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AN UPDATED DATABASE OF SERPENTINE ENDEMISMIN THE CALIFORNIA FLORA

Authors: Safford, Hugh, and Miller, Jesse E. D.

Source: Madroño, 67(2) : 85-104

Published By: California Botanical SocietyURL: https://doi.org/10.3120/0024-9637-67.2.85

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MADRONO, Vol. 67, No. 2, pp. 85–104, 2020

AN UPDATED DATABASE OF SERPENTINE ENDEMISM IN THE CALIFORNIA FLORA

HUGH SAFFORD

USDA Forest Service, Pacific Southwest Region, 1323 Club Drive, Vallejo, CA 94592;Department of Environmental Science and Policy, University of California, Davis, CA 95616

[email protected]

JESSE E. D. MILLER

Department of Biology, Stanford University, Palo Alto, CA 94305

ABSTRACT

We update and revise the serpentine affinity database of Safford et al. (2005), which documents levels ofplant taxon association with ultramafic (‘‘serpentine’’) substrates in the California flora. The revised databasereflects recent taxonomic changes consistent with the second edition of the Jepson Manual (TJM2) and morerecent updates to the Jepson eFlora, and includes additional species that were not previously documented asserpentine associates. We also include serpentine-associated species that have been described since thepublication of TJM2 and are not yet incorporated in the eFlora. A number of taxa were removed from theSafford et al. (2005) database due to new ecological information or taxonomic changes. As before, the newdatabase presents information on rarity, geographic distribution, taxonomy, and lifeform. Based on our newdatabase numbers and TJM2’s list of California endemic species, the percentage of California endemic fullspecies that are þ/� restricted to ultramafic substrates has risen to 14.7%. Of 255 total endemic taxa in ourdatabase (including strict and ‘‘broad’’ endemics), 148 (c. 60%) come from only ten plant families,concentrated mostly in one or two genera per family. The North Coast and Klamath Ranges continue tosupport more serpentine endemics than the rest of the State combined. The previous version of the databasehas been widely used in biodiversity research and conservation management, and we hope that this improvedand updated version will prove as valuable.

Key Words: California flora, edaphic, endemics, serpentine, ultramafic.

Soils derived from ultramafic parent materials–often referred to as ‘‘serpentine’’–support a greatdiversity of plant life in California. Serpentine soilsare generally ecologically stressful environmentsbecause they are poor in plant macronutrients, havehigh magnesium : calcium ratios, and contain toxicheavy metals (Alexander et al. 2007). Serpentineenvironments support high levels of native plantdiversity partly because they are less likely to supportthe (often non-native) dominant species that com-petitively exclude other plants on more fertilesubstrates (Brooks 1987). In addition, the stressfulenvironment of serpentine leads to high levels ofendemism. While serpentine soils occupy less than1.5% of California’s geographic area (Harrison et al.2000), a high number of California’s endemic plantspecies are partially or entirely restricted to serpen-tine soils (described below, also see Safford et al.2005; Safford 2011). Understanding the plant diver-sity associated with serpentine environments isimportant for facilitating research into basic ecolog-ical and evolutionary questions (of which there is along history in California serpentine; Harrison andRajakaruna 2011), as well as for informing manage-ment decisions involving landscapes, natural distur-bances, and conservation (Safford and Harrison2004, 2008).

To this point, the serpentine endemism databaseof Safford et al. (2005) has provided the mostcomprehensive documentation of plant species affin-

ities for serpentine substrates in California. Thisdatabase assigns plant taxa a ranking representingtheir degree of specialization to serpentine soils basedon herbarium records, botanical descriptions, fielddata, expert opinion, and published literature. Thedatabase has been widely used for a variety ofpurposes since its publication. The rankings havebeen cited by numerous ecological and evolutionarystudies, such as research on the evolutionary originsof serpentine adaptation (Anacker et al. 2010),biogeographic considerations of plant diversity andendemism in the California flora (Harrison et al.2006; Burge et al. 2011), autoecological studies ofserpentine species (Eckert 2006), and studies ofmycorrhizae on serpentine soils (Southworth et al.2014). The serpentine affinity rankings have beenintegrated into the CalFlora (http://www.calflora.org) web interface—a database on Californiaplants—where the rankings are displayed on thepages of serpentine-associated species. The serpen-tine database has also been used by land managers inconservation planning, environmental assessmentsand impact statements, and resource managementplans.

METHODS

Our understanding of the systematics and ecologyof the California flora has expanded substantiallysince the earlier version of this database was


published. Here, we provide a revised version of thedatabase (Appendix 1; Supplemental Table 1), withnomenclature updated to reflect The Jepson Manual,2nd edition (TJM2; Baldwin et al. 2012) and thenagain using the most recent Jepson eFlora (mostrecent version accessed November 18, 2019; JepsonFlora Project 2019). This contribution follows onSafford (2011), which provided a lay person’ssummary of the status of the endemism databasejust as TJM2 was being prepared for press. Thegeneral patterns discussed in Safford (2011) are stillvalid, but it was important to incorporate thepublished results from Baldwin et al. (2012) and themost recent Jepson eFlora (Jepson Flora Project2019), and to provide a completely revised version ofthe database itself, which was not part of Safford(2011).

In the current database (Appendix 1), we haveadded numerous additional species that were notincluded on the original 2005 list, many of which arerecently described serpentine-associated species. Wealso removed seven taxa from the list that weconstructed after TJM2 (Table 1) because recenttaxonomic changes and/or further study has indicat-ed they are not strongly associated with serpentinesubstrates. To revise the original list, we first updatednomenclature to current Jepson eFlora names (as ofNovember 18, 2019; Jepson Flora Project 2019). Wethen solicited feedback on the list from numerousexpert California botanists and requested suggestionsfor additional species that should be included. Wealso conducted a literature review for recentlydescribed species that have affinities for serpentine.We then conducted further research on all speciesthat were identified by experts or through the

literature review as possibilities for addition to orremoval from the list, as described below.

We have ranked new additions to the list using thesame semi-quantitative scale as the original database(see Safford et al. 2005 for details), with thedifference that we have reduced all rankings .6 to6, as those resulted from the addition of CalFlorainformation (which at the time could be consideredindependent of our other sources). Today CalFloradirectly references the serpentine affinity database inits species descriptions. Rankings of new additions tothe list presented here are based primarily onherbarium records (mostly accessed via the Consor-tium of California Herbaria (CCH1 2019; CCH22019), habitat descriptions by Baldwin et al. (2012)and the Jepson eFlora (Jepson Flora Project 2019),CalFlora observation records (CalFlora 2019), theprimary literature, and in some cases, direct inputfrom botanical experts. In the CCH herbariumrecords, we accessed at least 150 collection recordsfor each taxon (except where records totaled ,150, inwhich case we accessed all records), using randomselection where necessary to ensure no bias inregional coverage. Where CCH or CalFlora recordswere ,5, we combined the two and treated them as asingle information source. We used the BerkeleyMapper (linked to the CCH website), Google Earth,our field experience, and various geologic andjurisdictional (e.g., US Forest Service) maps tovisualize the locations of herbarium records and todetermine, when possible, whether botanical recordslacking geological information were found on or offof ultramafic substrates. Taxa with .50% of usablerecords (‘‘usable’’ ¼ the records identified thesubstrate, or we were able to determine the substrateby another fashion) from serpentine made their way

TABLE 1. TAXA DROPPED FROM SERPENTINE AFFINITY DATABASE AFTER 2012. TJM2 (Baldwin et al. 2012) includedmany taxonomic changes and new taxa. These changes were incorporated into the database at the time of TJM2. This tableincludes taxa dropped from the list since that time.

Species Family Reason and source

Chlorogalum purpureum var. reductumHoover

Agavaceae Taxon does not grow on serpentine soils (Kofron et al. 2013)

Cardamine pachystigma var.pachystigma (S. Watson) Rollins

Brassicaceae Taxon is not a serpentine indicator; reanalysis of herbariumand collection records

Sedum obtusatum ssp. obtusatum A.gray.

Crassulaceae Taxon is not a serpentine indicator; reanalysis of herbariumand collection records

Carex amplectens Steud. Cyperaceae Moved to Carex fracta Mack. by Baldwin et al. (2012), whichis not a serpentine indicator

Swertia fastigiata Pursh Gentianaceae Moved to Frasera umquaensis M. Peck & Applegate byBaldwin et. al (2012), which is not a serpentine indicator

Calochortus greenei S. Watson Liliaceae Taxon does not grow on serpentine soils; government agency(BLM, USFWS) reports on species ecology

Ceanothus papillosus Torr. & A. Grayvar. roweanus McMinn

Rhamnaceae Taxon not recognized by TJM2 or Jepson eFlora, herbariumcollections and CalFlora observations do not support taxonas serpentine indicator

Brodiaea pallida Hoover Themidaceae New collections show that only 1 of 5 population groupsoccurs on serpentine; R. Preston, UC Davis, pers. comm.

Brodiaea purdyi Eastw. Themidaceae Taxon subsumed within B. minor (Benth.) S. Watson, whichis not a serpentine indicator (Preston 2006; Baldwin et al.2012)

86 [Vol. 67MADRONO


onto the final list. Rankings of new list additions(e.g., recently described species) should be usedcautiously since many of them are based on limiteddata, and we suggest that all species ranked based onthree or fewer sources are in need of further study.All species in the updated list follow the nomencla-ture of the current (November 2019) Jepson eFloraexcept for 13 taxa (Table 2), most of which arerecently described.

We emphasize that we did not carry out acomplete search of new collection records for everytaxon since 2005. This was only carried out for taxawhere recent publications, taxonomic changes, orpersonal communications from California botanistsled us to believe that such a search might change thestatus of the species. For new additions to the list, weno longer required three separate sources of infor-mation supporting their affinity for serpentine ifevidence for their affinity to serpentine appearedstrong (for example, in the herbarium records).

Finally, it is important to note that for species forwhich all described infraspecific taxa (subspecies,varieties) exhibit the same level of serpentine affinity,we only include the full species name in our database.

RESULTS AND DISCUSSION

As in Safford et al. (2005) and Safford (2011), wedefine serpentine endemic taxa as those that scored atleast 4.5 in our ranking process. Put another way, ourclass of ‘‘serpentine endemics’’ includes strict endem-ics (more than c. 95% of recorded occurrences onultramafics) and broad endemics (c. 85–94%). Theupdated serpentine affinity database (Appendix 1)contains 255 endemic taxa (strict plus broad endem-ics), 193 of which are full species (this includes fivespecies for which all of the subspecies are listed in thedatabase due to differing levels of serpentine affinity:Arctostaphylos bakeri Eastw., A. montana Eastw.,Calystegia collina (Greene) Brummitt, Cirsium fonti-

nale (Greene) Jeps., and Lessingia micradeniaGreene). Thus, serpentine endemic species accountfor about 14.7% – more than 1/7 – of the endemicspecies in California (based on 1315 endemic speciesin Baldwin 2012). This is a notable increase from the12.5% reported by Safford et al. (2005), and is due toadditions of newly described species and a decrease inthe number of California endemic species accordingto Baldwin et al. (2012).

Taxonomic revisions have led to some changesfrom Safford et al. (2005) in the ordering of familyand genera with the most serpentine endemic taxa.Asteraceae remains the family with the most serpen-tine endemics, with 25 strict endemics, 13 broadendemics, and 106 total serpentine-associated taxa(Table 3). Liliaceae, which historically contained thesecond most serpentine endemics of any family(considering both strict and broad endemics), hasbeen split into several smaller families, many ofwhich are further down the current list, includingAlliaceae – also in the top ten families, Agavaceae,Themidaceae, and so on. As a result, Brassicaceae isnow the second most endemic-rich family (Table 3).Scrophulariaceae was also split in Baldwin et al.(2012) into numerous plant families, and it hasdropped from a fifth place ranking in the Safford etal. (2005) rankings to a complete absence from thelist now. Families derived from Scrophulariaceae arestill important components of serpentine endemismin California however: for example, Orobanchaceaecurrently occupies sixth place on the list, andPhrymaceae and Plantaginaceae are further downthe list. Eriogonum is now the most endemic-richgenus, followed by Streptanthus (Table 4).

As discussed by Safford et al. (2005; see their Figs1 and 2), the distribution of taxa among plantfamilies and genera is very different when comparingserpentine endemics and California endemics. Fam-ilies which support many more serpentine endemictaxa than expected include Linaceae, Polygonaceae,

TABLE 2. TAXA INCLUDED IN THE SERPENTINE AFFINITY LIST THAT ARE NOT IN THE CURRENT JEPSON EFLORA (AS OF

NOVEMBER 18, 2019). These include recently described taxa as well as taxa that are widely recognized by botanists inCalifornia, but are not included in the current eFlora because of taxonomic uncertainty.

Taxon Source

Brodiaea rosea ssp. rosea (Greene) Baker Preston 2013Calystegia vanzuukiae Brummitt & Namoff Brummitt and Namoff 2013Cardamine pachystigma var. dissectifolia (S. Watson) Rollins R. Preston and L. Janeway, pers comm and CCH

and CalFlora recordsClaytonia serpenticola T.R. Stoughton Stoughton 2017Dicentra formosa ssp. oregana (Haw.) Walp. Baldwin et al. 2012; L. Hoover, U.S. Forest

Service, pers. comm.Erythranthe willisii G.L. Nesom Nesom 2017Hesperolinon sharsmithiae R. O’Donnell O’Donnell 2006Sedum eastwoodiae (Britton) A. Berger Zika et al. 2018Sedum flavidum B.L. Wilson & Zika Zika et al. 2018Sedum kiersteadiae B.L. Wilson & R.E. Brainerd Wilson et al. 2014Sedum patens Zika Zika et al. 2018Sedum rubiginosum Zika & B.L. Wilson Zika et al. 2018Vaccinium coccineum Piper L. Hoover and J. Nelson, U.S. Forest Service,

pers comm.

2020] 87SAFFORD AND MILLER: SERPENTINE AFFINITY DATABASE


Brassicaceae, Apiaceae, Caryophyllaceae, and Lilia-ceae. Genera that follow the same pattern of notableenrichment in serpentine endemics include Streptan-thus, Minuartia, Cordylanthus, Packera, and Hesper-olinon.

Of the five geographic regions identified bySafford et al. (2005), the North Coast region remainsthe geographic leader in number of serpentineendemic taxa (127), and the Klamath region supportsthe most serpentine endemics that are restricted to a

TABLE 3. NUMBERS OF SERPENTINE ENDEMIC AND NEAR

ENDEMIC TAXA BY FAMILY. 1 Strict endemics. 2 Strictendemics plus broad endemics (‘‘endemic taxa’’ in thispaper). 3 Strict and broad endemics plus ‘‘near endemic’’taxa (taxa transitional from strong indicators to broadendemics). 4 Total taxa in our database, including weakindicators and taxa transitional from indifferent to weakindicators. Families ranked by numbers of endemics (strictþ broad).

