

600

12001000800

800

800

600800

800

600

1200

1400

1000

400

800

600

1200

1400

1000

400

800600 1200

14001000

400

800

800

1000

600

1000

1000

600

1000

800

600

18 000 15 000 12 000 9000 6000 3000 Present

Figure 24.15Ecological Climatology © 2008 G. Bonan

Page 16: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.16Ecological Climatology © 2008 G. Bonan

Page 17: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

400 km 400 km

Spruce Migration Oak MigrationThousand Years BP Thousand Years BP

12

1010

14

14

13

15

15

14

11

15

13

14

12

1011

11

12

98

8

7

Contour From 7 To 15 By 1Contour From 10 To 15 By 1

14

15

13

109

12

13

11

12

Figure 24.17Ecological Climatology © 2008 G. Bonan

Page 18: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

0

0.3

-10 25 60Temperature (C)

g C

O2 g

-1 s

-1

Root

HeterotrophicRespiration

Growth Respiration

g C

O2 g

-1 s

-1

0 1 2

Foliage Nitrogen (%)

0 15 30

Temperature (C)

g C

O2 g

-1 s

-1

0 500 1000

Ambient CO2 (ppm)

Photosynthesis

0 -1 -2

Foliage Water Potential (MPa)

g C

O2 g

-1 s

-1

0 1500 3000

Vapor Pressure Deficit (Pa)

46

20

0 500 1000

PPFD (mol m-2 s-1)

46

20

46

20

Sapwood

0

0.01

-10 25 60Temperature (C)

g C

O2 g

-1 s

-1

Foliage

0

0.5

-10 25 60Temperature (C)

g C

O2 g

-1 s

-1

0 15 30Temperature

(C)

Re

lativ

e R

ate

1

8

Soil Water (% saturation)

Re

lativ

e R

ate

0 1000

1

AutotrophicRespiration

Litterfall

NutrientUptake

Foliage

Stem

Root

Soil CarbonMineralization

Figure 24.18Ecological Climatology © 2008 G. Bonan

Page 19: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Duke Forest, NCNiwot Ridge, CO Metolius, OR

ET

(m

m m

onth

-1)

GP

P (

g C

m-2 m

onth

-1)

0

20

40

60

80

100

120

0

20

40

60

80

100

120

0

20

40

60

80

100

120

0

-50

-100

-150

-200

-250

0

-50

-100

-150

-200

-250

0

-50

-100

-150

-200

-250

0 2 4 6 8 10 12 0 2 4 6 8 10 120 2 4 6 8 10 12

0 2 4 6 8 10 12 0 2 4 6 8 10 120 2 4 6 8 10 12

Month MonthMonth

Month Month Month

Figure 24.19Ecological Climatology © 2008 G. Bonan

Page 20: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

200

0

-200

-400

-600

-800

-1000

Net

Eco

syst

em P

rodu

ctio

n (g

C m

-2 y

r-1)

Bio

me

tric

Edd

yco

vari

an

ce

BG

C+

+

BIO

ME

-BG

C

CA

NO

AK

EA

LC

O

eco

sys

INT

RA

STA

ND

La

RS

Lo

TE

C

PnE

T-I

I

400

500

600

700

800

900

1000

1100

Eva

potr

ansp

iratio

n (m

m y

r-1)

Wa

ters

he

d

Edd

yco

vari

an

ce

BG

C+

+

BIO

ME

-BG

C

CA

NO

AK

EA

LC

O

eco

sys

INT

RA

STA

ND

La

RS

Lo

TE

C

PnE

T-I

I

Figure 24.20Ecological Climatology © 2008 G. Bonan

Page 21: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.21Ecological Climatology © 2008 G. Bonan

Page 22: Tundra desert woodland shrubland grassland boreal forest temperate forest tropical seasonal forest tropical rain forest thorn scrub savanna 302520151050-5-10-15

Figure 24.22

100

80

60

40

20

0

Per

cent

Cov

er (

%)

0 200 400 600 800 1000

Years

Arctic grass

Boreal deciduous tree

Boreal needleleaf evergreen tree

Ecological Climatology © 2008 G. Bonan