Family

Serpentine affinity score

Total4�5.51 �4.52 �3.53

Asteraceae 25 38 46 106Brassicaceae 19 24 28 38Polygonaceae 11 19 22 41Liliaceae 5 13 16 38Polemoniaceae 9 10 11 21Orobanchaceae 7 10 10 19Apiaceae 7 9 12 33Crassulaceae 8 9 9 16Linaceae 8 9 9 14Ericaceae 5 8 10 15Fabaceae 4 7 11 25Alliaceae 4 7 11 24Lamiaceae 6 7 9 17Boraginaceae 6 6 11 24Caryophyllaceae 5 6 8 18Campanulaceae 3 5 8 13Rhamnaceae 4 5 7 14Onagraceae 3 5 7 13Cyperaceae 3 5 7 12Convolvulaceae 1 5 6 7Phrymaceae 4 4 6 8Rubiaceae 3 4 4 8Montiaceae 0 3 6 17Rosaceae 1 3 6 11Plantaginaceae 1 3 5 15Poaceae 1 3 3 18Iridaceae 2 3 3 5Violaceae 0 2 3 7Cupressaceae 0 2 3 6Malvaceae 2 2 2 7Agavaceae 2 2 2 5Salicaceae 2 2 2 3Gentianaceae 2 2 2 2Garryaceae 1 2 2 2Themidaceae 0 1 5 8Ranunculaceae 1 1 3 7Papaveraceae 0 1 1 5Berberidaceae 1 1 1 4Pteridaceae 0 1 1 4Fagaceae 1 1 1 3Orchidaceae 0 1 1 3Dryopteridaceae 1 1 1 2Lentibulariaceae 1 1 1 1Apocynaceae 1 1 1 1Verbenaceae 0 1 1 1Melanthiaceae 0 0 2 4Pinaceae 0 0 1 6Parnassiaceae 0 0 1 2Saxifragaceae 0 0 1 1Sarraceniaceae 0 0 1 1Tecophilaeaceae 0 0 0 1Primulaceae 0 0 0 1Polygalaceae 0 0 0 1Cistaceae 0 0 0 1

TABLE 4. GENERA WITH MORE THAN THREE TAXA

ENDEMIC TO SERPENTINE (MEAN AFFINITY SCORE � 4.5).

Genus Family Number of taxa

Eriogonum Polygonaceae 19Streptanthus Brassicaceae 18Allium Alliaceae 11Arctostaphylos Ericaceae 9Hesperolinon Linaceae 9Carex Cyperaceae 7Lomatium Apiaceae 7Erythranthe Phrymaceae 6Navarretia Polemoniaceae 6Packera Asteraceae 6Phacelia Boraginaceae 6Sedum Crassulaceae 6Calystegia Convolvulaceae 5Campanula Campanulaceae 5Cordylanthus Orobanchaceae 5Erigeron Asteraceae 5Fritillaria Liliaceae 5Monardella Lamiaceae 5Boechera Brassicaceae 4Calochortus Liliaceae 4Castilleja Orobanchaceae 4Ceanothus Rhamnaceae 4Cirsium Asteraceae 4Galium Rubiaceae 4Harmonia Asteraceae 4Horkelia Rosaceae 4Lessingia Asteraceae 4

FIG. 1. Geographic distribution of serpentine endemictaxa (mean score � 4.5) in California. Dark areas of barsrepresent taxa restricted to a single geographic region, andlight areas represents taxa that also occur in other regions.Geographic regions are from Baldwin et al. (2012).

88 [Vol. 67MADRONO


single region (60; Fig. 1). Together, the North Coastand Klamath regions support 146 serpentine endemictaxa that are found nowhere else in the state. SeeSafford (2011) for a figure and short discussionregarding the elevational distribution of serpentineendemic taxa in California. In short, most taxa growin habitats below 1000 m elevation and few are foundabove 1500 m, primarily due to the relative rarity ofultramafic outcrops above that elevation.

Among the endemic taxa, 216 are dicots, 35 aremonocots, two are gymnosperms, and two arepteridophytes (Appendix 1). The distribution of lifeforms is shown in Fig. 2. Serpentine endemic taxa inCalifornia are primarily perennial forbs (153 of 255taxa, including three that can also be annuals), mostof the rest are annual forbs (74 taxa, including thesame three taxa); less than 10% of the endemic taxain our database are woody species (Fig. 2).

192 (77%) of the endemic taxa in our database arelisted in the CNPS Inventory of Rare and Endan-gered Plants (CNPS 2019). 108 serpentine endemictaxa are assigned California Rare Plant Rank 1B,which includes Federal and State threatened andendangered taxa as well as those taxa that theCalifornia Native Plant Society believes warrantlisting (Appendix 1).

The richness of serpentine endemic taxa inCalifornia is astonishing, given the limited area ofthe state that supports ultramafic rocks and soils.Yet, amazingly, new species continue to be describedfrom ultramafic soils in California. Only 15 yearsafter embarking on this exercise for the first time, weexpanded the list of California serpentine endemicsby 3.7%. Given the outsized contribution made toCalifornia’s natural riches by ultramafic substratesand their biota, we hope and expect that botanicaland geoecological exploration will continue toexpand the California serpentine flora, as well asour understanding of the factors that make it sodiverse.

ACKNOWLEDGEMENTS

We are grateful to the California botanists and landmanagers who have shared their observations and expertiseon serpentine taxa over the years, and helped us torecognize the need to update the serpentine endemismdatabase. We thank them also for suggesting correctionsand reviewing them as they were made. Many botanistscontributed to this endemics database update; we wouldespecially like to thank Lisa Hoover, Mike Kauffman,Lawrence Janeway, Julie Kierstead, Ryan O’Dell, and RobPreston. Ellen Dean and Barbara Ertter provided advice onnewer taxa that had not yet been listed in the Jepson eFlora.We thank Maaike Wielenga for her assistance in databasemanagement and conducting background research on newlydescribed species. Finally, we thank the Jepson Herbariumstaff for assistance with nomenclature.

LITERATURE CITED

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ANACKER, B. L., W. JUSTEN, E. E. GOLDBERG, AND S. P.HARRISON. 2010. Origins and consequences of serpen-tine endemism in the California Flora. Evolution65:365–376.

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(eds.). 2012. The Jepson manual: vascular plants ofCalifornia. 2nd edition. University of California Press,Berkeley, CA.

BROOKS, R. R. 1987. Serpentine and its vegetation.Dioscorides Press, Portland, OR.

BRUMMITT, R. K., AND S. NAMOFF. 2013. Calystegiavanzuukiae (Convolvulaceae), a remarkable new speciesfrom central California. Aliso 31:15–18.

BURGE, D. O., J. H. THORNE, S. P. HARRISON, L.HARDISON, B. O’BRIEN. 2016. Plant diversity andendemism in the California Floristic Province.Madrono 63:3–216.

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CALIFORNIA NATIVE PLANT SOCIETY, RARE PLANT

PROGRAM. 2020. Inventory of Rare and EndangeredPlants of California (online edition, v8-03 0.39).Website: http://www.rareplants.cnps.org [accessed 18November, 20219].

CCH1. 2019. Consortium of California Herbaria, CCH1Portal. Website: http://ucjeps.berkeley.edu/consortium/[accessed on 19 November 2019].

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HARRISON, S., AND N. RAJAKARUNA (eds.). 2011. Serpen-tine: the evolution and ecology of a model system.University of California Press, Berkeley, CA.

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FIG. 2. Lifeforms of serpentine endemic and near endemictaxa (mean score .¼ 3.5) in California.



serpentine plants of California. Diversity and Distri-butions 6:153–161.

HARRISON, S. P., H. D. SAFFORD, J. GRACE, J. H. VIERS,AND K. F. DAVIES. 2006. Regional and local speciesrichness in an insular environment: serpentine plants inCalifornia. Ecological Monographs 76:41–56.

JEPSON FLORA PROJECT (eds.). 2019. Jepson eFlora, http://ucjeps.berkeley.edu/eflora/ [accessed on 10 January2019].

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BURY, J. OLSON, AND R.F. HOLLAND. 2013. Review ofthe purple amole Chlorogalum purpureum (Agavaceae):a threatened plant in the coast ranges of CentralCalifornia. Bulletin, Southern California Academy ofSciences 112:38–48.

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SAFFORD, H. D., AND S. P. HARRISON. 2008. The effects offire on serpentine vegetation and implications formanagement. Proceedings of the 2002 Fire Conferenceon Managing Fire and Fuels in the RemainingWildlands and Open Spaces of the SouthwesternUnited States. December 2–5, 2002, San Diego, CA.General Technical Report PSW-189. USDA-ForestService, Pacific Southwest Research Station, Albany,CA.

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APPENDIX

1.Updatedlistoftaxaassociatedwithserpentineordered

byfamilyandtaxon.1Names

asin

theJepsoneF

lora

(exceptforspecieslisted

inTable2);

2Aff¼Affinity:SE¼

strict

endem

ic,BE¼

broadendem

ic,BE/SI¼

broadendem

ic/strongindicator,SI¼

strongindicator,WI¼

weakindicator,WI/IN¼

weakindicator/indifferent.

3Meanaffinity

score.4Sum

ofallaffinityscores.

5Medianofaffinityscores.

6Standard

deviationofaffinityscores.

7California

NativePlantSocietyrarity

codes

(CNPS2019).

8Geographic

distribution:KL¼

Klamath

Ranges,NC¼

NorthCoast

Ranges,BA¼

SanFranciscoBayArea,SC¼

South

Coast

Ranges,SN¼

SierraNevada.9Lifeform

sare

abbreviatedas

follows:carn¼

carnivorous,cesp¼

cespitose,hem

ipar¼

hem

iparasitic,

paras¼

parasitic,

rhiz¼

rhizomatous.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Chlorogalum

angustifolium

Kellogg

Agavaceae

WI

2.4

9.5

41.8

2.8

11

Perennialforb

(bulb)

Chlorogalum

grandiflorum

Hoover

Agavaceae

WI/IN

12

21

-1B.2

1Perennialforb

(bulb)

Chlorogalum

pomeridianum

var.minus

(DC.)Kunth

Agavaceae

SE

630

56

01B.2

11

1Perennialforb

(bulb)

Hastingsiaalba

(Durand)S.Watson

Agavaceae

SI

3.4

17

53

1.5

11

1Perennialforb

(bulb)

Hastingsiaserpentinicola

R.Becking

Agavaceae

SE

618

36

01

1Perennialforb

(bulb)

Allium

acuminatum

Hook.

Alliaceae

WI

1.5

4.5

32

0.9

11

1Perennialforb

(bulb)

Allium

amplectens

Torr.

Alliaceae

WI

2.3

11.3

52

2.2

11

11

1Perennialforb

(bulb)

Allium

bolanderivar.bolanderi

S.Watson

Alliaceae

WI/IN

1.1

4.5

41

0.6

11

1Perennialforb

(bulb)

Allium

bolanderivar.mirabile

S.Watson

Alliaceae

WI

24

22

-1

1Perennialforb

(bulb)

Allium

cratericola

Eastw.

Alliaceae

SI

2.6

15.8

62.5

1.9

CBR

11

1Perennialforb

(bulb)

Allium

crispum

Greene

Alliaceae

WI/IN

1.3

3.75

31

0.7

11

Perennialforb

(bulb)

Allium

diabolense

(Ownbey

&Aase

exTraub)

McN

eal

Alliaceae

SE

618

36

01

Perennialforb

(bulb)

Allium

falcifolium

Hook.&

Arn.

Alliaceae

BE/SI

4.2

38

94

1.6

11

1Perennialforb

(bulb)

Allium

fimbriatum

var.purdyi

S.Watson

Alliaceae

BE

5.4

21.5

46

1.5

4.3

1Perennialforb

(bulb)

Allium

hoffmanii

Ownbey

exTraub

Alliaceae

SE

630

56

04.3

11

Perennialforb

(bulb)

Allium

howelliivar.sanbenitense

Eastw.

Alliaceae

BE/SI

412

34

11B.3

1Perennialforb

(bulb)

Allium

jepsonii

(Ownbey

&Aase

exTraub)

S.S.Denison&

McN

eal

Alliaceae

BE

5.4

37.5

76

11B.2

1Perennialforb

(bulb)

Allium

lacunosum

var.lacunosum

S.Watson

Alliaceae

BE/SI

3.8

15.3

44.5

2.8

11

Perennialforb

(bulb)

Allium

lacunosum

var.micranthum

S.Watson

Alliaceae

BE/SI

4.3

13

36

2.9

CBR

1Perennialforb

(bulb)

Allium

mem

branaceum

Ownbey

exTraub

Alliaceae

WI/IN

1.3

43

11.5

CBR

11

Perennialforb

(bulb)

Allium

obtusum

var.conspicuum

Lem

mon

Alliaceae

WI/IN

12

21

-1

Perennialforb

(bulb)

Allium

peninsulare

var.franciscanum

Lem

monex

Greene

Alliaceae

WI

1.8

3.5

21.8

-1B.2

11

Perennialforb

(bulb)

Allium

sanborniivar.congdonii

Alph.Wood

Alliaceae

SE

5.6

22.5

46

14.3

1Perennialforb

(bulb)

Allium

sanborniivar.sanbornii

Alph.Wood

Alliaceae

SI

3.4

27

83.5

2.2

4.2

1Perennialforb

(bulb)

Allium

serra

McN

eal&

Ownbey

Alliaceae

SI

2.6

10

43

1.5

11

1Perennialforb

(bulb)

Allium

sharsmithiae

(Ownbey

&Aase

exTraub)

McN

eal

Alliaceae

BE

5.1

20.5

46

21B.3

11

Perennialforb

(bulb)

Allium

siskiyouense

Ownbey

exTraub

Alliaceae

SI

2.8

14

52

1.8

4.3

11

Perennialforb

(bulb)

Allium

tuolumnense

(Ownbey

&Aase

exTraub)

S.S.Denison&

McN

eal

Alliaceae

SE

618

36

01B.2

1Perennialforb

(bulb)

Allium

unifolium

Kellogg

Alliaceae

WI/IN

13

31

11

11

Perennialforb

(bulb)

Angelicatomentosa

S.Watson

Apiaceae

SI

2.7

83

31.5

11

1Perennialforb

Apiastrum

angustifolium

Nutt.

Apiaceae

WI

1.5

7.6

50.1

2.5

11

11

Annualforb

Ligusticum

californicum

J.M.Coult.&

Rose

Apiaceae

WI/IN

1.4

5.75

41.4

1.3

11

1Perennialforb

Lomatium

ciliolatum

Jeps.

Apiaceae

SE

618

36

01

11

Perennialforb

Lomatium

congdonii

J.M.Coult.&

Rose

Apiaceae

SE

618

36

01B.2

1Perennialforb

Lomatium

dasycarpum

ssp.dasycarpum

(Torr.&

A.Gray)J.

M.

Coult.&

Rose

Apiaceae

BE/SI

3.6

21.5

63.5

1.9

11

1Perennialforb

Lomatium

engelmannii

Mathias

Apiaceae

SE

5.8

34.5

66

0.8

4.3

11

Perennialforb

Lomatium

hooveri

(Mathias&

Constance)

Constance

&Ertter

Apiaceae

SE

5.9

29.5

56

0.4

4.3

1Perennialforb

Lomatium

howellii

(S.Watson)Jeps.

Apiaceae

SE

624

46

04.3

1Perennialforb

Lomatium

macrocarpum

(Torr.&

A.Gray)J.

M.

Coult.&

Rose

Apiaceae

SI

2.7

83

30.6

11

11

1Perennialforb

Lomatium

marginatum

var.marginatum

(Benth.)J.

M.Coult.&

Rose

Apiaceae

WI

2.3

94

2.5

2.1

11

1Perennialforb



APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Lomatium

marginatum

var.purpureum

(Benth.)J.

M.Coult.&

Rose

Apiaceae

SI

3.3

13

43

2.2

CBR

11

Perennialforb

Lomatium

observatorium

Constance

&Ertter

Apiaceae

WI/IN

1.4

2.75

21.4

-1B.2

11

Perennialforb

Lomatium

parvifolium

(Hook.&

Arn.)Jeps.

Apiaceae

SI

3.3

13

43

2.3

4.2

1Perennialforb

Lomatium

repostum

(Jeps.)Mathias

Apiaceae

SI

3.2

12.6

43

2.4

4.3

1Perennialforb

Lomatium

tracyi

Mathias&

Constance

Apiaceae

SE

642

76

04.3

11

Perennialforb

Lomatium

triternatum

var.triternatum

(Pursh)J.

M.Coult.&

Rose

Apiaceae

SI

2.8

11

42

2.4

11

Perennialforb

Lomatium

utriculatum

(Nutt.ex

Torr.&

A.Gray)

J.M.Coult.&

Rose

Apiaceae

WI

1.7

8.5

51

1.4

11

11

1Perennialforb

Perideridia

bacigalupii

Chuang&

Constance

Apiaceae

BE

4.6

23

56

2.4

4.2

1Perennialforb

Perideridia

kelloggii

(A.Gray)Mathias

Apiaceae

WI

2.1

10.6

52

21

11

1Perennialforb

Perideridia

leptocarpa

T.I.Chuang&

Constance

Apiaceae

SE

5.6

22.5

46

14.3

1Perennialforb

Perideridia

oregana

(S.Watson)Mathias

Apiaceae

WI

1.7

53

11.2

11

11

Perennialforb

Perideridia

pringlei

(J.M.Coult.&

Rose)A.

Nelson&

J.F.Macbr.

Apiaceae

BE/SI

3.7

18.5

53

2.3

4.3

1Perennialforb

Sanicula

bipinnatifida

Hook.

Apiaceae

WI

1.8

7.1

42

1.5

11

11

1Perennialforb

Sanicula

hoffmannii

(Munz)

R.H.Shan&

Constance

Apiaceae

WI

1.8

3.5

21.5

-4.3

1Perennialforb

Sanicula

maritima

S.Watson

Apiaceae

WI

2.3

4.5

22

-1B.1

11

Perennialforb

Sanicula

peckiana

J.F.Macbr.

Apiaceae

BE

5.3

26.5

56

1.3

4.3

1Perennialforb

Sanicula

tracyi

R.H.Shan&

Constance

Apiaceae

WI

2.1

8.5

41

2.6

4.2

1Perennialforb

Sanicula

tuberosa

Torr.

Apiaceae

WI/IN

1.3

3.75

31

0.7

11

1Perennialforb

Tauschia

glauca

(J.M.Coult.&

Rose)

Mathias&

Constance

Apiaceae

BE/SI

3.5

10.5

33

0.6

4.3

1Perennialforb

Tauschia

hartwegii

(A.Gray)J.

F.Macbr.

Apiaceae

WI/IN

1.3

43

11.5

11

1Perennialforb

Tauschia

howellii

(J.M.Coult.&

Rose)J.

F.

Macbr.

Apiaceae

WI

2.3

73

13.2

1B.3

1Perennialforb

Tauschia

kelloggii

(A.Gray)J.

F.Macbr.

Apiaceae

SI

2.6

12.8

52

2.2

11

1Perennialforb

Asclepiassolanoana

Woodson

Apocynaceae

SE

642

76

04.2

1Perennialforb

Agoserisheterophylla

(Nutt.)Greene

Asteraceae

WI/IN

1.4

4.1

31

1.5

11

11

1Annualforb

Ancistrocarphusfilagineus

A.Gray

Asteraceae

SI

3.3

13

43

0.5

11

11

1Annualforb

Antennariaargentea

Benth.

Asteraceae

WI

1.9

7.75

40.8

2.7

11

Perennialforb

Antennariasawyeri

R.J.Bayer

&Figura

Asteraceae

SE

618

36

01B.2

1Perennialforb

Antennariasuffrutescens

Greene

Asteraceae

SE

5.6

22.5

46

14.3

11

Perennialforb

Arnicacernua

Howell

Asteraceae

SE

624

46

04.3

1Perennialforb

(rhiz.)

Arnicaspathulata

Greene

Asteraceae

SE

5.5

16.5

36

1.2

4.3

1Perennialforb

(rhiz.)

Balsamorhizamacrolepis

W.M.Sharp

Asteraceae

SI

2.5

15

62

1.1

1B.2

11

Perennialforb

Balsamorhizasericea

W.A

.Weber

Asteraceae

SE

618

36

01B.3

1Perennialforb

Benitoaoccidentalis

(H.M.Hall)D.D.Keck

Asteraceae

BE/SI

4.1

16.5

44

1.8

4.3

1Annualforb

Brickelliagreenei

A.Gray

Asteraceae

BE/SI

3.7

11

34

0.6

11

1Perennialforb

Cacaliopsisnardosm

ia(A

.Gray)A.Gray

Asteraceae

WI/IN

1.3

43

21.2

11

Perennialforb

Calycadenia

multiglandulosa

DC.

Asteraceae

SI

3.1

15.5

53

1.2

11

11

Annualforb

Calycadenia

oppositifolia

(Greene)

Greene

Asteraceae

SI

2.6

18

72

1.6

4.2

1Annualforb

Calycadenia

pauciflora

A.Gray

Asteraceae

BE

5.3

21

45.5

11

Annualforb

Calycadenia

truncata

DC.

Asteraceae

WI

2.1

12.5

62.5

1.1

11

11

1Annualforb

Chaenactisglabriuscula

var.glabriuscula

DC.

Asteraceae

WI

1.7

5.1

32

1.5

11

1Perennialforb

Chaenactisglabriuscula

var.heterocarpha

DC.

Asteraceae

SI

2.5

10

42.5

0.6

11

11

Annualforb

Chaenactissuffrutescens

A.Gray

Asteraceae

SE

630

56

01B.3

1Perennialforb

Cirsium

andrewsii

(A.Gray)Jeps.

Asteraceae

WI

1.7

53

20.9

1B.2

1Perennialforb

Cirsium

cymosum

(Greene)

J.T.Howell

Asteraceae

SI

312

42

21

11

11

Perennialforb

Cirsium

douglasiivar.breweri

DC.

Asteraceae

SI

312

43

1.6

11

1Perennialforb

Cirsium

fontinale

var.campylon

(Greene)

Jeps.

Asteraceae

SE

5.9

29.5

56

0.4

1B.2

11

Perennialforb

Cirsium

fontinale

var.fontinale

(Greene)

Jeps.

Asteraceae

SE

630

56

01B.1

11

Perennialforb

92 [Vol. 67MADRONO


APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Cirsium

fontinale

var.obispoense

(Greene)

Jeps.

Asteraceae

SE

624

46

01B.2

1Perennialforb

Cirsium

hydrophilum

var.vaseyi

(Greene)

Jeps.

Asteraceae

SE

624

46

01B.2

11

Perennialforb

Cirsium

remotifolium

(Hook.)DC.

Asteraceae

WI/IN

13.1

31

11

11

Perennialforb

Corethrogynefilaginifolia

(Hook.&

Arn.)Nutt.

Asteraceae

WI/IN

1.3

43

11.5

11

11

1Perennialforb,Shrub

Crepispleurocarpa

A.Gray

Asteraceae

WI

210

52

0.7

11

1Perennialforb

Deinandra

halliana

(D.D.Keck)B.G.

Baldwin

Asteraceae

SI

312

43

2.4

1B.2

1Annualforb

Ericameria

arborescens

(A.Gray)Greene

Asteraceae

WI/IN

1.3

43

11.5

11

11

Shrub

Ericameria

greenei

(A.Gray)G.L.Nesom

Asteraceae

WI

28.1

41.5

2.1

11

1Shrub

Ericameria

nauseosa

var.oreophila

(Pall.ex

Pursh)G.L.

Nesom

&G.I.Baird

Asteraceae

WI

1.8

8.85

52

1.5

11

1Shrub

Ericameria

ophitidis

(J.T.Howell)G.L.

Nesom

Asteraceae

SE

5.5

38.5

76

14.3

11

Shrub

Erigeronbloomerivar.nudatus

A.Gray

Asteraceae

SE

618

36

02B.3

1Perennialforb

Erigeroncervinus

Greene

Asteraceae

SI

3.3

10

34

3.1

4.3

1Perennialforb

(rhiz.)

Erigeronfoliosusvar.confinis

Nutt.

Asteraceae

BE/SI

3.7

11

33

1.2

1Perennialforb

Erigerongreenei

G.L.Nesom

Asteraceae

BE

4.8

24

54

1.1

1B.2

1Perennialforb

Erigeronlassenianusvar.deficiens

Greene

Asteraceae

WI

1.7

53

21.5

1B.3

1Perennialforb

Erigeronpetrophilusvar.sierrensis

Greene

Asteraceae

BE

4.8

28.5

66

2.1

4.3

1Perennialforb

(rhiz.)

Erigeronpetrophilusvar.viscidulus

Greene

Asteraceae

WI

2.4

9.5

42

0.5

4.3

1Perennialforb

(rhiz.)

Erigeronreductus

(Cronquist)

G.L.Nesom

Asteraceae

WI

28

42

1.6

11

1Perennialforb

Erigeronrobustior

(Cronquist)

G.L.Nesom

Asteraceae

WI

1.5

4.5

32

1.2

4.3

1Perennialforb

Erigeronserpentinus

G.L.Nesom

Asteraceae

SE

618

36

01B.3

1Perennialforb

Eriophyllum

confertiflorum

var.

tanacetiflorum

(DC.)A.Gray

Asteraceae

WI

1.9

3.75

21.9

-4.3

1Shrub

Eriophyllum

jepsonii

Greene

Asteraceae

BE/SI

3.5

17.5

53

1.5

4.3

11

Shrub

Eriophyllum

lanatum

var.achilleoides

(Pursh)J.

Forbes

Asteraceae

WI

2.3

73

20.6

11

11

1Shrub

Eriophyllum

lanatum

var.lanceolatum

(Pursh)J.

Forbes

Asteraceae

WI

1.7

53

21.5

11

Shrub

Eriophyllum

latilobum

Rydb.

Asteraceae

SE

5.5

16.5

36

1.2

1B.1

11

Shrub

Grindelia

camporum

Greene

Asteraceae

WI

1.8

5.25

32

1.4

11

Perennialforb

Grindelia

hirsutula

Hook.&

Arn.

Asteraceae

WI/IN

1.4

8.3

61

11

1Perennialforb

Gutierreziacalifornica

(DC.)Torr.&

A.Gray

Asteraceae

WI

1.8

5.25

32

1.4

CBR

11


Harm

onia

doris-nilesiae

(T.W.Nelson&

J.P.

Nelson)B.G.Baldwin

Asteraceae

BE

5.4

32.5

65.5

0.8

1B.1

1Annualforb

Harm

onia

guggolziorum

B.G.Baldwin

Asteraceae

SE

618

36

01B.1

1Annualforb

Harm

onia

hallii

(D.D.Keck)B.G.

Baldwin

Asteraceae

SE

624

46

01B.2

1Annualforb

Harm

onia

stebbinsii

(T.W.Nelson&

J.P.

Nelson)B.G.Baldwin

Asteraceae

SE

642

76

01B.2

1Annualforb

Hazardia

stenolepis

(H.M.Hall)Hoover

Asteraceae

WI

26.1

33

1.7

CBR

1Shrub(stem

succulent)

Hazardia

whitneyivar.discoidea

(A.Gray)Greene

Asteraceae

WI

1.9

5.75

32

1.1


Hazardia

whitneyivar.whitneyi

(A.Gray)Greene

Asteraceae

WI/IN

12

21

-1

Perennialforb,shrub

Helenium

bigelovii

A.Gray

Asteraceae

SI

2.9

11.5

42.5

2.3

11

11

Perennialforb

Helianthusexilis

A.Gray

Asteraceae

SE

5.7

45.5

86

1.1

4.2

11

11

Annual,Perennialforb

Hem

izonia

congesta

ssp.calyculata

DC.

Asteraceae

WI

1.5

4.5

32

1.2

4.3

1Annualforb

Hem

izonia

congesta

ssp.congesta

DC.

Asteraceae

WI/IN

1.3

43

21.2

1B.2

11

1Annualforb

Hem

izonia

congesta

ssp.tracyi

DC.

Asteraceae

WI

1.8

5.25

32

0.7

4.3

1Annualforb

Hesperevaxsparsiflora

var.sparsiflora

(A.Gray)Greene

Asteraceae

WI

1.8

7.25

41.5

1.6

11

1Annualforb

Heterothecaoregonavar.oregona

(Nutt.)Shinners

Asteraceae

WI

26

33

1.7

11

1Perennialforb

Hieracium

bolanderi

A.Gray

Asteraceae

BE/SI

3.8

15

44.5

2.6

11

Perennialforb

Hieracium

greenei

A.Gray

Asteraceae

WI

2.2

6.5

33

1.4

11

Perennialforb

Lagophyllaglandulosa

A.Gray

Asteraceae

WI

26.1

33

1.7

11

Annualforb

Lagophyllaminor

(D.D.Keck)D.D.Keck

Asteraceae

BE

4.7

23.5

55

1.7

CBR

11

Annualforb



APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Layia

discoidea

D.D.Keck

Asteraceae

SE

5.7

28.5

56

0.8

1B.1

1Annualforb

Layia

jonesii

A.Gray

Asteraceae

BE/SI

3.5

10.5

33

0.6

1B.2

1Annualforb

Layia

septentrionalis

D.D.Keck

Asteraceae

SI

3.2

19

63.5

1.4

1B.2

1Annualforb

Leptosynestillm

anii

A.Gray

Asteraceae

SI

2.7

83

30.6

11

1Annualforb

Lessingia

arachnoidea

Greene

Asteraceae

SE

618

36

01B.2

1Annualforb

Lessingia

hololeuca

Greene

Asteraceae

SI

2.5

7.5

33

1.2

31

1Annualforb

Lessingia

micradenia

var.glabrata

Greene

Asteraceae

BE

5.1

30.5

65.5

1.3

1B.2

1Annualforb

Lessingia

micradenia

var.micradenia

Greene

Asteraceae

BE

5.3

31.5

65.5

11B.2

1Annualforb

Lessingia

nem

aclada

Greene

Asteraceae

WI

26

32

11

11

11

Annualforb

Lessingia

ramulosa

A.Gray

Asteraceae

BE

5.4

27

56

1.3

11

Annualforb

Luinahypoleuca

Benth.

Asteraceae

WI/IN

1.4

4.25

32

11

11

Perennialforb

Madia

exigua

(Sm.)A.Gray

Asteraceae

WI

1.8

7.25

42

1.4

11

11

1Annualforb

Malacothrixclevelandii

A.Gray

Asteraceae

SI

39.1

33

31

11

11

Annualforb

Malacothrixfloccifera

(DC.)S.F.Blake

Asteraceae

WI

2.1

6.25

33

1.6

11

11

1Annualforb

Micropusamphibolus

A.Gray

Asteraceae

WI

2.4

7.25

31

3.1

3.2

11

1Annualforb

Microserisdouglasii

(DC.)Sch.Bip.

Asteraceae

WI/IN

1.3

43

10.6

11

11

Annualforb

Monolopia

gracilens

A.Gray

Asteraceae

WI

2.4

4.75

22.4

-1B.2

11

Annualforb

Packeraclevelandii

(Greene)

W.A.Weber

&A.Love

Asteraceae

SE

5.8

46.5

86

0.7

1Perennialforb

Packeraeurycephala

var.eurycephala

(Torr.&

A.Gray)W.A

.Weber

&A.Love

Asteraceae

BE/SI

3.8

15

43

1.5

11

Perennialforb

Packeraeurycephala

var.lewisrosei

(Torr.&

A.Gray)W.A

.Weber

&A.Love

Asteraceae

SE

5.8

40.5

76

0.8

1B.2

1Perennialforb

Packeragreenei

(A.Gray)W.A.Weber

&A.Love

Asteraceae

BE

5.3

32

66

1.6

11

Perennialforb

Packeralayneae

(Greene)

W.A.Weber

&A.Love

Asteraceae

BE

4.9

29.5

65

1.3

1Perennialforb

Packeramacounii

(Greene)

W.A.Weber

&A.Love

Asteraceae

BE

5.1

20.5

46

24.3

1Perennialforb

Pentachaetabellidiflora

Greene

Asteraceae

WI

2.4

7.25

33

1.3

1B.1

1Annualforb

Pyrrocomaracemosa

var.congesta

(Nutt.)Torr.&

A.Gray

Asteraceae

SE

618

36

02B.3

1Perennialforb

Pyrrocomaracemosa

var.pinetorum

(Nutt.)Torr.&

A.Gray

Asteraceae

BE/SI

416

44.5

2.4

4.2

1Perennialforb

Pyrrocomaracemosa

var.racemosa

(Nutt.)Torr.&

A.Gray

Asteraceae

WI

1.7

53

12.1

11

1Perennialforb

Raillardella

pringlei

Greene

Asteraceae

SE

630

56

01B.2

1Perennialforb

Rigiopappusleptocladus

A.Gray

Asteraceae

WI

1.9

7.5

42

1.3

11

11

1Annualforb

Rudbeckia

glaucescens

Eastw.

Asteraceae

BE

5.3

21

46

1.5

11

Perennialforb

Rudbeckia

klamathensis

P.B.Cox&

Urbatsch

Asteraceae

BE

510

25

-1

Perennialforb

Sericocarpusoregonensis

Nutt.

Asteraceae

WI/IN

1.1

3.25

31

0.9

11

1Perennialforb

Solidagoguiradonis

A.Gray

Asteraceae

SE

618

36

04.3

1Perennialforb

Solidagomultiradiata

Aiton

Asteraceae

WI/IN

1.1

2.1

21.1

-1

1Perennialforb

Stebbinsoserisdecipiens

(K.L.Chambers)

K.L.

Chambers

Asteraceae

WI

1.8

5.5

32

0.6

1B.2

11

Annualforb

Wyethia

bolanderi

(A.Gray)W.A.Weber

Asteraceae

WI

1.5

32

1.5

-1

Perennialforb

Berberisaquifolium

var.aquifolium

Pursh

Berberidaceae

WI

1.6

4.75

31

1.2

11

1Shrub

Berberisaquifolium

var.repens

Pursh

Berberidaceae

WI

1.7

53

11.2

11

1Shrub

Vancouveriachrysantha

Greene

Berberidaceae

SE

618

36

04.3

11

Perennialforb

(rhiz.)

Vancouveriaplanipetala

Calloni

Berberidaceae

WI

1.7

53

11.2

11

11

Perennialforb

(rhiz.)

Cryptanthadissita

I.M.Johnst.

Boraginaceae

BE/SI

4.4

17.5

44.5

2.1

1B.2

1Annualforb

Cryptanthaexcavata

Brandegee

Boraginaceae

WI

1.5

32

1.5

-1B.1

1Annualforb

Cryptanthaflaccida

(Lehm.)Greene

Boraginaceae

WI

1.6

4.75

32

0.7

11

11

1Annualforb

Cryptanthahispidula

Brand

Boraginaceae

SE

624

46

0CBR

11

Annualforb

Cryptanthainterm

edia

var.interm

edia

(A.Gray)Greene

Boraginaceae

WI/IN

1.4

4.1

31

1.5

11

11

1Annualforb

Cryptanthamariposae

I.M.Johnst.

Boraginaceae

SE

618

36

01B.3

1Annualforb

Cryptanthamilobakeri

I.M.Johnston

Boraginaceae

SI

3.3

9.75

33

2.6

11

1Annualforb

Emmenanthependuliflora

var.penduliflora

Benth.

Boraginaceae

WI

1.8

74

20.5

11

11

Annualforb

94 [Vol. 67MADRONO


APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Emmenanthependuliflora

var.rosea

Benth.

Boraginaceae

BE/SI

4.3

17

44.5

1.7

11

Annualforb

Hackelia

bella

(J.F.Macbr.)I.M.Johnst.

Boraginaceae

WI/IN

1.3

43

11.5

CBR

11

Perennialforb

Howellanthusdalesianus

(J.T.H

owell)Walden

&R.Patt.

Boraginaceae

SE

630

56

04.3

1Perennialforb

Pectocaryapusilla

(A.DC.)A.Gray

Boraginaceae

WI/IN

1.3

2.5

21.3

-1

11

11

Annualforb

Phaceliabreweri

A.Gray

Boraginaceae

SE

5.5

11

25.5

-1

1Annualforb

Phaceliacalifornica

Cham.

Boraginaceae

WI/IN

1.4

4.25

31

1.4

11

1Perennialforb

Phaceliacorymbosa

Jeps.

Boraginaceae

SE

5.5

33

66

0.8

11

Perennialforb

Phaceliadistans

Benth.

Boraginaceae

WI/IN

1.1

2.1

21.1

-1

11

Annualforb

Phaceliadivaricata

(Benth.)A.Gray

Boraginaceae

WI

2.3

73

31.2

11

1Annualforb

Phaceliaegena

(Brand)J.

T.Howell

Boraginaceae

WI

2.1

6.25

33

1.6

11

11

1Perennialforb

Phaceliagreenei

J.T.Howell

Boraginaceae

SE

636

66

01B.2

1Annualforb

Phaceliaim

bricata

var.im

bricata

Greene

Boraginaceae

WI

1.7

53

11.2

11

11

Perennialforb

Phacelialeonis

J.T.Howell

Boraginaceae

BE/SI

3.9

27.5

74

1.1

1B.3

1Annualforb

Phaceliaphacelioides

(Benth.)Brand

Boraginaceae

BE/SI

4.2

12.5

36

3.2

1B.2

11

Annualforb

Phaceliapringlei

A.Gray

Boraginaceae

BE/SI

412

34

1CBR

1Annualforb

Phaceliapurpusii

Brandegee

Boraginaceae

WI/IN

1.1

2.1

21.1

-1

Annualforb

Arabismcdonaldiana

Eastw.

Brassicaceae

BE

5.4

32.5

66

11B.1

11

Perennialforb

Arabisoregana

Rollins

Brassicaceae

BE/SI

3.8

11.5

33

2.1

4.3

11

Perennialforb

Boecheraconstancei

(Rollins)

Al-Shehbaz

Brassicaceae

SE

5.9

41.5

76

0.4

1B.1

1Perennialforb

Boecherakoehleri

(Howell)Al-Shehbaz

Brassicaceae

BE

5.3

21

45.5

11B.3

1Perennialforb

Boecherarollei

(Rollins)

Al-Shehbaz

Brassicaceae

SE

612

26

-1B.1

1Perennialforb

Boecheraserpenticola

Windham

&Al-Shehbaz

Brassicaceae

BE

5.3

16

36

1.2

1B.2

11

Perennialforb

Boecherasubpinnatifida

(S.Watson)Al-Shehbaz

Brassicaceae

WI

2.3

94

2.5

11

1Perennialforb

Cardaminenuttallii

Greene

Brassicaceae

WI

22

11

11

Perennialforb

(rhiz.)

Cardaminepachystigmavar.dissectifolia

(S.Watson)Rollins

Brassicaceae

SE

5.5

60

11

60.9

1B.2

11

Perennialforb

(rhiz.)

Caulanthusflavescens

(Hook.)Payson

Brassicaceae

WI

2.3

9.25

42.5

1.6

11

1Annualforb

Drabaaureola

S.Watson

Brassicaceae

SI

2.7

83

30.6

1B.3

1Perennialforb

Drabacarnosula

O.E.Schulz

Brassicaceae

SE

614

46

01B.3

1Perennialforb

Drabahowellii

S.Watson

Brassicaceae

WI/IN

1.4

4.25

31

1.4

4.3

1Perennialforb

Erysimum

franciscanum

Rossbach

Brassicaceae

SI

39

33

04.2

11

1Perennialforb

Noccaea

fendlerissp.californica

(A.Gray)Holub

Brassicaceae

SE

630

56

01B.1

1Perennialforb

Noccaea

fendlerissp.glauca

(A.Gray)Holub

Brassicaceae

BE/SI

4.4

22

54

1.5

11

Perennialforb

Streptanthusbarbatus

S.Watson

Brassicaceae

SE

5.6

28

56

0.5

CBR

1Perennialforb

Streptanthusbarbiger

Greene

Brassicaceae

SE

624

46

04.2

1Annualforb

Streptanthusbatrachopus

J.L.Morrison

Brassicaceae

SE

624

46

01B.3

11

Annualforb

Streptanthusbrachiatus

F.W.Hoffm.

Brassicaceae

SE

5.9

23.5

86

0.5

1Annual,Perennialforb

Streptanthusbreweri

A.Gray

Brassicaceae

SE

5.7

40

76

0.8

11

11

Annualforb

Streptanthusdrepanoides

Kruckeb.&

J.L.Morrison

Brassicaceae

SE

636

66

04.3

11

1Annualforb

Streptanthusglandulosusssp.albidus

Hook.

Brassicaceae

BE

5.3

31.5

66

1.3

11

Annualforb

Streptanthusglandulosusssp.glandulosus

Hook.

Brassicaceae

BE/SI

3.7

40.3

11

31.7

11

1Annualforb

Streptanthusglandulosusssp.hoffmanii

Hook.

Brassicaceae

SI

33

13

-1B.3

1Annualforb

Streptanthusglandulosusssp.niger

Hook.

Brassicaceae

SE

630

56

01B.1

11

Annualforb

Streptanthusglandulosusssp.pulchellus

Hook.

Brassicaceae

BE

4.9

24.5

56

1.8

11

Annualforb

Streptanthusglandulosusssp.secundus

Hook.

Brassicaceae

SI

3.3

20

63

1.5

11

Annualforb

Streptanthusglandulosusssp.sonomensis

Hook.

Brassicaceae

SE

612

26

-1

Annualforb

Streptanthushesperidis

Jeps.

Brassicaceae

SE

624

46

01B.2

1Annualforb

Streptanthushowellii

S.Watson

Brassicaceae

SE

630

56

01B.2

1Perennialforb

Streptanthusinsignisssp.insignis

Jeps.

Brassicaceae

BE/SI

420

54

2.4

1Annualforb

Streptanthusinsignisssp.lyonii

Jeps.

Brassicaceae

SI

3.3

16.5

52

2.7

1B.2

1Annualforb

Streptanthusmorrisonii

F.W.Hoffm.

Brassicaceae

SE

6110

18

60

CBR


Streptanthuspolygaloides

A.Gray

Brassicaceae

SE

5.7

28.5

56

0.9

1Annualforb

Streptanthusvernalis

R.O’D

onnell&

R.W.

Dolan

Brassicaceae

SE

612

26

-1B.2

1Annualforb



APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Streptanthusvimineus

(Greene)

Al-Shehbaz&

D.W

.Taylor

Brassicaceae

SE

612

26

-1

Annualforb

Thelypodium

brachycarpum

Torr.

Brassicaceae

SI

3.3

10

33

0.6

4.2

11


Campanula

angustiflora

Eastw.

Campanulaceae

BE/SI

3.9

19.3

54

2.4

11

Annualforb

Campanula

exigua

Rattan

Campanulaceae

BE/SI

3.9

19.5

54

1.5

1B.2

11

Annualforb

Campanula

griffinii

Morin

Campanulaceae

SE

618

36

0CBR

11

1Annualforb

Campanula

rotundifolia

L.

Campanulaceae

BE

515

36

1.7

1Perennialforb

Campanula

scabrella

Engelm.

Campanulaceae

SI

2.5

10

42.5

1.3

4.3

1Perennialforb

(rhiz.)

Campanula

sharsmithiae

Morin

Campanulaceae

SE

618

36

01B.2

11

Annualforb

Campanula

wilkinsiana

Greene

Campanulaceae

WI/IN

15

50

1.7

1B.2

1Perennialforb

(rhiz.)

Githopsisdiffusa

ssp.candida

A.Gray

Campanulaceae

WI/IN

12

21

-CBR

1Annualforb

Githopsispulchella

ssp.campestris

Vatke

Campanulaceae

WI

1.6

3.25

21.6

-1

Annualforb

Githopsispulchella

ssp.serpentinicola

Vatke

Campanulaceae

BE

5.3

21

45.5

14.3

1Annualforb

Githopsispulchella

var.glabra

Vatke

Campanulaceae

BE/SI

3.8

19

53

21

Annualforb

Githopsispulchella

var.pulchella

Vatke

Campanulaceae

WI/IN

11

11

-1

Annualforb

Nem

acladusmontanus

Greene

Campanulaceae

SE

618

36

0CBR

11

1Annualforb

Cerastium

arvense

ssp.strictum

L.

Caryophyllaceae

WI

2.1

8.5

40.9

2.6

11

11

1Perennialforb

Eremogonekingiivar.glabrescens

(S.Watson)Ikonn.

Caryophyllaceae

WI/IN

1.4

4.1

32

1.1

1Perennialforb

Minuartia

californica

(A.Gray)Mattf.

Caryophyllaceae

WI

1.7

53

21.5

11

11

1Annualforb

Minuartia

cism

ontana

Meinke&

Zika

Caryophyllaceae

WI

1.8

3.5

21.8

-1

11

11


Minuartia

decumbens

T.W.Nelson&

J.P.Nelson

Caryophyllaceae

SE

624

46

01

1Perennialforb

Minuartia

douglasii

(Fenzl

exTorr.&

A.Gray)

Mattf.

Caryophyllaceae

SI

315

53

0.7

11

11

1Annualforb

Minuartia

howellii

(S.Watson)Mattf.

Caryophyllaceae

SE

5.7

28.5

56

0.9

1Perennialforb

Minuartia

nuttalliivar.gregaria

(Pax)Briq.

Caryophyllaceae

SI

3.2

16

53

1.9

11

Perennialforb

Minuartia

rosei

(Maguire&

Barneby)

McN

eill

Caryophyllaceae

SE

630

56

01

1Perennialforb

Minuartia

stolonifera

T.W.Nelson&

J.P.Nelson

Caryophyllaceae

SE

630

56

01

Perennialforb

Moehringia

macrophylla

(Hook.)Fenzl

Caryophyllaceae

SI

2.7

83

30.6

11

11

1Perennialforb

Sileneantirrhina

L.

Caryophyllaceae

WI/IN

1.1

3.25

31

0.9

11

11

1Annualforb

Silenecampanulata

ssp.campanulata

S.Watson

Caryophyllaceae

BE

5.3

31.5

65.5

14.2

11

Perennialforb

Silenecampanulata

ssp.glandulosa

S.Watson

Caryophyllaceae

BE/SI

3.8

19

53

1.3

11

Perennialforb

Silenegrayi

S.Watson

Caryophyllaceae

WI

1.8

5.5

32

1.3

1Perennialforb

Silenehookeri

Nutt.

Caryophyllaceae

BE/SI

3.5

72

3.5

-1

1Perennialforb

Silenesalm

onacea

T.W.Nelson,J.

P.

Nelson,&

S.A.Erw

inCaryophyllaceae

SI

36

23

-1B.2

1Perennialforb

Sileneserpentinicola

T.W.Nelson&

J.P.Nelson

Caryophyllaceae

SE

66

16

-1

Perennialforb

(rhiz.)

Crocanthem

um

scoparium

(Nutt.)Millsp.

Cistaceae

WI/IN

12

21

-1

Shrub

Calystegia

collinassp.collina

(Greene)

Brummitt

Convolvulaceae

BE

4.7

33

76

1.6

11

Perennialforb

Calystegia

collinassp.oxyphylla

(Greene)

Brummitt

Convolvulaceae

SE

5.6

33.5

66

1.2

4.2

1Perennialforb

Calystegia

collinassp.tridactylosa

(Greene)

Brummitt

Convolvulaceae

BE

4.5

18

44.5

1.7

1B.2

1Perennialforb

Calystegia

collinassp.venusta

(Greene)

Brummitt

Convolvulaceae

BE

4.9

24.5

55

1.3

4.3

1Perennialforb

Calystegia

malacophylla

(Greene)

Munz

Convolvulaceae

WI

1.5

4.5

31

1.3

11

1Perennialforb

Calystegia

vanzuukiae

Brummitt&

Namoff

Convolvulaceae

BE

515

35

01B.3

1Perennialforb

Convolvulussimulans

L.M.Perry

Convolvulaceae

BE/SI

3.7

14.8

44

2.4

4.2

11

Annualforb

Dudleyaabramsiissp.bettinae

Rose

Crassulaceae

SE

618

36

01B.2

1Perennialforb

Dudleyaabramsiissp.murina

Rose

Crassulaceae

SE

618

36

01B.3

1Perennialforb

Dudleyaabramsiissp.setchellii

Rose

Crassulaceae

SE

624

46

01B.1

11

Perennialforb

Dudleyablochmaniaessp.blochmaniae

(Eastw.)Moran

Crassulaceae

SI

3.2

9.5

33

01B.1

1Perennialforb

Sedella

pentandra

H.Sharsm.

Crassulaceae

WI

28.1

41.5

2.1

11

1Annualforb

Sedella

pumila

(Benth.)Britton&

Rose

Crassulaceae

WI

1.7

5.1

32

1.5

11

Annualforb

Sedum

albomarginatum

R.T.Clausen

Crassulaceae

SE

5.9

47

86

0.4

1B.2

1Perennialforb

Sedum

citrinum

Zika

Crassulaceae

SE

618

36

01B.2

1Perennialforb

Sedum

eastwoodiae

(Britton)A.Berger

Crassulaceae

SE

630

56

01B.2

1Perennialforb

96 [Vol. 67MADRONO


APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Sedum

flavidum

B.L.Wilson&

Zika

Crassulaceae

SI

318

63

0.6

4.3

11

Perennialforb

Sedum

kiersteadiae

B.L.Wilson&

R.E.

Brainerd

Crassulaceae

SI

3.3

10

33

0.6

1B.2

1Perennialforb

Sedum

laxum

ssp.heckneri

(Britton)A.Berger

Crassulaceae

SI

3.3

10

33

0.6

4.3

11

Perennialforb

Sedum

laxum

ssp.laxum

(Britton)A.Berger

Crassulaceae

BE

4.8

23

55

0.8

11

Perennialforb

Sedum

patens

Zika

Crassulaceae

SE

66

16

-1

Perennialforb

Sedum

radiatum

S.Watson

Crassulaceae

WI

26

32

21

11

11

Annualforb

Sedum

rubiginosum

Zika&

B.L.Wilson

Crassulaceae

SE

66

16

-1

Perennialforb

Calocedrusdecurrens

(Torr.)Florin

Cupressaceae

SI

39

33

01

11

11

Tree

Chamaecyparislawsoniana

(A.Murraybis)Parl.

Cupressaceae

SI

315

53

0.7

CBR

11

Tree

Hesperocyparisbakeri

(Jeps.)Bartel

Cupressaceae

SI

2.6

13

53

0.5

4.2

11

Tree

Hesperocyparismacnabiana

(A.Murraybis)Bartel

Cupressaceae

BE

4.7

28

64.5

1.2

11

Tree

Hesperocyparissargentii

(Jeps.)Bartel

Cupressaceae

BE

4.9

34

75

1.2

11

1Tree

Juniperuscommunisvar.jackii

L.

Cupressaceae

BE/SI

48

24

-CBR

1Shrub

Calliscirpuscriniger

(A.Gray)C.N

.Gilmour,

J.R.Starr,&

Naczi

Cyperaceae

WI

1.7

53

12.1

11

1Perennialgram.

Carexbrainerdii

Mack.

Cyperaceae

WI/IN

1.4

4.25

32

11

11

1Perennialgram.(rhiz.)

Carexconcinnoides

Mack.

Cyperaceae

BE/SI

48

24

-1


Carexhirtissim

aW.Boott

Cyperaceae

WI

1.8

3.5

21.8

-1

1Perennialgram.(cesp.)

Carexklamathensis

B.L.Wilson&

Janew

ay

Cyperaceae

SE

612

26

-1B.2

11

Perennialgram.(rhiz.)

Carexmendocinensis

Olney

exW.Boott

Cyperaceae

BE/SI

3.8

23

63.5

1.2

11


Carexobispoensis

Stacey

Cyperaceae

BE

4.9

24.5

56

1.6

1B.2


Carexscabriuscula

Mack.

Cyperaceae

SE

612

26

-4.3

11


Carexserpenticola

Zika

Cyperaceae

SE

5.5

11

25.5

-1


Carexserratodens

W.Boott

Cyperaceae

BE

4.9

39

85

1.1

11

11

Perennialgram.(cesp.)

Carexspissa

L.H.Bailey

Cyperaceae

SI

2.8

8.25

32

2.9


Carexxerophila

Janew

ay&

Zika

Cyperaceae

WI

1.7

53

20.6


Polystichum

lemmonii

Underw.

Dryopteridaceae

SE

624

46

01

11

Perennialforb

(rhiz.)

Polystichum

scopulinum

(D.C.Eaton)Maxon

Dryopteridaceae

WI

1.7

5.1

32

1.5

11

1Perennialforb

(rhiz.)

Arctostaphylosbakerissp.bakeri

Eastw.

Ericaceae

SE

5.5

27.5

56

1.3

1B.1

1Shrub

Arctostaphylosbakerissp.sublaevis

Eastw.

Ericaceae

SE

612

26

-1B.2

1Shrub

Arctostaphyloscanescensssp.sonomensis

Eastw.

Ericaceae

SI

2.5

12.5

53

1.5

CBR

11

Shrub

Arctostaphylosfranciscana

Eastw.

Ericaceae

SE

618

36

01B.1

11

Shrub

Arctostaphyloshispidula

Howell

Ericaceae

BE

4.5

22.5

54

1.1

4.2

11

Shrub

Arctostaphylosklamathensis

S.W.Edwards,Keeler-

Wolf&

W.Knight

Ericaceae

BE/SI

3.9

19.5

54

1.8

1B.2

1Shrub

Arctostaphylosmontanassp.montana

Eastw.

Ericaceae

BE

4.9

19.5

44.5

11B.3

11

Shrub

Arctostaphylosmontanassp.ravenii

Eastw.

Ericaceae

SE

618

36

01B.1

11

Shrub

Arctostaphylosnortensis

(P.V.Wells)P.V.Wells

Ericaceae

SI

2.8

5.5

22.5

-4.3

1Shrub

Arctostaphylosobispoensis

Eastw.

Ericaceae

SE

5.7

28.5

56

0.9

4.3

1Shrub

Arctostaphylosstanfordianassp.raichei

Parry

Ericaceae

SI

2.6

10.5

43

1.7

1B.1

1Shrub

Arctostaphylosviscidassp.pulchella

Parry

Ericaceae

BE

525

55

11

1Shrub

Arctostaphylosviscidassp.viscida

Parry

Ericaceae

WI

2.2

10.8

53

1.2

11

Shrub

Pyrola

dentata

Sm.

Ericaceae

WI

26

33

1.7

11

Perennialforb

(rhiz.)

Vaccinium

coccineum

Piper

Ericaceae

BE/SI

3.5

3.5

13

-3.3

1Shrub

Acm

isponjunceusvar.junceus

(Benth.)Brouillet

Fabaceae

WI

1.5

32

1.5

-1

Perennialforb

Astragalusbreweri

A.Gray

Fabaceae

SI

3.2

15.8

53

24.2

11

Annualforb

Astragalusclaranus

Jeps.

Fabaceae

SI

36

23

-1

Annualforb

Astragalusclevelandii

Greene

Fabaceae

SE

624

46

04.3

11

Perennialforb

Astragaluscurtipes

A.Gray

Fabaceae

WI

1.8

3.5

21.8

-CBR

1Perennialforb

Astragalusmacrodon

(Hook.&

Arn.)A.Gray

Fabaceae

WI/IN

1.3

3.75

31

0.7

4.3

1Perennialforb

Astragalusrattaniivar.jepsonianus

A.Gray

Fabaceae

BE/SI

4.3

25.5

64

1.2

1B.2

1Annualforb

Astragaluswhitneyivar.siskiyouensis

A.Gray

Fabaceae

BE

4.6

23

55

1.1

11

Perennialforb

Hoitastrobilina

(Hook.&

Arn.)Rydb.

Fabaceae

SI

2.5

52

2.5

-1B.1

1Perennialforb



APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Lathyrusbiflorus

T.W.Nelson&

J.P.Nelson

Fabaceae

SE

624

46

01B.1

1Perennialforb

Lathyrusdelnorticus

C.L.Hitchc.

Fabaceae

BE

5.3

10.5

25

-4.3

1Perennialforb

Lathyrusvestitusvar.vestitus

Nutt.

Fabaceae

WI

1.8

7.2

40.6

2.8

11

1Perennialforb

Lupinusconstancei

T.W.Nelson&

J.P.Nelson

Fabaceae

SE

624

46

01B.1

1Perennialforb

Lupinuscroceus

Eastw.

Fabaceae

BE/SI

412

34

01

Perennialforb

Lupinuslapidicola

A.Heller

Fabaceae

SI

315

53

31

1Perennialforb

Lupinusonustus

S.Watson

Fabaceae

SI

3.1

15.3

53

2.9

11

Perennialforb

Lupinusspectabilis

Hoover

Fabaceae

SE

624

46

01B.2

1Annualforb

Pediomelum

californicum

(S.Watson)Rydb.

Fabaceae

BE/SI

4.4

21.8

56

2.4

11

1Perennialforb

Trifolium

amoenum

Greene

Fabaceae

WI/IN

1.3

2.5

21

-1B.1

11

Annualforb

Trifolium

fucatum

Lindl.

Fabaceae

WI/IN

1.3

43

10.6

11

Annualforb

Trifolium

gracilentum

Torr.&

A.Gray

Fabaceae

WI/IN

13.1

31

11

11

11

Annualforb

Trifolium

longipes

ssp.elmeri

Nutt.

Fabaceae

BE

5.3

21

46

1.5

11

Perennialforb

Trifolium

longipes

ssp.oreganum

Nutt.

Fabaceae

BE/SI

412

34

21

1Perennialforb

Trifolium

microcephalum

Pursh

Fabaceae

WI/IN

1.4

75

11.1

11

1Annualforb

Trifolium

willdenovii

Spreng.

Fabaceae

WI/IN

1.3

43

10.6

11

11

1Annualforb

Notholithocarpusdensiflorusvar.

echinoides

(Hook.&

Arn.)Manos,

C.H

.Cannon,&

S.Oh

Fagaceae

SI

2.5

12.3

51

2.5

11

Shrub

Quercusdurata

var.durata

Jeps.

Fagaceae

SE

5.8

40.5

76

0.8

11

11

Shrub

Quercusvacciniifolia

Kellogg

Fagaceae

SI

2.5

12.3

52

2.2

11

1Shrub

Garryabuxifolia

A.Gray

Garryaceae

SE

5.8

29

56

0.4

11

Shrub

Garryacongdonii

Eastw.

Garryaceae

BE

530

65.5

1.3

11

11

Shrub

Gentianasetigera

A.Gray

Gentianaceae

SE

5.8

17.5

36

0.3

1B.2

11

Perennialforb

Zeltneratrichantha

(Griseb.)G.Mans.

Gentianaceae

SE

5.5

11

25.5

-1

1Annualforb

Irisbracteata

S.Watson

Iridaceae

SE

5.8

11.5

25.5

-3.3

1Perennialforb

(rhiz.)

Irisinnominata

L.F.Hend.

Iridaceae

SE

5.7

17

36

0.6

4.3

1Perennialforb

(rhiz.)

Irismacrosiphon

Torr.

Iridaceae

WI/IN

1.1

3.25

31

0.9

11

1Perennialforb

(rhiz.)

Iristenuissimassp.purdyiform

isDykes

Iridaceae

WI

1.5

32

1.5

-1

1Perennialforb

(rhiz.)

Iristhompsonii

R.C.Foster

Iridaceae

BE

4.5

92

4.5

-4.3

11

Perennialforb

Acanthominthaduttonii

(Abrams)

Jokerst

Lamiaceae

SE

630

56

01B.1

11

Annualforb

Acanthominthailicifolia

(A.Gray)A.Gray

Lamiaceae

WI/IN

1.3

33

01.7

1B.1

1Annualforb

Acanthominthalanceolata

Curran

Lamiaceae

SI

3.4

16.8

53

2.1

4.2

11

Annualforb

Acanthominthaobovata

ssp.obovata

Jeps.

Lamiaceae

BE/SI

3.5

10.5

33

2.5

4.2

1Annualforb

Monardella

douglasii

Benth.

Lamiaceae

WI/IN

12

21

-1

11

Annualforb

Monardella

follettii

(Jeps.)Jokerst

Lamiaceae

SE

5.8

34.5

66

0.8

1B.2

1Shrub

Monardella

palm

eri

A.Gray

Lamiaceae

BE

4.8

28.5

66

2.2

1B.2

1Perennialforb

(rhiz.)

Monardella

purpurea

Howell

Lamiaceae

SE

624

46

01

1Perennialforb

Monardella

sheltonii

Torr.

Lamiaceae

SI

318

63

1.7

11

1Perennialforb

Monardella

stebbinsii

Hardham

&Bartel

Lamiaceae

SE

630

56

01B.2

1Perennialforb

(rhiz.)

Monardella

villosa

ssp.villosa

Benth.

Lamiaceae

SE

624

46

01

Perennialforb

(rhiz.)

Monardella

viridis

Jeps.

Lamiaceae

WI/IN

12

21

-4.3

1Perennialforb

Salvia

sonomensis

Greene

Lamiaceae

WI

1.6

9.5

61.5

1.3

11

11

1Shrub

Scutellariaantirrhinoides

Benth.

Lamiaceae

WI

2.3

11.5

53

1.5

11

1Perennialforb

Stachyspycnantha

Benth.

Lamiaceae

WI

2.2

11

51

2.4

CBR

11

11

1Perennialforb

Trichostem

alaxum

A.Gray

Lamiaceae

BE/SI

416

44.5

2.4

11

Annualforb

Trichostem

arubisepalum

Elm

erLamiaceae

SE

5.6

28

56

0.9

4.3

11

Annualforb

Pinguicula

macroceras

Link

Lentibulariaceae

SE

618

36

02B.2

1Perennialforb

(carn.)

Calochortusclavatusvar.clavatus

S.Watson

Liliaceae

BE

4.5

13.5

34

0.6

4.3

1Perennialforb

(bulb)

Calochortusclavatusvar.pallidus

S.Watson

Liliaceae

WI

1.6

3.1

2-

11

Perennialforb

(bulb)

Calochortuselegans

Pursh

Liliaceae

WI

24

22

-1

Perennialforb

(bulb)

Calochortusfimbriatus

H.P.McD

onald

Liliaceae

WI

1.5

4.5

31

1.3

1B.3

11

1Perennialforb

(bulb)

Calochortusfimbriatus

H.P.McD

onald

Liliaceae

WI/IN

13

30

1.7

1B.3

1Perennialforb

(bulb)

Calochortusnudus

S.Watson

Liliaceae

WI

2.1

8.5

42.5

1.2

11

Perennialforb

(bulb)

Calochortusobispoensis

Lem

mon

Liliaceae

BE

5.4

21.5

46

1.5

1B.2

1Perennialforb

(bulb)

98 [Vol. 67MADRONO


APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Calochortusraichei

Farw

ig&

V.Girard

Liliaceae

SE

618

36

01B.2

1Perennialforb

(bulb)

Calochortustiburonensis

A.J.

Hill

Liliaceae

SE

624

46

01B.1

11

Perennialforb

(bulb)

Calochortusumbellatus

Alph.Wood

Liliaceae

SI

2.9

14.5

53

1.1

4.2

11

Perennialforb

(bulb)

Calochortusuniflorus

Hook.&

Arn.

Liliaceae

WI

1.7

53

11.2

4.2

11

11

Perennialforb

(bulb)

Calochortusvestae

Purdy

Liliaceae

WI

26

32

11

1Perennialforb

(bulb)

Erythronium

californicum

Purdy

Liliaceae

SI

2.7

83

22.1

11

Perennialforb

(bulb)

Erythronium

citrinum

var.citrinum

S.Watson

Liliaceae

BE/SI

4.3

21.5

54

0.4

4.3

1Perennialforb

(bulb)

Erythronium

citrinum

var.roderickii

S.Watson

Liliaceae

BE

4.7

37.5

84.5

1.4

1B.3

1Perennialforb

(bulb)

Erythronium

helenae

Applegate

Liliaceae

BE

4.5

18

44.5

1.7

4.2

1Perennialforb

(bulb)

Erythronium

hendersonii

S.Watson

Liliaceae

SI

2.5

52

2.5

-2B.3

1Perennialforb

(bulb)

Erythronium

multiscapideum

(Kellogg)A.Nelson&

P.

B.Kenn.

Liliaceae

SI

315

52

1.7

1Perennialforb

(bulb)

Erythronium

purpurascens

S.Watson

Liliaceae

WI/IN

12

21

-1

Perennialforb

(bulb)

Erythronium

tuolumnense

Applegate

Liliaceae

SI

2.5

52

2.5

-1B.2

1Perennialforb

(bulb)

Fritillariaaffinis

(Schult.&

Schult.f.)Sealy

Liliaceae

WI

26

32

01

11

1Perennialforb

(bulb)

Fritillariaagrestis

Greene

Liliaceae

SI

2.7

13.3

52

1.6

4.2

11

11

Perennialforb

(bulb)

Fritillariabiflora

var.biflora

Lindl.

Liliaceae

WI

2.3

94

2.5

1.7

CBR

11

1Perennialforb

Fritillariabiflora

var.ineziana

Lindl.

Liliaceae

BE

5.4

21.5

46

1.5

1B.1

1Perennialforb

(bulb)

Fritillariaeastwoodiae

R.M.MacF

arl.

Liliaceae

WI

2.3

13.5

62

0.4

3.2

1Perennialforb

(bulb)

Fritillariafalcata

(Jeps.)D.E.Beetle

Liliaceae

SE

624

46

01B.2

11

Perennialforb

(bulb)

Fritillariaglauca

Greene

Liliaceae

BE/SI

4.3

17.3

45.5

2.7

4.2

11

Perennialforb

Fritillarialiliacea

Lindl.

Liliaceae

WI

1.8

74

1.5

1.1

1B.2

11

Perennialforb

(bulb)

Fritillariapluriflora

Torr.ex

Benth.

Liliaceae

WI

2.4

9.5

42.5

1.5

1B.2

11

Perennialforb

(bulb)

Fritillariapurdyi

Eastw.

Liliaceae

BE

4.5

31.5

74

1.8

4.3

11

Perennialforb

(bulb)

Fritillariarecurva

Benth.

Liliaceae

SI

2.7

83

2.5

1.9

11

1Perennialforb

(bulb)

Fritillariaviridea

Kellogg

Liliaceae

SE

618

36

01B.2

1Perennialforb

(bulb)

Lilium

bolanderi

S.Watson

Liliaceae

SE

618

36

04.2

1Perennialforb

(bulb)

Lilium

kelloggii

Purdy

Liliaceae

SI

2.5

10

42

1.9

4.3

11

Perennialforb

(bulb)

Lilium

pardalinum

ssp.vollmeri

Kellogg

Liliaceae

BE

515

35

14.3

1Perennialforb

(bulb)

Lilium

rubescens

S.Watson

Liliaceae

WI

29.75

52

1.4

4.2

11

1Perennialforb

(bulb)

Lilium

washingtonianum

ssp.purpurascens

Kellogg

Liliaceae

BE/SI

3.5

10.5

33

2.5

4.3

1Perennialforb

(bulb)

Prosartes

parvifolia

S.Watson

Liliaceae

BE

55

15

-1B.2

1Perennialforb

Hesperolinonadenophyllum

(A.Gray)Small

Linaceae

SE

5.7

28.5

56

0.9

1B.2

1Annualforb

Hesperolinonbicarpellatum

(H.Sharsm.)H.Sharsm.

Linaceae

SE

618

36

01B.2

1Annualforb

Hesperolinonbreweri

(A.Gray)Small

Linaceae

SI

2.5

10

42.5

1.5

1B.2

11

Annualforb

Hesperolinoncalifornicum

(Benth.)Small

Linaceae

SI

2.8

8.5

33

0.6

11

11

Annualforb

Hesperolinonclevelandii

(Greene)

Small

Linaceae

WI

28

42

1.8

11

Annualforb

Hesperolinoncongestum

(A.Gray)Small

Linaceae

SE

624

46

01B.1

11

Annualforb

Hesperolinondidymocarpum

H.Sharsm.

Linaceae

SE

618

36

01B.2

1Annualforb

Hesperolinondisjunctum

H.Sharsm.

Linaceae

SE

618

36

01

11

Annualforb

Hesperolinondrymarioides

(Curran)Small

Linaceae

SE

624

46

01B.2

1Annualforb

Hesperolinonmicranthum

(A.Gray)Small

Linaceae

WI

2.4

11.8

53

11

11

11

Annualforb

Hesperolinonsharsmithiae

R.O’D

onnell

Linaceae

SE

5.7

17

36

0.6

11

Annualforb

Hesperolinonspergulinum

(A.Gray)Small

Linaceae

BE

4.7

14

36

2.3

CBR

11

Annualforb

Hesperolinontehamense

H.Sharsm.

Linaceae

SE

5.8

34.5

66

0.8

1B.3

1Annualforb

Linum

lewisiivar.lewisii

Pursh

Linaceae

WI/IN

1.3

43

11.5

11

11

1Perennialforb

Fremontodendrondecumbens

R.M.Lloyd

Malvaceae

WI

28

41.5

2.4

1B.2

1Shrub

Sidalcea

asprellassp.nana

Greene

Malvaceae

SI

2.5

52.5

2.5

-1

11

Perennialforb

Sidalcea

diploscypha

(Torr.&

A.Gray)A.Gray

Malvaceae

SI

2.6

13

53

2.3

11

11

Annualforb

Sidalcea

hartwegii

A.Gray

Malvaceae

WI

1.6

4.75

32

0.7

11

Annualforb

Sidalcea

hickmaniissp.anomala

Greene

Malvaceae

SE

5.6

22.5

46

11B.2

1Perennialforb

Sidalcea

hickmaniissp.viridis

Greene

Malvaceae

SE

612

26

-1B.1

11

Perennialforb

Sidalcea

keckii

Wiggins

Malvaceae

SI

36

23

-1B.1

1Annualforb

Pseudotrillium

rivale

(S.Watson)S.BFarm

erMelanthiaceae

BE/SI

412

34

11

Perennialforb



APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Toxicoscordionfontanum

(Eastw.)Zomlefer&

Judd

Melanthiaceae

BE/SI

3.8

23

64

0.8

4.2

11

1Perennialforb

(bulb)

Toxicoscordionpaniculatum

(Nutt.)Rydb.

Melanthiaceae

WI

1.6

4.75

32

0.7

11

1Perennialforb

(bulb)

Xerophyllum

tenax

(Pursh)Nutt.

Melanthiaceae

WI

1.6

85

10.9

11

11

1Perennialforb

(rhiz.)

Calyptridium

quadripetalum

S.Watson

Montiaceae

BE

4.6

27.5

64

1.2

4.3

1Annualforb

Calyptridium

umbellatum

(Torr.)Greene

Montiaceae

WI/IN

1.3

43

11.5

11

11

Perennialforb

Claytonia

exiguassp.exigua

Torr.&

A.Gray

Montiaceae

SI

3.4

24

73

1.1

11

11

1Annualforb

Claytonia

exiguassp.glauca

Torr.&

A.Gray

Montiaceae

BE/SI

3.6

18

53

1.3

11

11

Annualforb

Claytonia

gypsophiloides

Fisch.&

C.A.Mey.

Montiaceae

SI

3.1

15.5

53

0.2

11

1Annualforb

Claytonia

saxosa

Brandegee

Montiaceae

BE/SI

4.4

21.8

55

2.2

CBR

11

Annualforb

Claytonia

serpenticola

T.R

.Stoughton

Montiaceae

BE/SI

44

14

-1

Perennialforb

Lew

isia

cantelovii

J.T.Howell

Montiaceae

WI/IN

16

61

1.1

1B.2

11

Perennialforb

Lew

isia

cotyledonvar.cotyledon

(S.Watson)B.L.Rob.

Montiaceae

WI

26

33

1.7

CBR

1Perennialforb

Lew

isia

cotyledonvar.heckneri

(S.Watson)B.L.Rob.

Montiaceae

WI/IN

14

40.5

1.4

1B.2

1Perennialforb

Lew

isia

cotyledonvar.howellii

(S.Watson)B.L.Rob.

Montiaceae

WI/IN

1.3

43

11.5

3.2

1Perennialforb

Lew

isia

leeana

(Porter)B.L.Rob.

Montiaceae

SI

36

23

-1

11

Perennialforb

Lew

isia

nevadensis

(A.Gray)B.L.Rob.

Montiaceae

WI

1.7

53

21.5

11

1Perennialforb

Lew

isia

oppositifolia

(S.Watson)B.L.Rob.

Montiaceae

BE

5.3

21

46

1.5

2B.2

1Perennialforb

Lew

isia

rediviva

Pursh

Montiaceae

WI/IN

1.4

75

11.1

11

11

1Perennialforb

Lew

isia

stebbinsii

Gankin

&W.R.Hildreth

Montiaceae

BE

4.7

14

36

2.3

1Perennialforb

Lew

isia

triphylla

(S.Watson)B.L.Rob.

Montiaceae

WI

1.7

53

21.5

11

1Perennialforb

Camissonia

benitensis

P.H.Raven

Onagraceae

SE

5.9

29.5

56

0.4

1B.1

1Annualforb

Camissonia

lacustris

P.H.Raven

Onagraceae

SI

39

33

31

1Annualforb

Clarkia

arcuata

(Kellogg)A.Nelson&

J.F.Macbr.

Onagraceae

WI

2.3

73

20.6

1Annualforb

Clarkia

bilobassp.biloba

(Durand)A.Nelson&

J.F.Macbr.

Onagraceae

WI/IN

1.4

2.75

21.4

-1

1Annualforb

Clarkia

breweri

(A.Gray)Greene

Onagraceae

BE/SI

3.8

11.5

33

2.1

4.2

11

Annualforb

Clarkia

franciscana

H.Lew

is&

P.H.Raven

Onagraceae

SE

624

46

01B.1

11

Annualforb

Clarkia

gracilisssp.albicaulis

(Piper)A.Nelson&

J.F.

Macbr.

Onagraceae

WI

2.2

6.5

32

11B.2

1Annualforb

Clarkia

gracilisssp.tracyi

(Piper)A.Nelson&

J.F.

Macbr.

Onagraceae

BE

525

55

14.2

1Annualforb

Epilobium

minutum

Lindl.

Onagraceae

WI

26

32

11

11

11

Annualforb

Epilobium

oreganum

Greene

Onagraceae

BE/SI

3.8

23

64

2.2

1B.2

11

Perennialforb

Epilobium

rigidum

Hausskn.

Onagraceae

BE

5.1

20.5

46

24.3

1Perennialforb

Epilobium

septentrionale

(D.D.Keck)R.N.

Bowman&

Hoch

Onagraceae

WI

24

2-

4.3

1Perennialforb

Epilobium

siskiyouense

(Munz)Hoch

&P.H.Raven

Onagraceae

SE

5.5

38.5

76

11B.3

1Perennialforb

Cypripedium

californicum

A.Gray

Orchidaceae

BE

4.5

40.5

94

1.3

4.2

11

11

Perennialforb

(bulb)

Cypripedium

fasciculatum

Kellogg

Orchidaceae

SI

2.5

12.3

52

1.6

4.2

11

11

Perennialforb

(bulb)

Piperia

candida

Ran

d.Morgan

&Ackerman

Orchidaceae

WI/IN

1.2

3.5

31

11B.2

11

1Perennialforb

(bulb)

Aphyllonvalidum

ssp.howellii

Jeps.

Orobanchaceae

SI

3.4

13.5

43

1.3

4.3

1Perennialforb

(paras.)

Castilleja

affinisssp.neglecta

Hook.&

Arn.

Orobanchaceae

SE

630

56

01

1Perennialforb

(hem

ipar.)

Castilleja

brevilobata

Piper

Orobanchaceae

SE

618

36

04.2

1Perennialforb

(hem

ipar.)

Castilleja

foliolosa

Hook.&

Arn.

Orobanchaceae

WI

2.3

94

2.5

11

11


Castilleja

miniata

ssp.elata

Douglasex

Hook.

Orobanchaceae

BE

4.6

27.5

64.5

1.4

1Perennialforb

(hem

ipar.)

Castilleja

minorssp.spiralis

(A.Gray)A.Gray

Orobanchaceae

SI

3.3

16.5

53

2.6

11

11

Annualforb

(hem

ipar.)

Castilleja

pruinosa

Fernald

Orobanchaceae

SI

3.2

15.8

53

1.9

11

1Perennialforb

(hem

ipar.)

Castilleja

rubicundula

ssp.lithospermoides

(Jeps.)T.I.Chuang&

Heckard

Orobanchaceae

WI

2.4

9.75

42

1.8

1Annualforb

(hem

ipar.)

Castilleja

rubicundula

ssp.rubicundula

(Jeps.)T.I.Chuang&

Heckard

Orobanchaceae

SE

5.6

28

56

0.9

1Annualforb

(hem

ipar.)

Cordylanthusnidularius

J.T.Howell

Orobanchaceae

SE

618

36

01B.1

11

Annualforb

(hem

ipar.)

Cordylanthuspilosusssp.pilosus

A.Gray

Orobanchaceae

SI

2.5

10

42.5

0.6

11

Annualforb

(hem

ipar.)

100 [Vol. 67MADRONO


APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Cordylanthuspringlei

A.Gray

Orobanchaceae

SE

5.6

28

56

0.9

CBR

1Annualforb

(hem

ipar.)

Cordylanthustenuisssp.brunneus

A.Gray

Orobanchaceae

BE

5.1

25.5

55

14.3

1Annualforb

(hem

ipar.)

Cordylanthustenuisssp.capillaris

A.Gray

Orobanchaceae

SE

624

46

01B.2

1Annualforb

(hem

ipar.)

Cordylanthustenuisssp.tenuis

A.Gray

Orobanchaceae

WI

2.3

94

20.5

1Annualforb

(hem

ipar.)

Cordylanthustenuisssp.viscidus

A.Gray

Orobanchaceae

BE

4.5

27

64.5

1.4

11

1Annualforb

(hem

ipar.)

Orthocarpuspachystachyus

A.Gray

Orobanchaceae

SE

618

36

01B.1

1Annualforb

(hem

ipar.)

Pedicularishowellii

A.Gray

Orobanchaceae

SI

2.5

7.5

33

1.2

4.3

1Perennialforb

(hem

ipar.)

Triphysariafloribunda

(Benth.)T.I.Chuang&

Heckard

Orobanchaceae

WI

2.3

6.75

32

1.9

1B.2

11

1Annualforb

Dicentraform

osa

ssp.oregana

(Haw.)Walp.

Papaveraceae

BE

5.2

31

66

1.3

4.2

1Perennialforb

Dicentrapauciflora

S.Watson

Papaveraceae

WI

2.2

6.5

33

1.4

CBR

11

Perennialforb

Ehrendorferia

chrysantha

(Hook.&

Arn.)Rylander

Papaveraceae

WI/IN

1.1

3.25

31

0.9

11

11

1Perennialforb

Eschscholzia

hypecoides

Benth.

Papaveraceae

SI

2.6

7.75

31

3.1

4.3

1Annualforb

Platystem

oncalifornicus

Benth.

Papaveraceae

WI

1.7

53

20.6

11

11

1Annualforb

Parnassia

cirrata

var.interm

edia

Piper

Parnassiaceae

BE/SI

3.5

72

3.5

-1

Perennialforb

Parnassia

palustris

L.

Parnassiaceae

WI

26

32

01

11

1Perennialforb

Diplacusdouglasii

(Benth.)G.L.Nesom

Phrymaceae

SI

2.7

13.5

53

0.5

11

11

1Annualforb

Diplacuslayneae

(Greene)

Jeps.

Phrymaceae

SI

2.9

14.3

53

1.2

11

1Annualforb

Erythrantheglaucescens

(Greene)

G.L.Nesom

Phrymaceae

BE/SI

3.8

18.8

54

2.1

1Annualforb

Erythranthelinearifolia

Benth.

Phrymaceae

BE/SI

416

44.5

2.4

1Perennialforb

(rhiz.)

Erythranthenudata

(Curranex

Greene)

G.L.

Nesom

Phrymaceae

SE

5.6

33.5

66

1.2

1Annualforb

Erythranthepercaulis

G.L.Nesom

Phrymaceae

SE

618

36

01B.1

1Annualforb

Erythranthetrinitiensis

G.L.Nesom

Phrymaceae

SE

618

36

01B.3

11

Annualforb

Erythranthewillisii

G.L.Nesom

Phrymaceae

SE

5.5

11

25.5

-1

Perennialforb

Picea

breweriana

S.Watson

Pinaceae

WI

2.2

6.5

33

1.4

CBR

1Tree

Pinusattenuata

Lem

mon

Pinaceae

SI

2.5

12.6

53

2.4

11

11

1Tree

Pinusbalfourianassp.balfouriana

Grev.&

Balf.

Pinaceae

BE/SI

4.3

26

64

1.5

1Tree

Pinuscoulteri

D.Don

Pinaceae

WI/IN

1.3

43

11.5

11

Tree

Pinusjeffreyi

Grev.&

Balf.

Pinaceae

SI

2.7

83

30.6

11

11

Tree

Pinussabiniana

D.Don

Pinaceae

WI/IN

1.4

4.25

31

1.4

11

11

1Tree

Antirrhinum

cornutum

Benth.

Plantaginaceae

WI

2.2

11

52

0.8

CBR

1Annualforb

Antirrhinum

leptaleum

A.Gray

Plantaginaceae

WI

1.6

3.1

21.6

-1

Annualforb

Antirrhinum

subcordatum

A.Gray

Plantaginaceae

BE/SI

4.3

21.5

54

1.8

4.3

1Annualforb

Antirrhinum

vexillocalyculatum

Kellogg

Plantaginaceae

SI

2.5

20

82.5

1.3

11

11

1Annualforb

Antirrhinum

virga

A.Gray

Plantaginaceae

SI

2.8

8.5

33

0.6

4.3

1Perennialforb

Collinsiagreenei

A.Gray

Plantaginaceae

BE

5.2

31

66

1.3

CBR

11

Annualforb

Collinsiamulticolor

Lindl.&

Paxton

Plantaginaceae

WI/IN

1.1

2.25

21.1

-1B.2

11

Annualforb

Collinsiasparsiflora

Fisch.&

C.A.Mey.

Plantaginaceae

WI

1.7

53

11.2

11

11

1Annualforb

Keckiellalemmonii

(A.Gray)Straw

Plantaginaceae

WI/IN

1.1

3.25

31

0.9

11

1Shrub

Penstem

onazureusvar.azureus

Benth.

Plantaginaceae

SI

2.7

83

30.6

11

1Perennialforb

Penstem

onfiliform

is(D

.D.Keck)D.D.Keck

Plantaginaceae

BE

530

65.5

1.3

1B.3

1Perennialforb

Penstem

onparvulus

(A.Gray)Krautter

Plantaginaceae

BE/SI

3.7

11

34

0.6

11

Perennialforb

Penstem

onpurpusii

Brandegee

Plantaginaceae

SI

2.8

11

42

2.4

CBR

11

Perennialforb

Plantagoerecta

E.Morris

Plantaginaceae

WI/IN

13

31

11

11

11

Annualforb

Veronicacopelandii

Eastw.

Plantaginaceae

SE

634

46

04.3

1Perennialforb

Agrostismicrophylla

Steud.

Poaceae

WI/IN

1.1

4.25

41.1

1.1

11

1Annualgram.

Bromuslaevipes

Shear

Poaceae

WI

1.7

53

20.6

11

11


Calamagrostisfoliosa

Kearney

Poaceae

WI

1.7

53

21.5

4.2


Calamagrostisophitidis

(J.T.Howell)Nygren

Poaceae

SE

624

46

04.3

11


Calamagrostisstrictassp.inexpansa

(Tim

m)Koeler

Poaceae

WI

1.5

32

1.5

-1

11

11


Danthonia

californica

Bol.

Poaceae

SI

3.3

13

43

2.2

11

11


Elymustrachycaulusssp.trachycaulus

(Link)Shinners

Poaceae

WI

1.6

3.1

21.6

-1

11

11


Festuca

californica

Vasey

Poaceae

WI

2.4

11.8

52

1.6

11

11




APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Festuca

idahoensis

Elm

erPoaceae

WI/IN

1.3

5.25

41

1.2

11

11


Hordeum

brachyantherum

ssp.

californicum

Nevski

Poaceae

SI

3.1

9.25

33

2.9

11

11


Melicageyeri

Munro

Poaceae

WI/IN

1.2

65

10.4

11

11


Poapiperi

Hitchc.

Poaceae

BE

5.4

21.5

45.5

14.3


Poarhizomata

Hitchc.

Poaceae

WI

1.8

3.5

21.5

-4.3


Poatenerrima

Scribn.

Poaceae

SI

3.3

13

43

1.3

CBR

11


Scribneria

bolanderi

(Thurb.)Hack.

Poaceae

WI

1.7

5.1

31

21

11

11

Annualgram.

Stipalemmoniivar.pubescens

(Vasey)Scribn.

Poaceae

BE

4.8

14.5

36

2.3

3.2

11


Stipanelsoniivar.dorei

Scribn.

Poaceae

WI/IN

12

21

-1


Stipastillm

anii

Bol.

Poaceae

WI/IN

1.1

2.1

21.1

-CBR


Collomia

diversifolia

Greene

Polemoniaceae

SE

5.6

33.5

66

1.2

4.3

11

Annualforb

Collomia

tinctoria

Kellogg

Polemoniaceae

WI

1.8

7.1

42

1.5

11

1Annualforb

Gilia

capitata

ssp.capitata

Sim

sPolemoniaceae

WI

1.6

4.75

31

1.2

11


Leptosiphonambiguus

(Rattan)J.

M.Porter

&L.

A.Johnson

Polemoniaceae

SE

5.8

17.5

36

0.6

11

Annualforb

Leptosiphonbolanderi

(A.Gray)J.M.Porter

&L.A

.Johnson

Polemoniaceae

WI/IN

1.3

2.5

21.3

-1

11

Annualforb

Leptosiphonlatisectus

(E.G.Buxton)J.

M.

Porter

&L.A.Johnson

Polemoniaceae

WI

26

32

01

Annualforb

Leptosiphonliniflorus

(Benth.)J.

M.Porter

&L.

A.Johnson

Polemoniaceae

WI

1.6

6.25

41.5

1.2

11

11

1Annualforb

Leptosiphonnuttalliissp.howellii

(A.Gray)J.

M.Porter

&L.A.Johnson

Polemoniaceae

BE

5.3

31.5

66

1.3

1Perennialforb

Linanthusdichotomus

Benth.

Polemoniaceae

SI

2.5

12.4

53

2.4

11

11

1Annualforb

Navarretia

heterodoxa

(Greene)

Greene

Polemoniaceae

SI

2.8

14

53

2.4

1Annualforb

Navarretia

jepsonii

V.L.Bailey

exJeps.

Polemoniaceae

SE

5.6

22.5

45.5

0.6

4.3

1Annualforb

Navarretia

linearifoliassp.pinnatisecta

(Howell)L.A

.Johnson

Polemoniaceae

BE/SI

3.8

19

53

2.2


Navarretia

mitracarpa

Greene

Polemoniaceae

SE

5.9

23.5

46

0.5

1Annualforb

Navarretia

paradoxiclara

L.A

.Johnson&

D.Gowen

Polemoniaceae

SE

618

36

01B.3

1Annualforb

Navarretia

paradoxinota

L.A

.Johnson&

D.Gowen

Polemoniaceae

SE

618

36

01B.3

1Annualforb

Navarretia

pubescens

(Benth.)Hook.&

Arn.

Polemoniaceae

WI

26

32

11

11

1Annualforb

Navarretia

rosulata

Brand

Polemoniaceae

SE

618

36

01B.2

11

Annualforb

Navarretia

sinistra

(M.E.Jones)L.A.Johnson

Polemoniaceae

SI

2.5

7.5

33

1.8

11

1Annualforb

Phloxhirsuta

E.E.Nelson

Polemoniaceae

SE

618

36

01B.2

1Perennialforb

Polemonium

chartaceum

H.Mason

Polemoniaceae

WI

1.6

8.1

52

1.5

1B.3

11

Perennialforb

Polemonium

eddyense

Stubbs

Polemoniaceae

SE

618

36

01B.2

1Perennialforb

Polygala

cornuta

var.cornuta

Kellogg

Polygalaceae

WI

2.3

94

21.3

11

Perennialforb,shrub

Chorizanthebreweri

S.Watson

Polygonaceae

BE

5.4

21.5

45.5

11B.3

1Annualforb

Chorizanthepalm

eri

S.Watson

Polygonaceae

BE

4.9

24.5

56

1.6

4.2

1Annualforb

Chorizantheuniaristata

Torr.&

A.Gray

Polygonaceae

SI

2.7

10.8

42.5

21


Chorizantheventricosa

Goodman

Polygonaceae

BE

5.3

16

36

1.2

4.3

1Annualforb

Eriogonum

alpinum

Engelm.

Polygonaceae

SE

630

56

01B.2

1Perennialforb

Eriogonum

argillosum

J.T.Howell

Polygonaceae

SI

3.1

12.5

43

2.6

4.3

11

Annualforb

Eriogonum

cedrorum

Reveal&

Raiche

Polygonaceae

SE

612

26

-1B.3

1Perennialforb

Eriogonum

clavatum

Small

Polygonaceae

SI

3.3

10

33

2.5

1Annualforb

Eriogonum

compositum

var.compositum

Douglasex

Benth.

Polygonaceae

WI

1.7

5.1

32

1.5

CBR

11

Perennialforb

Eriogonum

congdonii

(S.Stokes)Reveal

Polygonaceae

BE

5.1

35.5

76

1.7

4.3

1Shrub

Eriogonum

covilleanum

Eastw.

Polygonaceae

SI

312

43

1.6

CBR

1Annualforb

Eriogonum

dasyanthem

um

Torr.&

A.Gray

Polygonaceae

SI

36

23

-CBR

1Annualforb

Eriogonum

diclinum

Rev.

Polygonaceae

SI

3.2

9.5

33

32B.3

1Perennialforb

Eriogonum

elatum

var.villosum

Douglasex

Benth.

Polygonaceae

SI

3.3

13

43.5

3.2

11

Perennialforb

Eriogonum

hirtellum

J.T.Howell&

Bacig.

Polygonaceae

SE

618

36

01B.3

1Perennialforb

(rhiz.)

Eriogonum

hirtiflorum

A.Grayex

S.Watson

Polygonaceae

SI

3.3

13

43.5

3.2

CBR

11

11

1Annualforb

102 [Vol. 67MADRONO


APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Eriogonum

kelloggii

A.Gray

Polygonaceae

SE

624

46

01B.2

1Perennialforb

Eriogonum

libertini

Reveal

Polygonaceae

SE

636

66

04.2

11

Perennialforb

Eriogonum

luteolum

var.caninum

Greene

Polygonaceae

SE

5.5

27.5

56

0.9

1B.2

11

Annualforb

Eriogonum

luteolum

var.luteolum

Greene

Polygonaceae

BE/SI

3.8

15

43

1.5

11

11

Annualforb

Eriogonum

luteolum

var.pedunculatum

Greene

Polygonaceae

BE

510

25

-CBR

1Annualforb

Eriogonum

nervulosum

(S.Stokes)Reveal

Polygonaceae

SE

618

36

01B.2

1Perennialforb

(rhiz.)

Eriogonum

nudum

var.indictum

Douglasex

Benth.

Polygonaceae

WI

1.5

4.5

31

0.6

4.2

1Perennialforb

Eriogonum

nudum

var.oblongifolium

Douglasex

Benth.

Polygonaceae

WI

26

32

01

11

Perennialforb

Eriogonum

pendulum

S.Watson

Polygonaceae

SE

618

36

02B.2

1Perennialforb

Eriogonum

pyrolifolium

Hook.

Polygonaceae

WI/IN

13

30

1.7

11

Perennialforb

Eriogonum

siskiyouense

Small

Polygonaceae

BE

5.4

32.5

66

1.2

4.3

1Perennialforb

Eriogonum

strictum

var.greenei

Benth.

Polygonaceae

SE

5.9

29.5

56

0.4

4.3

11

Perennialforb

Eriogonum

ternatum

Howell

Polygonaceae

SE

618

36

04.3

11

Perennialforb

Eriogonum

tripodum

Greene

Polygonaceae

BE

5.3

26.5

56

1.3

4.2

11

Shrub

Eriogonum

umbellatum

var.argus

Torr.

Polygonaceae

SI

312

43.5

1.4

CBR

11

Perennialforb

Eriogonum

umbellatum

var.bahiiform

eTorr.

Polygonaceae

BE/SI

3.5

21

63

1.2

4.2

11

1Perennialforb

Eriogonum

umbellatum

var.goodmanii

Torr.

Polygonaceae

SI

3.3

10

33

2.5

CBR

1Perennialforb

Eriogonum

umbellatum

var.humistratum

Torr.

Polygonaceae

BE

4.5

27.3

65

2.1

4.3

11

Perennialforb

Eriogonum

umbellatum

var.nelsoniorum

Torr.

Polygonaceae

SE

5.5

11

25.5

-1

1Perennialforb

Eriogonum

umbellatum

var.speciosum

Torr.

Polygonaceae

BE/SI

4.2

21

54

1.3

CBR

11

Perennialforb

Eriogonum

ursinum

S.Watson

Polygonaceae

WI/IN

1.1

2.25

21.1

-1

1Perennialforb

Eriogonum

vimineum

Douglasex

Benth.

Polygonaceae

WI/IN

13

31

01

1Annualforb

Polygonum

majus

(Meisn.)Piper

Polygonaceae

WI

1.5

4.5

32

0.9

11

Annualforb

Polygonum

spergulariiform

eMeisn.ex

Small

Polygonaceae

SI

318.3

62.5

2.1

11

11

Annualforb

Systenothecavortriedei

(Brandegee)Reveal&

Hardham

Polygonaceae

SI

3.1

12.3

42.5

2.2

4.3

1Annualforb

Primula

clevelandiivar.patula

(Greene)

Mast

&Reveal

Primulaceae

SI

39

33

01

11

Perennialforb

Adiantum

aleuticum

(Rupr.)C.A.Paris

Pteridaceae

WI

2.4

11.8

52

1.2

11

11

1Perennialforb

Aspidotiscarlotta-halliae

(W.H.Wagner

&E.F.

Gilbert)

Lellinger

Pteridaceae

BE

5.3

26.5

56

1.1

4.2

11

Perennialforb

(rhiz.)

Aspidotisdensa

(Brack.)Lellinger

Pteridaceae

SI

3.4

31

93

1.2

11

11

1Perennialforb

Pellaea

brachyptera

(T.Moore)Baker

Pteridaceae

WI

1.5

4.5

32

0.9

11

1Perennialforb

Anem

onedrummondiivar.drummondii

S.Watson

Ranunculaceae

WI

2.3

6.75

32

1.6

11

Perennialforb

Anem

oneoreganavar.oregana

A.Gray

Ranunculaceae

SI

36

23

-1

Perennialforb

Aquilegia

exim

iaVanHoutteex

Planch..

Ranunculaceae

BE/SI

4.2

25

63.5

1.5

11

1Perennialforb

Delphinium

hesperium

ssp.hesperium

A.Gray

Ranunculaceae

SI

2.7

83

30.6

11

1Perennialforb

Delphinium

nuttallianum

Pritz.

Ranunculaceae

WI/IN

1.4

4.1

31

1.5

11

Perennialforb

Delphinium

parryissp.eastwoodiae

A.Gray

Ranunculaceae

BE/SI

3.7

11

34

2.5

1B.2

1Perennialforb

Delphinium

uliginosum

Curran

Ranunculaceae

SE

5.7

28.5

56

0.9

4.2

1Perennialforb

Ceanothusarcuatus

McM

inn

Rhamnaceae

BE/SI

3.5

72

3.5

-1

11

Shrub

Ceanothusconfusus

J.T.Howell

Rhamnaceae

WI/IN

1.3

2.5

21.3

-1B.1

1Shrub

Ceanothuscuneatusvar.cuneatus

(Hook.)Nutt.

Rhamnaceae

WI

1.5

6.1

41.5

1.3

11

11

1Shrub

Ceanothusdivergens

Parry

Rhamnaceae

WI

24

22

-1B.2

1Shrub

Ceanothusferrisiae

McM

inn

Rhamnaceae

SE

624

46

01B.1

1Shrub

Ceanothusfoliosusvar.medius

Parry

Rhamnaceae

BE/SI

412

33

1.7

11

Shrub

Ceanothusjepsonii

Greene

Rhamnaceae

SE

618

36

01

1Shrub

Ceanothusmasonii

McM

inn

Rhamnaceae

SI

3.3

6.5

23

-1B.2

1Shrub

Ceanothuspumilus

Greene

Rhamnaceae

SE

5.7

28.5

56

0.9

11

Shrub

Ceanothusroderickii

W.Knight

Rhamnaceae

WI

1.7

53

21.5

1B.1

1Shrub

Ceanothussonomensis

J.T.Howell

Rhamnaceae

WI/IN

1.3

43

21.2

1B.2

1Shrub

Frangula

californicassp.crassifolia

(Eschsch.)A.Gray

Rhamnaceae

BE

4.8

19

46

2.5

11

Shrub

Frangula

californicassp.occidentalis

(Eschsch.)A.Gray

Rhamnaceae

SE

624

46

01

11

Shrub

Frangula

californicassp.tomentella

(Eschsch.)A.Gray

Rhamnaceae

WI

1.5

64

0.8

1.7

11

11

1Shrub

Adenostomafasciculatum

Hook.&

Arn.

Rosaceae

WI/IN

1.3

5.2

41.1

1.4

11

11

Shrub



APPENDIX

1.CONTIN

UED.

Taxon1

Authority

Family

Aff2

Mean3

Sum

4Sourc.

Med

5SD

6Rarity

7

Geog.Distribution8

Lifeform

9KL

NC

BA

SC

SN

Holodiscusdiscolor

(Pursh)Maxim

.Rosaceae

WI/IN

13

31

11

11

11

Shrub

Horkelia

congesta

var.nem

orosa

Hook.

Rosaceae

BE/SI

3.8

7.5

23.5

-2B.1

1Perennialforb

Horkelia

daucifolia

(Greene)

Rydb.

Rosaceae

BE/SI

3.8

15

43

1.5

11

Perennialforb

Horkelia

daucifoliavar.daucifolia

(Greene)

Rydb.

Rosaceae

BE

4.5

92

4.5

-1

Perennialforb

Horkelia

sericata

S.Watson

Rosaceae

SE

5.6

22.5

46

14.3

1Perennialforb

Horkelia

tridentata

var.flavescens

Torr.

Rosaceae

SI

39

32

1.7

11

1Perennialforb

Ivesia

gordonii

(Hook.)Torr.&

A.Gray

Rosaceae

WI

1.6

3.25

21.6

-1

11

Perennialforb

Ivesia

pickeringii

Torr.ex

A.Gray

Rosaceae

BE

5.4

32.5

66

11B.2

1Perennialforb

Potentillacristae

Ferlatte&

Strother

Rosaceae

SI

3.1

12.5

43

01B.3

1Perennialforb

Sanguisorbaofficinalis

L.

Rosaceae

BE/SI

4.2

12.5

33

1.7

2B.2

11

Perennialforb

(rhiz.)

Galium

ambiguum

ssp.ambiguum

W.Wight

Rubiaceae

SI

3.3

10

33

2.5

11

Perennialforb

Galium

ambiguum

ssp.siskiyouense

W.Wight

Rubiaceae

SE

5.5

27.5

56

0.9

11

Perennialforb

Galium

andrewsiissp.andrewsii

A.Gray

Rubiaceae

SI

3.2

16

53

1.9

11

1Perennialforb

Galium

andrewsiissp.gatense

A.Gray

Rubiaceae

BE

5.1

20.5

45

0.8

4.2

11

Perennialforb

Galium

andrewsiissp.interm

edium

A.Gray

Rubiaceae

WI/IN

1.4

2.75

21.4

-1

Perennialforb

Galium

clem

entis

Eastw.

Rubiaceae

WI/IN

12

21

-1B.3

1Perennialforb

Galium

hardhamiae

Dem

pster

Rubiaceae

SE

624

46

01B.3

1Perennialforb

Galium

serpenticum

ssp.scotticum

Dem

pster

Rubiaceae

SE

5.9

29.5

56

0.4

1B.2

1Perennialforb

Salixbreweri

Bebb

Salicaceae

SE

630

56

01

11

Shrub

Salixdelnortensis

C.K.Schneid.

Salicaceae

SE

618

36

04.3

1Shrub

Salixsitchensis

Sansonex

Bong.

Salicaceae

WI

1.6

4.75

31

1.2

11

11

Tree,

shrub

Darlingtonia

californica

Torr.

Sarraceniaceae

BE/SI

4.1

32.5

84

1.4

4.2

11

Perennialforb

(carn.)

Micranthes

howellii

(Greene)

Small

Saxifragaceae

BE/SI

3.8

7.5

23.5

-4.3

1Perennialforb

Odontostomum

hartwegii

Torr.

Tecophilaeaceae

SI

2.7

83

30.6

11

Perennialforb

Brodiaea

roseassp.rosea

(Greene)

Baker

Them

idaceae

BE/SI

412

34

23.1

11

Perennialforb

Brodiaea

sierrae

R.E.Preston

Them

idaceae

WI

1.7

53

11.7

4.3

1Perennialforb

Brodiaea

stellaris

S.Watson

Them

idaceae

BE/SI

3.7

11

34

2.5

1Perennialforb

Muilla

maritima

(Torr.)S.Watson

Them

idaceae

WI

26

32

11

11

1Perennialforb

Triteleia

bridgesii

(S.Watson)Greene

Them

idaceae

SI

3.3

13

43.5

1.7

11

1Perennialforb

Triteleia

crocea

(Alph.Wood)Greene

Them

idaceae

BE/SI

4.1

32.5

83.7

1.8

1Perennialforb

Triteleia

ixioides

ssp.cookii

(W.T.Aiton)Greene

Them

idaceae

BE

4.5

13.5

36

2.9

1B.3

1Perennialforb

Triteleia

peduncularis

Lindl.

Them

idaceae

BE/SI

3.8

19

53

2.2

11

11

Perennialforb

Verbenacalifornica

Moldenke

Verbenaceae

BE

4.8

14.5

34

1.2

1B.1

1Perennialforb

Viola

cuneata

S.Watson

Violaceae

BE

5.2

31

66

1.3

CBR

11

Perennialforb

Viola

douglasii

Steud.

Violaceae

SI

2.8

13.8

52

21

11

11

Perennialforb

Viola

hallii

A.Gray

Violaceae

BE/SI

416

44

2.3

CBR

11

Perennialforb

Viola

lobata

ssp.lobata

Benth.

Violaceae

WI

2.3

11.4

52

2.4

11

1Perennialforb

(rhiz.)

Viola

ocellata

Torr.&

A.Gray

Violaceae

SI

2.5

12.5

53

0.9

11

11

Perennialforb

Viola

primulifoliassp.occidentalis

L.

Violaceae

BE

5.1

25.5

56

1.4

1B.2

1Perennialforb

(rhiz.)

Viola

purpureassp.integrifolia

Kellogg

Violaceae

WI/IN

1.3

43

21.2

11

1Perennialforb

104 [Vol. 67MADRONO


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